Jesper Hoffmeyer (21 February 1942 – 25 September 2019) [1] was a professor at the University of Copenhagen Institute of Biology, and a leading figure in the emerging field of biosemiotics. He was the president of the International Society for Biosemiotic Studies (ISBS) from 2005 to 2015, co-editor of the journal Biosemiotics and the Springer Book series in Biosemiotics. He authored the books Biosemiotics: An Examination into the Signs of Life and the Life of Signs and Signs of Meaning in the Universe and edited A Legacy for Living Systems: Gregory Bateson as Precursor to Biosemiotics.
Jesper Hoffmeyer was born in Slangerup, Denmark in 1942. He received his Cand. Scient. in biochemistry from the University of Copenhagen in 1967, and from 1967-1968 he was Science Fellow at the Institut de Biochemie Génerale et Comparée of the Collège de France, Paris.
He began his teaching career in 1968 as an assistant professor at the University of Copenhagen's Institute for Biological Chemistry, where he became an associate professor in 1972, and served as the Head of the Institute from 1978-1980. Hoffmeyer was the recipient of the Poul Henningsen Award in 1985, [2] the 1991 Mouton d’Or Award, and named a Thomas Sebeok Fellow by the Semiotic Society of America on the occasion of its 25th annual meeting in 2000.
Hoffmeyer was awarded a doctorate in philosophy (Dr. Phil.) at Aarhus University in 2005, and since 2007 has served as president of the International Society for Biosemiotic Studies.
Since 2009, Hoffmeyer was a professor emeritus at the University of Copenhagen.
Jesper Hoffmeyer's research interests changed considerably through the years. An early engagement in the social and political consequences of his own discipline, biochemistry, led him in the 1970s to take up studies of the technological, ecological and historical dimensions of science. The results of these investigations can be found in his 1982 book Samfundets naturhistorie (The Natural History of Society). [2]
At the same time, Hoffmeyer also embarked upon a parallel track of inquiry, based on a growing awareness that the reductionist tendencies of modern biology and science systematically legitimized and guided the technological horizon towards developments that were inherently damaging to natural systems, including that of human health. This inquiry led him towards questions of theoretical biology and philosophy. Eventually these two research tracks ran together, when, in a collaboration with Claus Emmeche, he initiated an analysis of the concepts of information in both its bio-ontological dimension, as well as in its applied contexts dealing with biological (e.g. genetic) and cultural information, and the then-upcoming 'smart' information and biotechnologies [3]
Hoffmeyer traced much of the development of his thinking during this period to the semiotic traditions of the late 1980s, as well as to such scientific systems theorists as Gregory Bateson, Michael Polanyi, Anthony Wilden, Howard H. Pattee and Peder Voetmann Christiansen. Eventually making the acquaintance of Thomas Sebeok in the United States, Thure von Uexküll in Germany and Kalevi Kull in Estonia, by the beginning of the 1990s, Hoffmeyer had formulated a new programme for a scientific biology that would define life as a signbased phenomenon. Hoffmeyer's first comprehensive essay outlining this of biosemiotics and its implication for a non-dualist understanding of the embodied self was his Signs of Meaning in the Universe (in Danish, En snegl på vejen, 1993). [4]
Beginning in 2001, when the yearly annual international ‘Gatherings in Biosemiotics’ conferences began, Hoffmeyer became a central figure in establishing biosemiotics as a scientific cross-disciplinary field, assembling scientists and humanities scholars to jointly investigate how a semiotic analysis can inform current biological thinking, and how the findings of biology provide general semiotics with a firmer ground for the naturalization of 'meaning'. [5]
In 2005, Hoffmeyer was conferred with a Danish doctoral degree for his treatise Biosemiotics: An Examination into the Signs of Life and the Life of Signs . Translated into an English-language edition in 2008, this work examines the semiotics of living nature, from the origin of life with its self-organizing code-duality in evolution and development, to the complex endosemiosis of living bodies, to the ‘semiotic niches’ in ecosystems, and the species-specific peculiarities of human semiosis, such as language.
Semiotic freedom. The increase in 'depth' of meaning that can be communicated or interpreted. [6] [7]
Semiome. In biosemiotics: the entirety of semiotic tool sets available to the species. [8]
Semiotic scaffolding operates by assuring higher level performance through semiotic interactions inside a living system or between living systems and cue elements in their environment. [9]
Semiotics is the systematic study of sign processes and the communication of meaning. In semiotics, a sign is defined as anything that communicates intentional and unintentional meaning or feelings to the sign's interpreter.
Semiosis, or sign process, is any form of activity, conduct, or process that involves signs, including the production of meaning. A sign is anything that communicates a meaning, that is not the sign itself, to the interpreter of the sign. The meaning can be intentional such as a word uttered with a specific meaning, or unintentional, such as a symptom being a sign of a particular medical condition. Signs can communicate through any of the senses, visual, auditory, tactile, olfactory, or taste.
Biosemiotics is a field of semiotics and biology that studies the prelinguistic meaning-making, biological interpretation processes, production of signs and codes and communication processes in the biological realm.
Zoosemiotics is the semiotic study of the use of signs among animals, more precisely the study of semiosis among animals, i.e. the study of how something comes to function as a sign to some animal. It is the study of animal forms of knowing.
Terrence William Deacon is an American neuroanthropologist. He taught at Harvard for eight years, relocated to Boston University in 1992, and is currently Professor of Anthropology and member of the Cognitive Science Faculty at the University of California, Berkeley.
Jakob Johann Freiherr von Uexküll was a Baltic German biologist who worked in the fields of muscular physiology and animal behaviour studies and was an influence on the cybernetics of life. However, his most notable contribution is the notion of Umwelt, used by semiotician Thomas Sebeok and philosopher Martin Heidegger. His works established biosemiotics as a field of research.
In the semiotic theories of Jakob von Uexküll and Thomas Sebeok, umwelt is the "biological foundations that lie at the very center of the study of both communication and signification in the human [and non-human] animal". The term is usually translated as "self-centered world". Uexküll theorised that organisms can have different umwelten, even though they share the same environment. The term umwelt, together with companion terms Umgebung and Innenwelt, have special relevance for cognitive philosophers, roboticists and cyberneticians because they offer a potential solution to the conundrum of the infinite regress of the Cartesian Theater.

Thomas Albert Sebeok was a Hungarian-born American polymath, semiotician, and linguist. As one of the founders of the biosemiotics field, he studied non-human and cross-species signaling and communication. He is also known for his work in the development of long-time nuclear waste warning messages, in which he worked with the Human Interference Task Force to create methods for keeping the inhabitants of Earth away from buried nuclear waste that will still be hazardous 10,000 or more years in the future.

Sign Systems Studies is a peer-reviewed academic journal on semiotics edited at the Department of Semiotics of the University of Tartu and published by the University of Tartu Press. It is the oldest periodical in the field. It was initially published in Russian and since 1998 in English with Russian and Estonian language abstracts. The journal was established by Juri Lotman as Trudy po Znakovym Sistemam in 1964. Since 1998 it has been edited by Kalevi Kull, Mihhail Lotman, and Peeter Torop. Since 2022, Ott Puumeister leads the editorial team. The journal is available online from the Philosophy Documentation Center, indexed by WoS and Scopus, and starting 2012 also on an open access platform.
The semiosphere is an idea in biosemiotic theory proposing that, contrary to ideas of nature determining sense and experience, the phenomenal world is a creative and logical structure of processes of semiosis where signs operate together to produce sense and experience.
Kalevi Kull is a biosemiotics professor at the University of Tartu, Estonia.
John Deely was an American philosopher and semiotician. He was a professor of philosophy at Saint Vincent College and Seminary in Latrobe, Pennsylvania. Prior to this, he held the Rudman Chair of Graduate Philosophy at the Center for Thomistic Studies, located at the University of St. Thomas (Houston).
Claus Emmeche is a Danish theoretical biologist and philosopher, one of founders of contemporary biosemiotics. He is associate professor at the University of Copenhagen, and is head of the Center for the Philosophy of Nature and Science Studies at the Faculty of Science.
Karl Kuno Thure Freiherr von Uexküll was a German scholar of psychosomatic medicine and biosemiotics. He developed the approach of his father, Jakob von Uexküll, in the study of living systems and applied it in medicine.
Giorgio Prodi was an Italian medical scientist, oncologist and semiotician.
The International Society for Biosemiotic Studies (ISBS) is an academic society for the researchers in semiotic biology. The Society was established in 2005. Its official journal is Biosemiotics, published by Springer and launched in 2008.
The following outline is provided as an overview of and topical guide to semiotics:
Copenhagen–Tartu school of biosemiotics is a loose network of scholars working within the discipline of biosemiotics at the University of Tartu and the University of Copenhagen.
Susan Petrilli is an Italian semiotician, professor of philosophy and theory of languages at the University of Bari, Aldo Moro, Italy, and the seventh Thomas A. Sebeok Fellow of the Semiotic Society of America. She is also International Visiting Research Fellow at the School of Psychology, the University of Adelaide, South Australia.
Paul Cobley is an English eminent semiotician and narratologist.