Ceratosauridae

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Ceratosaurids
Temporal range: Late Jurassic-Early Cretaceous, 155–112  Ma
O
S
D
C
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(Possible Early Jurassic record)
Genyodectes saurus jaws.jpg
Holotype jaws of Genyodectes
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Neoceratosauria
Family: Ceratosauridae
Marsh, 1884
Type species
Ceratosaurus nasicornis
Marsh, 1884
Genera

Ceratosauridae is an extinct family of theropod dinosaurs belonging to the infraorder Ceratosauria. The family's type genus, Ceratosaurus , was first found in Jurassic rocks from North America. Ceratosauridae is made up of the genera Ceratosaurus, found in North America, Tanzania, and Portugal, and Genyodectes , from the Early Cretaceous of Argentina. Unnamed probable ceratosaurids are known from limited material in the Middle Jurassic of Madagascar, the Late Jurassic of Switzerland, the Late Jurassic of Tanzania, and the Late Jurassic or possibly Early Cretaceous of Uruguay. [1]

Contents

Classification

Othniel Charles Marsh named the family Ceratosauridae in 1884 to contain the type species, Ceratosaurus nasicornis. Since then, a number of other species have been referred to this family, mainly from the genus Ceratosaurus. Currently recognized ceratosaurid species include Genyodectes serus and possibly Eoabelisaurus mefi,C. stechowi and Ostafrikasaurus are undiagnostic ceratosaurs. [2] Delcourt (2018) defined Ceratosauridae as "the most inclusive clade containing Ceratosaurus nasicornis but not Carnotaurus sastrei". [3]

Features

A historical skeletal reconstruction of Ceratosaurus. CeratosaurusSkeleton.jpg
A historical skeletal reconstruction of Ceratosaurus .

Due to the paucity of remains of Genyodectes, it is difficult to discern possible synapomorphies of the group from autapomorphies of Ceratosaurus; e.g. Ceratosaurus is different from other ceratosaurians by the very prominent horn on its snout; Genyodectes, however, was not found with a complete skull; whether it had a horn is unknown, so it cannot establish that the horn was a shared derived feature of the group. However, due to the shared similarities between the teeth of the two genera, synapomorphies have been recognized in the teeth. These are: overlap of the second and third premaxillary alveoli in palatal view, largest crown in subadults/adults higher than six centimeters, subquadrangular mesial denticles at two-thirds of the crown in lateral teeth. [4]

There are two known types of Ceratosaurus teeth: one with longitudinal ridges and the other with veined enamel. Both types of teeth have crowns with a teardrop-shaped cross section and carinae running up the middle. The cross section of the tooth's base depends on the position of the tooth in the mouth with front teeth having less symmetric cross sections.

Environment

Being found in the Morrison and Tendaguru put the family Ceratosauridae in the presence of other large predators. In North America, it is likely that members of the family such as C. nasicornis competed with allosaurids (A. fragilis) for food, such as sauropods common to the region at the time. In Africa and Europe members also competed with other large predators for similar food sources. The presence of C. nasicornis at the Cleveland-Lloyd Dinosaur Quarry along with the remains of several allosaurids is a good indication of just how close members of this family and other predators coexisted.

Sites containing ceratosaurids

See also

Related Research Articles

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Allosaurus is an extinct genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period. The name "Allosaurus" means "different lizard", alluding to its unique concave vertebrae. It is derived from the Greek words ἄλλος and σαῦρος. The first fossil remains that could definitively be ascribed to this genus were described in 1877 by famed paleontologist Othniel Charles Marsh. As one of the first well-known theropod dinosaurs, it has long attracted attention outside of paleontological circles.

<i>Ceratosaurus</i> Genus of theropod dinosaur from the Late Jurassic period

Ceratosaurus was a carnivorous theropod dinosaur that lived in the Late Jurassic period. The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is Ceratosaurus nasicornis.

<span class="mw-page-title-main">Ceratosauria</span> Extinct clade of dinosaurs

Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with Ceratosaurus than with birds. The oldest known ceratosaur, Saltriovenator, dates to the earliest part of the Jurassic, around 199 million years ago. Ceratosauria includes three major clades: Ceratosauridae, Noasauridae, and Abelisauridae, found primarily in the Southern Hemisphere. Originally, Ceratosauria included the above dinosaurs plus the Late Triassic to Early Jurassic Coelophysoidea and Dilophosauridae, implying a much earlier divergence of ceratosaurs from other theropods. However, most recent studies have shown that coelophysoids and dilophosaurids do not form a natural group with other ceratosaurs, and are excluded from this group.

<span class="mw-page-title-main">Tetanurae</span> Clade containing most theropod dinosaurs

Tetanurae is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans. Tetanurans are defined as all theropods more closely related to modern birds than to Ceratosaurus and contain the majority of predatory dinosaur diversity. Tetanurae likely diverged from its sister group, Ceratosauria, during the late Triassic. Tetanurae first appeared in the fossil record by the Early Jurassic about 190 mya and by the Middle Jurassic had become globally distributed.

<i>Torvosaurus</i> Megalosaurid theropod dinosaur genus from Late Jurassic Period

Torvosaurus is a genus of large megalosaurine theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle and Late Jurassic period in what is now Colorado, Portugal, Germany, and possibly England, Spain, Tanzania, and Uruguay. It contains two currently recognized species, Torvosaurus tanneri and Torvosaurus gurneyi, plus a third unnamed species from Germany.

<span class="mw-page-title-main">Carcharodontosauridae</span> Extinct family of dinosaurs

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<i>Rugops</i> Genus of dinosaur

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<span class="mw-page-title-main">Abelisauridae</span> Extinct family of dinosaurs

Abelisauridae is a family of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found on the modern continents of Africa and South America, as well as on the Indian subcontinent and the island of Madagascar. Isolated teeth were found in the Late Jurassic of Portugal, and the Late Cretaceous genera Tarascosaurus and Arcovenator have been described in France. Abelisaurids first appear in the fossil record of the early middle Jurassic period, and at least three genera survived until the end of the Mesozoic era 66 million years ago.

<span class="mw-page-title-main">Allosauridae</span> Extinct family of theropod dinosaurs

Allosauridae is a family of medium to large bipedal, carnivorous allosauroid theropod dinosaurs from the Late Jurassic. Allosauridae is a fairly old taxonomic group, having been first named by the American paleontologist Othniel Charles Marsh in 1878. Allosaurids are characterized by an astragalus with a restriction of the ascending process to the lateral part of the bone, a larger medial than lateral condyle, and a horizontal groove across the face of the condyles.

<i>Deltadromeus</i> Theropod dinosaur genus from mid-Cretaceous Period

Deltadromeus is a genus of theropod dinosaur from Northern Africa. It had long, unusually slender hind limbs for its size, suggesting that it was a swift runner. The skull is not known. One fossil specimen of a single species has been described, found in the Kem Kem Beds, which date to the mid-Cretaceous Period, about 95 million years ago. It may be a junior synonym of the contemporary Bahariasaurus. The classification of Deltadromeus has been in flux since its original description. In 2016, a South American theropod known as Gualicho shinyae was found to possess many similarities with Deltadromeus. Depending on the phylogenetic position of Gualicho, Deltadromeus may have been a neovenatorid carnosaur, a tyrannosauroid, or a basal coelurosaur if its close relation to Gualicho is legitimate. Other studies have considered it a ceratosaurian, more specifically a member of the family Noasauridae.

<span class="mw-page-title-main">Megalosauridae</span> Extinct family of dinosaurs

Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively primitive group of basal tetanurans containing two main subfamilies, Megalosaurinae and Afrovenatorinae, along with the basal genus Eustreptospondylus, an unresolved taxon which differs from both subfamilies.

<span class="mw-page-title-main">Abelisauroidea</span> Extinct clade of dinosaurs

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<i>Elaphrosaurus</i> Ceratosaurian theropod dinosaur genus from the Late Jurassic Period

Elaphrosaurus is a genus of ceratosaurian theropod dinosaur that lived approximately 154 to 150 million years ago during the Late Jurassic Period in what is now Tanzania in Africa. Elaphrosaurus was a medium-sized but lightly built member of the group that could grow up to 6.2 m (20 ft) long. Morphologically, this dinosaur is significant in two ways. Firstly, it has a relatively long body but is very shallow-chested for a theropod of its size. Secondly, it has very short hindlimbs in comparison with its body. Phylogenetic analyses indicate that this genus is likely a ceratosaur. Earlier suggestions that it is a late surviving coelophysoid have been examined but generally dismissed. Elaphrosaurus is currently believed to be a very close relative of Limusaurus, an unusual beaked ceratosaurian which may have been either herbivorous or omnivorous.

<i>Saurophaganax</i> Allosaurid theropod dinosaur genus from Late Jurassic period

Saurophaganax, also known as Allosaurus maximus or Saurophagus maximus, is a genus of large allosaurid dinosaur from the Morrison Formation of Late Jurassic Oklahoma, United States. Some paleontologists consider it to be a junior synonym and species of Allosaurus. Saurophaganax represents a very large Morrison allosaurid characterized by horizontal laminae at the bases of the dorsal neural spines above the transverse processes, and "meat-chopper" chevrons. It was the largest terrestrial carnivore of North America during the Late Jurassic, reaching 10.5 metres (34 ft) in length and 2.7–3.8 metric tons in body mass.

<i>Genyodectes</i> Extinct genus of dinosaurs

Genyodectes is a genus of ceratosaurian theropod dinosaur from the Lower Cretaceous (Aptian) of South America. The holotype material was collected from the Cerro Barcino Formation, Cañadón Grande, Departamento Paso de Indios in the Chubut Province of Argentina and consists of an incomplete snout, including the premaxillae, portions of both maxillas, the right and left dentary, many teeth, a fragment of the left splenial, and parts of the supradentaries. These elements are generally poorly preserved and some are in articulation. The premaxilla of Genyodectes possesses relatively large and protruding teeth, similar to those of Ceratosaurus. The specific name, serus, means "late". In 2016 it was estimated to be 6.25 meters in length and 790 kg in weight.

<i>Limusaurus</i> Genus of theropod dinosaur

Limusaurus is a genus of theropod dinosaur that lived in what is now China during the Late Jurassic, around 161 to 157 million years ago. The type and only species Limusaurus inextricabilis was described in 2009 from specimens found in the Upper Shishugou Formation in the Junggar Basin of China. The genus name consists of the Latin words for "mud" and "lizard", and the species name means "impossible to extricate", both referring to these specimens possibly dying after being mired. Limusaurus was a small, slender animal, about 1.7 m in length and 15 kg (33 lb) in weight, which had a long neck and legs but very small forelimbs. It underwent a drastic morphological transformation as it aged: while juveniles were toothed, these teeth were completely lost and replaced by a beak with age. Several of these features were convergently similar to the later ornithomimid theropods as well as the earlier non-dinosaurian shuvosaurids.

<i>Ostafrikasaurus</i> Genus of theropod dinosaur

Ostafrikasaurus is a genus of theropod dinosaur from the Late Jurassic period of what is now Lindi Region, Tanzania. It is known only from fossil teeth discovered sometime between 1909 and 1912, during an expedition to the Tendaguru Formation by the Natural History Museum of Berlin. Eight teeth were originally attributed to the dubious dinosaur genus Labrosaurus, and later to Ceratosaurus, both known from the North American Morrison Formation. Subsequent studies attributed two of these teeth to a spinosaurid dinosaur, and in 2012, Ostafrikasaurus crassiserratus was named by French palaeontologist Eric Buffetaut, with one tooth as the holotype, and the other referred to the same species. The generic name comes from the German word for German East Africa, the former name of the colony in which the fossils were found, while the specific name comes from the Latin words for "thick" and "serrated", in reference to the form of the animal's teeth.

<span class="mw-page-title-main">Orionides</span> Clade of dinosaurs

Orionides is a clade of tetanuran theropod dinosaurs from the Middle Jurassic to the Present. The clade includes most theropod dinosaurs, including birds.

<span class="mw-page-title-main">Timeline of ceratosaur research</span>

This timeline of ceratosaur research is a chronological listing of events in the history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became the apex predators of the southern hemisphere during the Late Cretaceous. The nature and taxonomic composition of the Ceratosauria has been controversial since the group was first distinguished in the late 19th century. In 1884 Othniel Charles Marsh described the new genus and species Ceratosaurus nasicornis from the Late Jurassic Morrison Formation of the western United States. He felt that it belonged in a new family that he called the Ceratosauridae. He created the new taxon Ceratosauria to include both the Ceratosauridae and the ostrich-like ornithomimids. The idea of the Ceratosauria was soon contested, however. Later that same decade both Lydekker and Marsh's hated rival Edward Drinker Cope argued that the taxon was invalid.

References

  1. Soto, Matías; Perea, Daniel (2008). "A ceratosaurid (Dinosauria, Theropoda) from the Late Jurassic-Early Cretaceous of Uruguay". Journal of Vertebrate Paleontology. 28 (2): 439–444. doi:10.1671/0272-4634(2008)28[439:ACDTFT]2.0.CO;2.
  2. Rauhut, Oliver W. M. (2011). "Theropod dinosaurs from the Late Jurassic of Tendaguru (Tanzania)". Special Papers in Palaeontology. 86: 195–239.
  3. Delcourt (2018). "Ceratosaur palaeobiology: New insights on evolution and ecology of the southern rulers". Scientific Reports. 8 (1): 9730. doi:10.1038/s41598-018-28154-x. PMC   6021374 . PMID   29950661.
  4. Hendrickx, Christophe; Mateus, Octávio; Araújo, Ricardo; Choiniere, Jonah (2019-11-24). "The distribution of dental features in non-avian theropod dinosaurs: Taxonomic potential, degree of homoplasy, and major evolutionary trends". Palaeontologia Electronica. 22 (3): 1–110. doi: 10.26879/820 . hdl: 10362/98436 . ISSN   1094-8074.