Lake Tanganyika | |
---|---|
Ikiyaga ca Tanganyika (Kirundi) | |
Coordinates | 6°06′S29°30′E / 6.1°S 29.5°E |
Lake type | Ancient lake, Rift Valley Lake |
Primary inflows | Ruzizi River Malagarasi River Kalambo River |
Primary outflows | Lukuga River |
Catchment area | 231,000 km2 (89,000 sq mi) |
Basin countries | Burundi, DR Congo, Tanzania, and Zambia |
Max. length | 673 km (418 mi) |
Max. width | 72 km (45 mi) |
Surface area | 32,900 km2 (12,700 sq mi) |
Average depth | 570 m (1,870 ft) |
Max. depth | 1,470 m (4,820 ft) |
Water volume | 18,750 km3 (4,500 cu mi) |
Residence time | 5500 years [1] |
Shore length1 | 1,828 km (1,136 mi) |
Surface elevation | 773 m (2,536 ft) [2] |
Settlements | Kigoma, Tanzania Kalemie, the DRC Bujumbura, Burundi Mpulungu, Zambia |
References | [2] |
Official name | Tanganyika |
Designated | 2 February 2007 |
Reference no. | 1671 [3] |
1 Shore length is not a well-defined measure. |
Lake Tanganyika ( /ˌtæŋɡənˈjiːkə,-ɡæn-/ TANG-gən-YEE-kə, -gan-; [4] Kirundi : Ikiyaga ca Tanganyika) is an African Great Lake. [5] It is the second-largest freshwater lake by volume and the second deepest, in both cases after Lake Baikal in Siberia. [6] [7] It is the world's longest freshwater lake. [6] The lake is shared among four countries—Tanzania, the Democratic Republic of the Congo (the DRC), Burundi, and Zambia—with Tanzania (46%) and the DRC (40%) possessing the majority of the lake. It drains into the Congo River system and ultimately into the Atlantic Ocean.[ citation needed ]
Lake Tanganyika is situated within the Albertine Rift, the western branch of the East African Rift, and is confined by the mountainous walls of the valley. It is the largest rift lake in Africa and the second-largest lake by volume in the world. It is the deepest lake in Africa and holds the greatest volume of fresh water on the continent, accounting for 16% of the world's available fresh water. It extends for 676 km (420 mi) in a general north–south direction and averages 50 km (31 mi) in width. The lake covers 32,900 km2 (12,700 sq mi), with a shoreline of 1,828 km (1,136 mi), a mean depth of 570 m (1,870 ft) and a maximum depth of 1,470 m (4,820 ft) (in the northern basin). It holds an estimated 18,750 km3 (4,500 cu mi). [8]
The catchment area of the lake is 231,000 km2 (89,000 sq mi). Two main rivers flow into the lake, as well as numerous smaller rivers and streams (whose lengths are limited by the steep mountains around the lake). The one major outflow is the Lukuga River, which empties into the Congo River drainage. Precipitation and evaporation play a greater role than the rivers. At least 90% of the water influx is from rain falling on the lake's surface and at least 90% of the water loss is from direct evaporation. [9]
The major river flowing into the lake is the Ruzizi River, formed about 10,000 years ago, which enters the north of the lake from Lake Kivu. [10] The Malagarasi River, which is Tanzania's second largest river, enters the east side of Lake Tanganyika. [10] The Malagarasi is older than Lake Tanganyika, and before the lake was formed, it probably was a headwater of the Lualaba River, the main Congo River headstream. [9]
The lake has a complex history of changing flow patterns, due to its high altitude, great depth, slow rate of refill, and mountainous location in a turbulently volcanic area that has undergone climate changes. Apparently, it has rarely in the past had an outflow to the sea. It has been described as "practically endorheic" for this reason. The lake's connection to the sea is dependent on a high water level allowing water to overflow out of the lake through the Lukuga River into the Congo. [10] When not overflowing, the lake's exit into the Lukuga River typically is blocked by sand bars and masses of weed, and instead this river depends on its own tributaries, especially the Niemba River, to maintain a flow. [9]
The lake may also have at times had different inflows and outflows; inward flows from a higher Lake Rukwa, access to Lake Malawi and an exit route to the Nile have all been proposed to have existed at some point in the lake's history. [11]
Lake Tanganyika is an ancient lake, one of only twenty more than a million years old. Its three basins, which in periods with much lower water levels were separate lakes, are of different ages. The central began to form 9–12 million years ago (Mya), the northern 7–8 Mya and the southern 2–4 Mya. [12]
The lake's water is alkaline with a pH around 9 at depths of 0–100 m (0–330 ft). [13] Below this, it is around 8.7, gradually decreasing to 8.3–8.5 in the deepest parts of Tanganyika. [13] A similar pattern can be seen in the electric conductivity, ranging from about 670 μS/cm in the upper part to 690 μS/cm in the deepest. [13]
Surface temperatures generally range from about 24 °C (75 °F) in the southern part of the lake in early August to 28–29 °C (82–84 °F) in the late rainy season in March—April. [14] At depths greater than 400 m (1,300 ft), the temperature is very stable at 23.1–23.4 °C (73.6–74.1 °F). [15] The water has gradually warmed since the 19th century and this has accelerated with global warming since the 1950s. [16]
The lake is stratified and seasonal mixing generally does not extend beyond depths of 150 m (490 ft). [14] The mixing mainly occurs as upwellings in the south and is wind-driven, but to a lesser extent, up- and downwellings also occur elsewhere in the lake. [17] As a consequence of the stratification, the deep sections contain "fossil water". [18] This also means it has no oxygen (it is anoxic) in the deeper parts, essentially limiting fish and other aerobic organisms to the upper part. Some geographical variations are seen in this limit, but it is typically at depths around 100 m (330 ft) in the northern part of the lake and 240–250 m (790–820 ft) in the south. [19] [20] The oxygen-devoid deepest sections contain high levels of toxic hydrogen sulphide and are essentially lifeless, [6] except for bacteria. [13] [21]
Lake Tanganyika and its associated wetlands are home to Nile crocodiles (including famous giant Gustave), Zambian hinged terrapins, serrated hinged terrapins, and pan hinged terrapins (last species not in the lake itself, but in adjacent lagoons). [22] Storm's water cobra, a threatened subspecies of banded water cobra that feeds mainly on fish, is only found in Lake Tanganyika, where it prefers rocky shores. [22] [23]
The lake holds at least 250 species of cichlid fish [27] and undescribed species remain. [28] Almost all (98%) of the Tanganyika cichlids are endemic to the lake and it is thus an important biological resource for the study of speciation in evolution. [29] [30] Some of the endemics do occur slightly into the upper Lukuga River, Lake Tanganyika's outflow, but further spread into the Congo River basin is prevented by physics (Lukuga has fast-flowing sections with many rapids and waterfalls) and chemistry (Tanganyika's water is alkaline, while the Congo's generally is acidic). [9] The cichlids of the African Great Lakes, including Tanganyika, represent the most diverse extent of adaptive radiation in vertebrates. [31]
Although Tanganyika has far fewer cichlid species than Lakes Malawi and Victoria which both have experienced relatively recent explosive species radiations (resulting in many closely related species), [32] its cichlids are the most morphologically and genetically diverse. [31] [33] This is linked to the high age of Tanganyika, as it is far older than the other lakes. [34] Tanganyika has the largest number of endemic cichlid genera of all African lakes. [31] All Tanganyika cichlids are in the subfamily Pseudocrenilabrinae. Of the 10 tribes in this subfamily, half are largely or entirely restricted to the lake (Cyprichromini, Ectodini, Lamprologini, Limnochromini and Tropheini) and another three have species in the lake (Haplochromini, Tilapiini and Tylochromini). [35] Others have proposed splitting the Tanganyika cichlids into as many as 12–16 tribes (in addition to previous mentioned, Bathybatini, Benthochromini, Boulengerochromini, Cyphotilapiini, Eretmodini, Greenwoodochromini, Perissodini and Trematocarini). [31]
Most Tanganyika cichlids live along the shoreline down to a depth of 100 m (330 ft), but some deep-water species regularly descend to 200 m (660 ft). [36] Trematocara species have exceptionally been found at more than 300 m (980 ft), which is deeper than any other cichlid in the world. [37] Some of the deep-water cichlids (e.g., Bathybates , Gnathochromis , Hemibates and Xenochromis ) have been caught in places virtually devoid of oxygen, but how they are able to survive there is unclear. [20] Tanganyika cichlids are generally benthic (found at or near the bottom) and/or coastal. [38] No Tanganyika cichlids are truly pelagic and offshore, except for some of the piscivorous Bathybates. [36] Two of these, B. fasciatus and B. leo , mainly feed on Tanganyika sardines. [36] [20] Tanganyika cichlids differ extensively in ecology and include species that are herbivores, detritivores, planktivores, insectivores, molluscivores, scavengers, scale-eaters and piscivores. [28] These dietary specializations, however, have been shown to be flexible. That is, many species of Tanganyikan cichlid with specialized diets showed opportunistic, episodic exploitation of Stolothrissa tanganicae and Limnothrissa miodon when prey concentrations were unusually high. [39] Their breeding behavior fall into two main groups, the substrate spawners (often in caves or rock crevices) and the mouthbrooders. [40] Among the endemic species are two of the world's smallest cichlids, Neolamprologus multifasciatus and N. similis (both shell dwellers) at up to 4–5 cm (1.6–2.0 in), [41] [42] and one of the largest, the giant cichlid (Boulengerochromis microlepis) at up to 90 cm (3.0 ft). [28] [43]
Many cichlids from Lake Tanganyika, such as species from the genera Altolamprologus , Cyprichromis , Eretmodus , Julidochromis , Lamprologus , Neolamprologus , Tropheus and Xenotilapia , are popular aquarium fish due to their bright colors and patterns, and interesting behaviors. [40] Recreating a Lake Tanganyika biotope to host those cichlids in a habitat similar to their natural environment is also popular in the aquarium hobby. [40] [44]
Lake Tanganyika is home to more than 80 species of non-cichlid fish and about 60% of these are endemic. [19] [27] [63] [64]
The open waters of the pelagic zone are dominated by four non-cichlid species: Two species of "Tanganyika sardine" ( Limnothrissa miodon and Stolothrissa tanganicae ) form the largest biomass of fish in this zone, and they are important prey for the forktail lates (Lates microlepis) and sleek lates (L. stappersii). [38] Two additional lates are found in the lake, the Tanganyika lates (L. angustifrons) and bigeye lates (L. mariae), but both these are primarily benthic hunters, although they also may move into open waters. [38] The four lates, all endemic to Tanganyika, have been overfished and larger individuals are rare today. [38]
Among the more unusual fish in the lake are the endemic, facultatively brood parasitic "cuckoo catfish", including at least Synodontis grandiops [65] and S. multipunctatus . [19] [40] A number of others are very similar (e.g., S. lucipinnis and S. petricola ) and have often been confused; it is unclear if they have a similar behavior. [19] [66] The facultative brood parasites often lay their eggs synchronously with mouthbroding cichlids. The cichlid pick up the eggs in their mouth as if they were their own. Once the catfish eggs hatch the young eat the cichlid eggs. [19] [40] Six catfish genera are entirely restricted to the lake basin: Bathybagrus , Dinotopterus , Lophiobagrus , Phyllonemus , Pseudotanganikallabes and Tanganikallabes . [53] [67] Although not endemic on a genus level, six species of Chrysichthys catfish are only found in the Tanganyika basin where they live both in shallow and relatively deep waters; [53] in the latter habitat they are the primary predators and scavengers. [20] A unique evolutionary radiation in the lake is the 15 species of Mastacembelus spiny eels, all but one endemic to its basin. [63] [68] Although other African Great Lakes have Synodontis catfish, endemic catfish genera and Mastacembelus spiny eels, the relatively high diversity is unique to Tanganyika, which likely is related to its old age. [68]
Among the non-endemic fish, some are widespread African species but several are only shared with the Malagarasi and Congo River basins, such as the Congo bichir (Polypterus congicus), goliath tigerfish (Hydrocynus goliath), Citharinus citharus, six-banded distichodus (Distichodus sexfasciatus) and mbu puffer (Tetraodon mbu). [53]
A total of 83 freshwater snail species (65 endemic) and 11 bivalve species (8 endemic) are known from the lake. [69] Among the endemic bivalves are three monotypic genera: Grandidieria burtoni , Pseudospatha tanganyicensis and Brazzaea anceyi . [69] Many of the snails are unusual for species living in freshwater in having noticeably thickened shells and/or distinct sculpture, features more commonly seen in marine snails. They are referred to as thalassoids, which can be translated to "marine-like". [70] All the Tanganyika thalassoids, which are part of Prosobranchia, are endemic to the lake. [70] Initially they were believed to be related to similar marine snails, but they are now known to be unrelated. Their appearance is now believed to be the result of the highly diverse habitats in Lake Tanganyika and evolutionary pressure from snail-eating fish and, in particular, Platythelphusa crabs. [27] [70] [71] A total of 17 freshwater snail genera are endemic to the lake, such as Hirthia , Lavigeria , Paramelania , Reymondia , Spekia , Stanleya , Tanganyicia and Tiphobia . [70] There are about 30 species of non-thalassoid snails in the lake, but only five of these are endemic, including Ferrissia tanganyicensis and Neothauma tanganyicense . [70] The latter is the largest Tanganyika snail and its shell is often used by small shell-dwelling cichlids. [72]
Crustaceans are also highly diverse in Tanganyika with more than 200 species, of which more than half are endemic. [27] They include 10 species of freshwater crabs (9 Platythelphusa and Potamonautes platynotus ; all endemic), [73] at least 11 species of small atyid shrimp ( Atyella , Caridella and Limnocaridina ), [74] an endemic palaemonid shrimp ( Macrobrachium moorei), [75] about 100 ostracods, [76] including many endemics, [77] [78] and several copepods. [79] Among these, Limnocaridina iridinae lives inside the mantle cavity of the unionid mussel Pleiodon spekei , making it one of only two known commensal species of freshwater shrimp (the other is the sponge-living Caridina spongicola from Lake Towuti, Indonesia). [80] [81]
Among Rift Valley lakes, Lake Tanganyika far surpasses all others in terms of crustacean and freshwater snail richness (both in total number of species and number of endemics). [82] For example, the only other Rift Valley lake with endemic freshwater crabs are Lake Kivu and Lake Victoria with two species each. [83] [84]
The diversity of other invertebrate groups in Lake Tanganyika is often not well-known, but there are at least 20 described species of leeches (12 endemics), [85] 9 sponges (7 endemic), 6 bryozoa (2 endemic), 11 flatworms (7 endemic), 20 nematodes (7 endemic), 28 annelids (17 endemic) [27] and the small hydrozoan jellyfish Limnocnida tanganyicae. [86]
Lake Tanganyika supports a major fishery, which, depending on source, provides 25–40% [87] or c. 60% of the animal protein in the diet of the people living in the region. [16] [88]
Lake Tanganyika fish can be found exported throughout East Africa. Major commercial fishing began in the mid-1950s and has, together with global warming, had a heavy impact on the fish populations, causing significant declines. [16] [88] [15] In 2016, it was estimated that the total catch was up to 200,000 tonnes. [16]
It is thought that early Homo sapiens were making an impact on the region during the Stone Age. The time period of the Middle Stone Age to Late Stone Age is described as an age of advanced hunter-gatherers. [89]
There are many methods in which the native people of the area were fishing. Most of them included using a lantern as a lure for fish that are attracted to light. There were three basic forms. One called Lusenga which is a wide net used by one person from a canoe. The second one is using a lift net. This was done by dropping a net deep below the boat using two parallel canoes and then simultaneously pulling it up. The third is called Chiromila which consisted of three canoes. One canoe was stationary with a lantern while another canoe holds one end of the net and the other circles the stationary one to meet up with the net. [90]
The first known Westerners to find the lake were the British explorers Richard Burton and John Speke, in 1858. They located it while searching for the source of the Nile River. Speke continued and found the actual source, Lake Victoria. Later David Livingstone passed by the lake. He noted the name "Liemba" for its southern part, a word probably from the Fipa language. [91] Tanganyika means "stars" in the Luvale language. [92] : 523
The lake was the scene of Battle for Lake Tanganyika during World War I. With the aid of the Graf Goetzen , the Germans had complete control of the lake in the early stages of the war. The ship was used both to ferry cargo and personnel across the lake, and as a base from which to launch surprise attacks on Allied troops. [93] It therefore became essential for the Allied forces to gain control of the lake themselves. Under the command of Lieutenant Commander Geoffrey Spicer-Simson the British Royal Navy achieved the monumental task of bringing two armed motor boats HMS Mimi and HMS Toutou from England to the lake by rail, road and river to Albertville (since renamed Kalemie in 1971) on the western shore of Lake Tanganyika. The two boats waited until December 1915, and mounted a surprise attack on the Germans, with the capture of the gunboat Kingani . Another German vessel, the Hedwig , was sunk in February 1916, leaving the Götzen as the only German vessel remaining to control the lake. [93] In order to avoid his prize ship falling into Allied hands, Zimmer scuttled the vessel on July 26, 1916. The vessel was later raised in 1924 and renamed MV Liemba. [93]
Cichlids are fish from the family Cichlidae in the order Cichliformes. Traditionally Cichlids were classed in a suborder, the Labroidei, along with the wrasses (Labridae), in the order Perciformes, but molecular studies have contradicted this grouping. On the basis of fossil evidence, it first appeared in Argentina during the Early Eocene epoch, about 48.6 million years ago; however, molecular clock estimates have placed the family's origin as far back as 67 million years ago, during the late Cretaceous period. The closest living relative of cichlids is probably the convict blenny, and both families are classified in the 5th edition of Fishes of the World as the two families in the Cichliformes, part of the subseries Ovalentaria. This family is large, diverse, and widely dispersed. At least 1,650 species have been scientifically described, making it one of the largest vertebrate families. New species are discovered annually, and many species remain undescribed. The actual number of species is therefore unknown, with estimates varying between 2,000 and 3,000.
Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, is an African Great Lake and the southernmost lake in the East African Rift system, located between Malawi, Mozambique and Tanzania.
Lake Victoria is one of the African Great Lakes. With a surface area of approximately 59,947 km2 (23,146 sq mi), Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest fresh water lake by surface area after Lake Superior in North America. In terms of volume, Lake Victoria is the world's ninth-largest continental lake, containing about 2,424 km3 (1.965×109 acre⋅ft) of water. Lake Victoria occupies a shallow depression in Africa. The lake has an average depth of 40 m (130 ft) and a maximum depth of 80–81 m (262–266 ft). Its catchment area covers 169,858 km2 (65,583 sq mi). The lake has a shoreline of 7,142 km (4,438 mi) when digitized at the 1:25,000 level, with islands constituting 3.7% of this length.
Lake Kivu is one of the African Great Lakes. It lies on the border between the Democratic Republic of the Congo and Rwanda, and is in the Albertine Rift, the western branch of the East African Rift. Lake Kivu empties into the Ruzizi River, which flows southwards into Lake Tanganyika. In 1894, German explorer and officer Gustav Adolf von Götzen was the first European to discover the lake.
Lamprologus ocellatus is a species of shell dwelling cichlid endemic to Lake Tanganyika. It is a popular aquarium fish due to its small size, appearance, and intelligence.
Synodontis multipunctatus, also known as the cuckoo catfish, cuckoo squeaker, or multipunk, is a small catfish from Lake Tanganyika, one of the lakes in the Great Rift Valley system in Africa. It is a brood parasite upon mouthbrooding cichlids. This species grows to a length of 27.5 centimetres (10.8 in) TL. This species is a minor component of local commercial fisheries.
Synodontis is the largest genus of mochokid catfishes. It is the biggest genus within the 10 genera and 190 different species in the family Mochokidae. Synodontis has over 131 different species within the genus. Synodontis are also known as squeakers, due to their ability to make stridulatory sounds through their pectoral fin spines when handled or disturbed. Synodontis make a sound that sounds like squeaking by rubbing their spines together. They do this when they have been frightened or when they become angry. Synodontis may also squeak when they are taken out of the water. These catfish are small- to medium-sized fish with many species exhibiting attractive spotted markings. Some species are also known for naturally swimming belly-up, earning the name upside-down catfish. Some of these species are Synodontis contractus and Synodontis nigriventris. While some of these species are known to swim upside down, another species, Synodontis multipunctatus, is a brood parasitic cuckoo catfish,there are two other species Synodontis petricola and Synodontis grandiops are also called brood parasitic cuckoo catfish.
Lepidophagy is a specialised feeding behaviour in fish that involves eating the scales of other fish. Lepidophagy is widespread, having evolved independently in at least five freshwater families and seven marine families. A related feeding behavior among fish is pterygophagy: feeding on the fins of other fish.
Altolamprologus compressiceps is a species of fish in the family Cichlidae. It is endemic to the shallow rocky areas of Lake Tanganyika. It is not considered threatened by the IUCN.
The convict julie is a cichlid species in the subfamily Pseudocrenilabrinae family endemic to Lake Tanganyika. Hence it is found in Burundi, the Democratic Republic of the Congo, Tanzania, and Zambia. The fish is named after Charles Tate Regan.
Perissodus is a genus of cichlids endemic to Lake Tanganyika in Africa. They feed on scales.
Plecodus elaviae is a species of cichlid endemic to Lake Tanganyika. This schooling species is a scale-eater, plucking scales from other fishes. Both parents care for the offspring in this mouthbrooder. This fish can reach a length of 32 centimetres (13 in) TL.
Plecodus is a genus of cichlids endemic to Lake Tanganyika in Africa. They feed on scales.
The spotfin goby cichlid is an African species of cichlid endemic to Lake Tanganyika where it is only known from the northern end of the lake. They live amongst pebbles in the surf-zone. This species can reach a length of 7 centimetres (2.8 in) TL. This species can also be found in the aquarium trade. Although presently considered the only species in the genus, another undescribed species is known from the Lukuga River.
Neothauma is a genus of freshwater snail with a gill and an operculum, an aquatic gastropod mollusc in the subfamily Bellamyinae of the family Viviparidae.
Synodontis petricola, known as the cuckoo catfish, or the pygmy leopard catfish, is a species of upside-down catfish endemic to Burundi, Zambia, the Democratic Republic of the Congo, and Tanzania where it is only known from Lake Tanganyika. It was first described by Belgian ichthyologist Hubert Matthes in 1959. The species name "petricola" is derived from a combination of the Latin petra, meaning stone or rock, and the Latin cola, meaning inhabitant. This refers to the rocky environment where this species is found.
Eretmodini is a tribe of African cichlids. It contains five species of freshwater fish endemic to Lake Tanganyika. They are small fish with reduced swim bladders that are found near the bottom in the turbulent, coastal surf zone. They are mouthbrooders.
Perissodini is a tribe of African cichlids, containing three genera of freshwater fish found only in Lake Tanganyika.
Synodontis dorsomaculatus is a species of upside-down catfish that is endemic to the Democratic Republic of the Congo where it occurs in the Lukusi and upper Lualaba Rivers and the Upemba lakes. It was first described by Max Poll in 1971. The original specimens were obtained in Kadia, Kisale, Katanga, in what is now the Democratic Republic of the Congo. The species name dorsomaculatus is derived from the word dorso, meaning "back", and the word maculatus, meaning "spotted", a reference to the dark round spots on the fish's back.
Mastacembelus ellipsifer is a species of spiny eel that is endemic to Lake Tanganyika in Africa and sometimes kept in aquariums. Although sometimes called the Tanganyikan spiny eel, it is only one of fifteen spiny eel species in the Tanganyikan basin.
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