Caracal

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Caracal
Caracl (01), Paris, decembre 2013.jpg
Caracal at Jardin des Plantes, Paris.
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Felinae
Genus: Caracal
Species:
C. caracal
Binomial name
Caracal caracal
(Schreber, 1776)
Subspecies

See text

Caracal distribution.jpg
Distribution of Caracal, 2016 [1]
Synonyms [2]
  • C. aharonii(Matschie, 1912)
  • C. bengalensis(J. B. Fischer, 1829)
  • C. berberorumMatschie, 1892
  • C. coloniae Thomas, 1926
  • C. corylinus(Matschie, 1912)
  • C. medjerdae(Matschie, 1912)
  • C. melanotis Gray, 1843
  • C. melanotixGray, 1843
  • C. michaelisHeptner, 1945
  • C. roothi(Roberts, 1926)
  • C. spatzi(Matschie, 1912)

The caracal (Caracal caracal) /ˈkærəkæl/ is a medium-sized wild cat native to Africa, the Middle East, Central Asia, and India. It is characterised by a robust build, long legs, a short face, long tufted ears, and long canine teeth. Its coat is uniformly reddish tan or sandy, while the ventral parts are lighter with small reddish markings. It reaches 40–50 cm (16–20 in) at the shoulder and weighs 8–19 kg (18–42 lb). It was first scientifically described by German naturalist Johann Christian Daniel von Schreber in 1776. Three subspecies are recognised.

Contents

Typically nocturnal, the caracal is highly secretive and difficult to observe. It is territorial, and lives mainly alone or in pairs. The caracal is a carnivore that typically preys upon small mammals, birds, and rodents. It can leap higher than 4 metres (12 ft) and catch birds in midair. It stalks its prey until it is within 5 m (16 ft) of it, after which it runs it down and kills its prey with a bite to the throat or to the back of the neck. Both sexes become sexually mature by the time they are one year old and breed throughout the year. Gestation lasts between two and three months, resulting in a litter of one to six kittens. Juveniles leave their mothers at the age of nine to ten months, though a few females stay back with their mothers. The average lifespan of captive caracals is nearly 16 years.

Caracals were tamed and used for hunting in ancient Egypt.

Taxonomy and phylogeny

Felis caracal was the scientific name used by Johann Christian Daniel von Schreber in 1776 who described a caracal skin from the Cape of Good Hope. [3] In 1843, British zoologist John Edward Gray placed it in the genus Caracal. It is placed in the family Felidae and subfamily Felinae. [2]

In the 19th and 20th centuries, several caracal specimens were described and proposed as subspecies. Since 2017, three subspecies are recognised as valid: [4]

Phylogeny

Results of a phylogenetic study indicates that the caracal and the African golden cat (Caracal aurata) diverged between 2.93 and 1.19 million years ago. These two species together with the serval (Leptailurus serval) form the Caracal lineage, which diverged between 11.56 and 6.66 million years ago. [7] [8] The ancestor of this lineage arrived in Africa between 8.5 and 5.6 million years ago. [9]

The relationship of the caracal is considered as follows: [7] [8]

Pardofelis

Marbled cat (P. marmorata)

Catopuma

Bay cat (C badia)

Asian golden cat (C temminckii)

 

Caracal
Leptailurus

Serval (L. serval)

Caracal

Caracal

African golden cat (C aurata)

lineage

Leopardus

Lynx

Acinonyx

Puma

Otocolobus

Prionailurus

Felis

Etymology

The name 'caracal' was proposed by Georges Buffon in 1761 who referred to its Turkish name 'Karrah-kulak' or 'Kara-coulac', meaning 'cat with black ears'. [10] [11] The 'lynx' of the Greeks and Romans was most probably the caracal, and the name 'lynx' is sometimes still applied to it, but the present-day lynx proper is a separate genus. [12]

The caracal is also known as desert lynx and Persian lynx. [13]

Local names

Its name in the Tigrinya language is ጭክ ኣንበሳ (ch’ok anbessa), which means 'bearded lion'. [14]

Characteristics

A close facial view of a caracal, with the typical tufted ears and the black facial markings Caracal001.jpg
A close facial view of a caracal, with the typical tufted ears and the black facial markings

The caracal is a slender, moderately sized cat characterised by a robust build, a short face, long canine teeth, tufted ears, and long legs. It reaches nearly 40–50 cm (16–20 in) at the shoulder. The tan, bushy tail extends to the hocks. [15] [16] The caracal is sexually dimorphic; the females are smaller than the males in most bodily parameters. [17]

The prominent facial features include the 4.5-cm-long black tufts on the ears, two black stripes from the forehead to the nose, the black outline of the mouth, the distinctive black facial markings, and the white patches surrounding the eyes and the mouth. The eyes appear to be narrowly open due to the lowered upper eyelid, probably an adaptation to shield the eyes from the sun's glare. The ear tufts may start drooping as the animal ages. The coat is uniformly reddish tan or sandy, though black caracals are also known. The underbelly and the insides of the legs are lighter, often with small reddish markings. [17] The fur, soft, short, and dense, grows coarser in the summer. The ground hairs (the basal layer of hair covering the coat) are denser in winter than in summer. The length of the guard hairs (the hair extending above the ground hairs) can be up to 3 cm (1.2 in) long in winter, but shorten to 2 cm (0.8 in) in summer. [18] These features indicate the onset of moulting in the hot season, typically in October and November. [19] The hind legs are longer than the forelegs, so the body appears to be sloping downward from the rump. [16] [17]

Male caracals measure in head-to-body length 78–108 cm (31–43 in) and have 21–34 cm (8.3–13.4 in) long tails; 77 male caracals ranged in weight between 7.2 and 19 kg (16 and 42 lb). The head-to-body length of females is 71–102.9 cm (28.0–40.5 in) with a tail of 18–31.5 cm (7.1–12.4 in); 63 females ranged in weight between 7 and 15.9 kg (15 and 35 lb). [20]

The caracal is often confused with the lynx, as both cats have tufted ears. However, a notable point of difference between the two is that the lynx is spotted and blotched, while the caracal shows no such markings on the coat. [17] The African golden cat has a similar build as the caracal's, but is darker and lacks the ear tufts. The sympatric serval can be distinguished from the caracal by the former's lack of ear tufts, white spots behind the ears, spotted coat, longer legs, longer tail, and smaller footprints. [18] [21]

The skull of the caracal is high and rounded, featuring large auditory bullae, a well-developed supraoccipital crest normal to the sagittal crest, and a strong lower jaw. The caracal has a total of 30 teeth; the dental formula is 3.1.3.13.1.2.1. The deciduous dentition is 3.1.23.1.2. The striking canines are up to 2 cm (0.8 in) long, heavy, and sharp; these are used to give the killing bite to the prey. The caracal lacks the second upper premolars, and the upper molars are diminutive. [19] The large paws, similar to those of the cheetah, [22] consist of four digits in the hind legs and five in the fore legs. [18] The first digit of the fore leg remains above the ground and features the dewclaw. The claws, sharp and retractable (able to be drawn in), are larger but less curved in the hind legs. [18]

Distribution and habitat

Caracals inhabit dry areas with some cover. SanWild07a.jpg
Caracals inhabit dry areas with some cover.

In Africa, the caracal is widely distributed south of the Sahara, but considered rare in North Africa. In Asia, it occurs from the Arabian Peninsula, Middle East, Turkmenistan, Uzbekistan to western India. [1] It inhabits forests, savannas, marshy lowlands, semideserts, and scrub forests, but prefers dry areas with low rainfall and availability of cover. In montane habitats such as the Ethiopian Highlands, it occurs up to an altitude of 3,000 m (9,800 ft). [18]

In Ethiopia's Degua Tembien massif, they can be seen along roads, sometimes as road kills. [14]

In the Emirate of Abu Dhabi, a male caracal was recorded by camera-traps in Jebel Hafeet National Park in the Al-Ain Region, Abu Dhabi in February 2019, the first such record since 1984. [23] [24]

In Uzbekistan, caracals were recorded only in the desert regions of the Ustyurt Plateau and Kyzylkum Desert. Between 2000 and 2017, 15 individuals were sighted alive, and at least 11 were killed by herders. [25]

In Iran, the caracal has been recorded in Abbasabad Naein Reserve, Bahram’gur Protected Area, Kavir National Park and in Yazd Province. [26] [27] [28] [29] [30] [31]

In India, the caracal occurs in Sariska Tiger Reserve and Ranthambhore Tiger Reserve. [32] [33]

Ecology and behaviour

Caracals are efficient climbers. 20120219 Olmense Zoo (64).jpg
Caracals are efficient climbers.

The caracal is typically nocturnal (active at night), though some activity may be observed during the day as well. However, the cat is so secretive and difficult to observe that its activity at daytime might easily go unnoticed. [19] A study in South Africa showed that caracals are most active when the air temperature drops below 20 °C (68 °F); activity typically ceases at higher temperatures. [34] A solitary cat, the caracal mainly occurs alone or in pairs; the only groups seen are of mothers with their offspring. [16] Females in oestrus temporarily pair with males. A territorial animal, the caracal marks rocks and vegetation in its territory with urine and probably with dung, which is not covered with soil. Claw scratching is prominent, and dung middens are typically not formed. [18] In Israel, males are found to have territories averaging 220 km2 (85 sq mi), while that of females averaged 57 km2 (22 sq mi). The male territories vary from 270–1,116 km2 (104–431 sq mi) in Saudi Arabia. In Mountain Zebra National Park, the home ranges of females vary between 4.0 and 6.5 km2 (1.5 and 2.5 sq mi). These territories overlap extensively. [17] The conspicuous ear tufts and the facial markings often serve as a method of visual communication; caracals have been observed interacting with each other by moving the head from side to side so that the tufts flicker rapidly. Like other cats, the caracal meows, growls, hisses, spits, and purrs. [16]

Diet and hunting

A caracal being fed Caracal caracal -Toronto Zoo, Ontario, Canada-8b.jpg
A caracal being fed

A carnivore, the caracal typically preys upon small mammals, birds, and rodents. Studies in South Africa have reported that it preys on the Cape grysbok, the common duiker, sheep, goats, bush vlei rats, rock hyraxes, hares, and birds. [35] [36] [37] A study in western India showed that rodents comprise a significant portion of the diet. [32] They will feed from a variety of sources, but tend to focus on the most abundant one. [38] Grasses and grapes are taken occasionally to clear their immune system and stomach of any parasites. [39] Larger antelopes such as young kudu, bushbuck, impala, mountain reedbuck, and springbok may also be targeted. Mammals generally comprise at least 80% of the diet. [18] Lizards, snakes, and insects are infrequently eaten. [1] In South Africa, caracals have been documented preying on small livestock in areas where sheep and goat are farmed. Livestock is however only a small and seasonal portion of their diet when wild prey is scarce. [35] [20]

Its speed and agility make it an efficient hunter, able to take down prey two to three times its size. [1] The powerful hind legs allow it to leap more than 3 m (10 ft) in the air to catch birds on the wing. [17] [40] [41] It can even twist and change its direction mid-air. [17] It is an adroit climber. [17] It stalks its prey until it is within 5 m (16 ft), following which it can launch into a sprint. While large prey such as antelopes are suffocated by a throat bite, smaller prey are killed by a bite on the back of the neck. [17] Kills are consumed immediately, and less commonly dragged to cover. It returns to large kills if undisturbed. [18] It has been observed to begin feeding on antelope kills at the hind parts. [19] It may scavenge at times, though this has not been frequently observed. [35] It often has to compete with foxes, wolves, leopards, and hyaena for prey. [22]

Reproduction

Caracal mother and kitten Caracal female and kitten playing.jpg
Caracal mother and kitten

Both sexes become sexually mature by the time they are a year old; production of gametes begins even earlier at seven to ten months. However, successful mating takes place only at 12 to 15 months. Breeding takes place throughout the year. Oestrus, one to three days long, recurs every two weeks unless the female is pregnant. Females in oestrus show a spike in urine-marking, and form temporary pairs with males. Mating has not been extensively studied; a limited number of observations suggest that copulation, lasting nearly four minutes on an average, begins with the male smelling the areas urine-marked by the female, which then rolls on the ground. Following this, the male approaches and mounts the female. The pair separate after copulation. [17] [18]

Gestation lasts about two to three months, following which a litter consisting of one to six kittens is born. Births generally peak from October to February. Births take place in dense vegetation or deserted burrows of aardvarks and porcupines. Kittens are born with their eyes and ears shut and the claws not retractable (unable to be drawn inside); the coat resembles that of adults, but the abdomen is spotted. Eyes open by ten days, but it takes longer for the vision to become normal. The ears become erect and the claws become retractable by the third or the fourth week. Around the same time, the kittens start roaming their birthplace, and start playing among themselves by the fifth or the sixth week. They begin taking solid food around the same time; they have to wait for nearly three months before they make their first kill. As the kittens start moving about by themselves, the mother starts shifting them every day. All the milk teeth appear in 50 days, and permanent dentition is completed in 10 months. Juveniles begin dispersing at nine to ten months, though a few females stay back with their mothers. The average lifespan of the caracal in captivity is nearly 16 years. [17] [22] [42]

In the 1990s, a captive caracal spontaneously mated with a domestic cat in the Moscow Zoo, resulting in a felid hybrid offspring. [43]

Threats

The caracal is listed as Least Concern on the IUCN Red List since 2002, as it is widely distributed in over 50 range countries, where the threats to caracal populations vary in extent. Habitat loss due to agricultural expansion, the building of roads and settlements is a major threat in all range countries. It is thought to be close to extinction in North Africa, Critically Endangered in Pakistan, Endangered in Jordan, but stable in central and Southern Africa. Local people kill caracal to protect livestock, or in retaliation for its preying on small livestock. Additionally, it is threatened by hunting for the pet trade on the Arabian Peninsula. In Turkey and Iran, caracals are frequently killed in road accidents. [1] In Uzbekistan, the major threat to caracal is killing by herders in retaliation for livestock losses. Guarding techniques and sheds are inadequate to protect small livestock like goats and sheep from being attacked by predators. Additionally, similarly to Ethiopia, heavy-traffic roads crossing caracal habitat pose a potential threat for the species. [25]

Conservation

A caracal in the San Diego Zoo Lightmatter caracal.jpg
A caracal in the San Diego Zoo

African caracal populations are listed under CITES Appendix II, while Asian populations come under CITES Appendix I. Hunting of caracal is prohibited in Afghanistan, Algeria, Egypt, India, Iran, Israel, Jordan, Kazakhstan, Lebanon, Morocco, Pakistan, Syria, Tajikistan, Tunisia, Turkey, Turkmenistan, and Uzbekistan. Caracals occur in a number of protected areas across their range. [1]

In South Africa, the caracal is considered Least Concern, as it is widespread and adaptable to a variety of habitats. It is tolerant to human-dominated areas, although it has been persecuted for many decades. Farmers are encouraged to report sightings of caracals, both dead and alive, as well as livestock killed by caracals to the national Predation Management Information Centre. [44]

The Central Asian caracal population is listed as Critically Endangered in Uzbekistan since 2009, and in Kazakhstan since 2010. [25] [45] [46]

In culture

Chinese emperors used caracals as gifts. In the 13th and the 14th centuries, Yuan dynasty rulers bought numerous caracals, cheetahs, and tigers from Muslim merchants in the western parts of the empire in return for gold, silver, cash, and silk. According to the Ming Shilu , the subsequent Ming dynasty continued this practice. Until as recently as the 20th century, the caracal was used in hunts by Indian rulers to hunt small game, while the cheetah was used for larger game. [47] In those times, caracals were exposed to a flock of pigeons and people would bet on which caracal would kill the largest number of pigeons. This probably gave rise to the expression "to put the cat among the pigeons". [41]

The caracal appears to have been religiously significant in the ancient Egyptian culture, as it occurs in paintings and as bronze figurines; sculptures are thought to have guarded the tombs of pharaohs. [48] Embalmed caracals have also been discovered. [49] Its pelt was used for making fur coats. [22]

Related Research Articles

Felidae Family of mammals

Felidae is a family of mammals in the order Carnivora, colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid. The term "cat" refers both to felids in general and specifically to the domestic cat.

Leopard A large cat native to Africa and Eurasia

The leopard is one of the five extant species in the genus Panthera, a member of the Felidae. It occurs in a wide range in sub-Saharan Africa, in small parts of Western and Central Asia, on the Indian subcontinent to Southeast and East Asia. It is listed as Vulnerable on the IUCN Red List because leopard populations are threatened by habitat loss and fragmentation, and are declining in large parts of the global range. In Hong Kong, Singapore, Kuwait, Syria, Libya, Tunisia and most likely in Morocco, leopard populations have already been extirpated. Contemporary records suggest that the leopard occurs in only 25% of its historical global range.

Bobcat Medium-sized North American wild cat

The bobcat, also known as the red lynx, is a medium-sized cat native to North America from southern Canada, most of the contiguous United States to Oaxaca in Mexico. It is listed as Least Concern on the IUCN Red List since 2002, because it is widely distributed and abundant. Although it has been hunted extensively both for sport and fur, populations have proven resilient though declining in some areas.

<i>Felis</i> Genus of mammals (cats)

Felis is a genus of small and medium-sized cat species native to most of Africa and south of 60° latitude in Europe and Asia to Indochina. The genus includes the domestic cat. The smallest Felis species is the black-footed cat with a head and body length from 38 to 42 cm. The largest is the jungle cat with a head and body length from 62 to 76 cm.

Wildcat Small wild cat

The wildcat is a species complex comprising two small wild cat species, the European wildcat and the African wildcat. The European wildcat inhabits forests in Europe and the Caucasus, while the African wildcat inhabits semi-arid landscapes and steppes in Africa, the Arabian Peninsula, Central Asia, into western India and western China. The wildcat species differ in fur pattern, tail, and size: the European wildcat has long fur and a bushy tail with a rounded tip; the smaller African wildcat is more faintly striped, has short sandy-gray fur and a tapering tail; the Asiatic wildcat is spotted.

Sand cat Small wild cat

The sand cat, also known as the sand dune cat, is a small wild cat living in sandy and stony deserts far from water sources. With its sandy to light grey fur, it is well camouflaged in a desert environment. Its head-and-body length ranges from 39–52 cm (15–20 in) with a 23–31 cm (9.1–12.2 in) long tail. Its 5–7 cm (2.0–2.8 in) long ears are set low on the sides of the head, aiding detection of prey moving underground. The long hair covering the soles of its feet insulate its foot pads against the extremely hot and cold temperatures in deserts.

Jungle cat Medium-sized wild cat

The jungle cat, also called reed cat and swamp cat, is a medium-sized cat native to the Middle East, South and Southeast Asia and southern China. It inhabits foremost wetlands like swamps, littoral and riparian areas with dense vegetation. It is listed as Least Concern on the IUCN Red List, and is mainly threatened by destruction of wetlands, trapping and poisoning.

Black-footed cat Small wild cat native to Southern Africa

The black-footed cat, also called the small-spotted cat, is the smallest wild cat in Africa, having a head-and-body length of 35–52 cm (14–20 in). Despite its name, only the soles of its feet are black or dark brown. With its bold small spots and stripes on the tawny fur, it is well camouflaged, especially on moonlit nights. It bears black streaks running from the corners of the eyes along the cheeks, and its banded tail has a black tip.

Chinese mountain cat Small wild cat

The Chinese mountain cat, also known as Chinese desert cat and Chinese steppe cat, is a wild cat endemic to western China that has been listed as Vulnerable on the IUCN Red List since 2002, as the effective population size may be fewer than 10,000 mature breeding individuals.

Pallass cat Small wild cat

Pallas's cat, also called the manul, is a small wild cat with a broad, but fragmented distribution in the grasslands and montane steppes of Central Asia. It is negatively affected by habitat degradation, prey base decline and hunting. It has been classified as Least Concern on the IUCN Red List since 2020.

African golden cat Small wild cat

The African golden cat is a wild cat endemic to the rainforests of West and Central Africa. It is threatened due to deforestation and bushmeat hunting and listed as Vulnerable on the IUCN Red List. It is a close relative of both the caracal and the serval. Previously, it was placed in the genus Profelis. Its body size ranges from 61 to 101 cm with a 16 to 46 cm long tail.

Serval Small wild cat

The serval is a wild cat native to Africa. It is rare in North Africa and the Sahel, but widespread in sub-Saharan countries except rainforest regions. On the IUCN Red List it is listed as Least Concern. Across its range, it occurs in protected areas, and hunting it is either prohibited or regulated in range countries.

Canada lynx Small wild cat

The Canada lynx is a medium-sized North American cat that ranges across Alaska, Canada and many of the contiguous United States. It is characterized by its long, dense fur, triangular ears with black tufts at the tips, and broad, snowshoe-like paws. Similar to the bobcat, the hindlimbs are longer than the forelimbs, so that the back slopes downward to the front. The Canada lynx stands 48–56 cm (19–22 in) tall at the shoulder and weighs between 5 and 17 kg. The lynx is a good swimmer and an agile climber. The Canada lynx was first described by Robert Kerr in 1792. Three subspecies have been proposed, but their validity is doubted.

Felinae subfamily of mammals

The Felinae are a subfamily of the family Felidae. This subfamily comprises the small cats having a bony hyoid, because of which they are able to purr but not roar.

Felid hybrid Offspring of two different species of Felidae

A felid hybrid is any of a number of hybrid between various species of the cat family, Felidae. This article deals with hybrids between the species of the subfamily Felinae.

European wildcat Small wild cat

The European wildcat is a wildcat species native to continental Europe, Scotland, Turkey and the Caucasus. It inhabits forests from the Iberian Peninsula, Italy, Central and Eastern Europe to the Caucasus. It has been extirpated in England and Wales.

Asiatic wildcat Small wild cat

The Asiatic wildcat is an African wildcat subspecies that occurs from the eastern Caspian Sea north to Kazakhstan, into western India, western China and southern Mongolia. It is also known as the Asian steppe wildcat and Indian desert cat. The status Least Concern in the IUCN Red List is attributed to the wildcat species complex. There is no information on current status or population numbers for the Asiatic wildcat's entire range, but populations are thought to be declining.

African wildcat Small wild cat

The African wildcat is a wildcat species native to Africa, West and Central Asia up to Rajasthan in India and Xinjiang in China. The IUCN Red List status Least Concern is attributed to the species Felis silvestris, which at the time of assessment also included the African wildcat as a subspecies.

Snow leopard species of mammal

The snow leopard, also known as the ounce, is a large cat native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is estimated to number less than 10,000 mature individuals and is expected to decline about 10% by 2040. It is threatened by poaching and habitat destruction following infrastructural developments. It inhabits alpine and subalpine zones at elevations from 3,000 to 4,500 m, ranging from eastern Afghanistan, the Himalayas and the Tibetan Plateau, to southern Siberia, Mongolia and western China. In the northern range countries, it also lives at lower elevations.

Sunda leopard cat Small wild cat

The Sunda leopard cat is a small wild cat species native to the Sundaland islands of Java, Bali, Borneo, Sumatra and the Philippines that is considered distinct from the leopard cat occurring in mainland South and Southeast Asia.

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