Einkorn wheat | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Poales |
Family: | Poaceae |
Subfamily: | Pooideae |
Genus: | Triticum |
Species: | T. monococcum |
Binomial name | |
Triticum monococcum | |
Synonyms | |
Triticum monococcum subsp. monococcum |
Wild einkorn | |
---|---|
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Poales |
Family: | Poaceae |
Subfamily: | Pooideae |
Genus: | Triticum |
Species: | T. boeoticum |
Binomial name | |
Triticum boeoticum | |
Synonyms | |
Einkorn wheat (from German Einkorn, literally "single grain") can refer either to a wild species of wheat (Triticum) or to its domesticated form. The wild form is T. boeoticum (syn. T. m. subsp. boeoticum), and the domesticated form is T. monococcum (syn. T. m. subsp. monococcum). Einkorn is a diploid species (2n = 14 chromosomes) of hulled wheat, with tough glumes ('husks') that tightly enclose the grains. The cultivated form is similar to the wild, except that the ear stays intact when ripe [1] and the seeds are larger. The domestic form is known as "petit épeautre" in French, "Einkorn" in German, "einkorn" or "littlespelt" in English, "piccolo farro" in Italian and "escanda menor" in Spanish. [2] The name refers to the fact that each spikelet contains only one grain.
Einkorn wheat was one of the first plants to be domesticated and cultivated. The earliest clear evidence of the domestication of einkorn dates from 10,600 to 9,900 years before present (8650 BCE to 7950 BCE) from Çayönü and Cafer Höyük, two Early Pre-Pottery Neolithic B archaeological sites in southern Turkey. [3] Remnants of einkorn were found with the iceman mummy Ötzi, dated to 3100 BCE. [4]
Einkorn wheat commonly grows wild in the hill country in the northern part of the Fertile Crescent and Anatolia although it has a wider distribution reaching into the Balkans and south to Jordan near the Dead Sea. It is a short variety of wild wheat, usually less than 70 centimetres (28 in) tall and is not very productive of edible seeds.[ citation needed ]
The principal difference between wild einkorn and cultivated einkorn is the method of seed dispersal. In the wild variety the seed head usually shatters and drops the kernels (seeds) of wheat onto the ground. [1] This facilitates a new crop of wheat. In the domestic variety, the seed head remains intact. While such a mutation may occasionally occur in the wild, it is not viable there in the long term: the intact seed head will only drop to the ground when the stalk rots, and the kernels will not scatter but form a tight clump which inhibits germination and makes the mutant seedlings susceptible to disease. But harvesting einkorn with intact seed heads was easier for early human harvesters, who could then manually break apart the seed heads and scatter any kernels not eaten. Over time and through selection, conscious or unconscious, the human preference for intact seed heads created the domestic variety, which also has slightly larger kernels than wild einkorn. Domesticated einkorn thus requires human planting and harvesting for its continuing existence. [5] This process of domestication might have taken only 20 to 200 years with the end product a wheat easier for humans to harvest. [6]
Einkorn wheat is one of the earliest cultivated forms of wheat, alongside emmer wheat (T. dicoccum). Hunter gatherers in the Fertile Crescent may have started harvesting einkorn as early as 30,000 years ago, according to archaeological evidence from Syria. [7] [8] [9] Although gathered from the wild for thousands of years, einkorn wheat was first domesticated approximately 10,000 years BP in the Pre-Pottery Neolithic A (PPNA) or B (PPNB) periods. [10] Evidence from DNA fingerprinting suggests einkorn was first domesticated near Karaca Dağ in southeast Turkey, an area in which a number of PPNB farming villages have been found. [11] One theory by Yuval Noah Harari suggests that the domestication of einkorn was linked to intensive agriculture to support the nearby Göbekli Tepe site. [12]
An important characteristic facilitating the domestication of einkorn and other annual grains is that the plants are largely self-pollinating. Thus, the desirable (for human management) traits of einkorn could be perpetuated at less risk of cross-fertilization with wild plants which might have traits – e.g. smaller seeds, shattering seed heads, [1] as less desirable for human management. [13]
From the northern part of the Fertile Crescent, the cultivation of einkorn wheat spread to the Caucasus, the Balkans, and central Europe. Einkorn wheat was more commonly grown in cooler climates than emmer wheat, the other domesticated wheat. Cultivation of einkorn in the Middle East began to decline in favor of emmer wheat around 2000 BC. Cultivation of einkorn was never extensive in Italy, southern France, and Spain. Einkorn continued to be cultivated in some areas of northern Europe throughout the Middle Ages and until the early part of the 20th century. [14]
Einkorn wheat is low-yielding but can survive on poor, dry, marginal soils where other varieties of wheat will not. It is primarily eaten boiled in whole grains or in porridge. [14] As with other ancient varieties of wheat, Einkorn is grouped with "the covered wheats" as its kernels do not break free from its seed coat (glume) with threshing and it is, therefore, difficult to separate the husk from the seed. [15]
Einkorn is a common food in northern Provence (France). [16] It is also used for bulgur or as animal feed in mountainous areas of France, India, Italy, Morocco, the former Yugoslavia, Turkey, and other countries. [15]
Einkorn contains gluten and has a higher percentage of protein than modern red wheats and is considered more nutritious because it has higher levels of fat, phosphorus, potassium, pyridoxine, and beta-carotene. [15]
Einkorn is the source of many potential introgressions for immunity – Nikolai Vavilov called it an "accumulator of complex immunities." [17] T. monococcum is the source of Sr21, a stem rust resistance gene which has been introgressed into hexaploid worldwide. [18] It is also the source of Yr34, a resistance gene for yellow rust. [19]
The salt-tolerance feature of T. monococcum has been bred into durum wheat. [20]
Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus Triticum ; the most widely grown is common wheat. The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BC. Botanically, the wheat kernel is a caryopsis, a type of fruit.
Tell Abu Hureyra is a prehistoric archaeological site in the Upper Euphrates valley in Syria. The tell was inhabited between 13,300 and 7,800 cal. BP in two main phases: Abu Hureyra 1, dated to the Epipalaeolithic, was a village of sedentary hunter-gatherers; Abu Hureyra 2, dated to the Pre-Pottery Neolithic, was home to some of the world's first farmers. This almost continuous sequence of occupation through the Neolithic Revolution has made Abu Hureyra one of the most important sites in the study of the origins of agriculture.
Emmer wheat or hulled wheat is a type of awned wheat. Emmer is a tetraploid. The domesticated types are Triticum turgidum subsp. dicoccum and T. t. conv. durum. The wild plant is called T. t. subsp. dicoccoides. The principal difference between the wild and the domestic forms is that the ripened seed head of the wild plant shatters and scatters the seed onto the ground, while in the domesticated emmer, the seed head remains intact, thus making it easier for humans to harvest the grain.
Durum wheat, also called pasta wheat or macaroni wheat, is a tetraploid species of wheat. It is the second most cultivated species of wheat after common wheat, although it represents only 5% to 8% of global wheat production. It was developed by artificial selection of the domesticated emmer wheat strains formerly grown in Central Europe and the Near East around 7000 BC, which developed a naked, free-threshing form. Like emmer, durum wheat is awned. It is the predominant wheat that grows in the Middle East.
The Neolithic Revolution, also known as the First Agricultural Revolution, was the wide-scale transition of many human cultures during the Neolithic period in Afro-Eurasia from a lifestyle of hunting and gathering to one of agriculture and settlement, making an increasingly large population possible. These settled communities permitted humans to observe and experiment with plants, learning how they grew and developed. This new knowledge led to the domestication of plants into crops.
Spelt, also known as dinkel wheat or hulled wheat, is a species of wheat that has been cultivated since approximately 5000 BCE.
Khorasan wheat or Oriental wheat is a tetraploid wheat species. The grain is twice the size of modern-day wheat, and has a rich, nutty flavor.
The founder crops or primary domesticates are a group of flowering plants that were domesticated by early farming communities in Southwest Asia and went on to form the basis of agricultural economies across Eurasia. As originally defined by Daniel Zohary and Maria Hopf, they consisted of three cereals, four pulses, and flax. Subsequent research has indicated that many other species could be considered founder crops. These species were amongst the first domesticated plants in the world.
Aegilops is a genus of Eurasian and North American plants in the grass family, Poaceae. They are known generally as goatgrasses. Some species are known as invasive weeds in parts of North America.
Common wheat, also known as bread wheat, is a cultivated wheat species. About 95% of wheat produced worldwide is common wheat; it is the most widely grown of all crops and the cereal with the highest monetary yield.
During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids, have developed under a combination of artificial and natural selection. This diversity has led to much confusion in the naming of wheats. Genetic and morphological characteristics of wheat influence its classification; many common and botanical names of wheat are in current use.
Farro refers to the grains of three wheat species, which are sold dried and cooked in water until soft. It is eaten plain or is often used as an ingredient in salads, soups, and other dishes.
Triticeae is a botanical tribe within the subfamily Pooideae of grasses that includes genera with many domesticated species. Major crop genera found in this tribe include wheat, barley, and rye; crops in other genera include some for human consumption, and others used for animal feed or rangeland protection. Among the world's cultivated species, this tribe has some of the most complex genetic histories. An example is bread wheat, which contains the genomes of three species with only one being a wheat Triticum species. Seed storage proteins in the Triticeae are implicated in various food allergies and intolerances.
Barley, a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikelets and making it much easier to harvest. Its use then spread throughout Eurasia by 2000 BC. Barley prefers relatively low temperatures to grow, and well-drained soil. It is relatively tolerant of drought and soil salinity, but is less winter-hardy than wheat or rye.
A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legumes.
Iraq ed-Dubb, or the Cave of the Bear, is an early Neolithic archeological site 7 km (4.3 mi) northwest of Ajlun in the Jordan Valley, in modern-day Jordan. The settlement existed before 8,000 BCE and experimented with the cultivation of founder crops, side by side with the harvesting of wild cereals. Along with Tell Aswad in Syria, the site shows the earliest reference to domestic hulled barley between 10,000 and 8,800 BCE. The site is located on a forested limestone escarpment above the Wadi el-Yabis in northwest Jordan. An oval-shaped stone structure was excavated along with two burials and a variety of animal and plant remains.
Hordeum spontaneum, commonly known as wild barley or spontaneous barley, is the wild form of the grass in the family Poaceae that gave rise to the cereal barley. Domestication is thought to have occurred on two occasions, first about ten thousand years ago in the Fertile Crescent and again later, several thousand kilometres further east.
Ancient grains is a marketing term used to describe a category of grains and pseudocereals that are purported to have been minimally changed by selective breeding over recent millennia, as opposed to more widespread cereals such as corn, rice and modern varieties of wheat, which are the product of thousands of years of selective breeding. Ancient grains are often marketed as being more nutritious than modern grains, though their health benefits over modern varieties have been disputed by some nutritionists.
Chogha Golan is an aceramic Neolithic archaeological site in the foothills of the Zagros Mountains in Iran, about 200 m (656 ft) from the right bank of the Konjan Cham River. Located in a semi-arid region about 30 km (19 mi) north of Mehran, Chogha Golan is one of the earliest aceramic Neolithic sites found in Iran. The people of Chogha Golan relied primarily on the exploitation of wild plants and hunting. Chogha Golan is notable for the early presence of domesticated emmer wheat, dating to around 9,800 BP as such the archaeobotanical remains from Chogha Golan shows the earliest record of long-term plant management in Iran.