Lichida Temporal range: | |
---|---|
Terataspis grandis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Trilobita |
Order: | † Lichida Moore, 1959 |
Families [1] | |
Lichida is an order of typically spiny trilobite that lived from the Furongian to the Devonian period. [2] These trilobites usually have 8–13 thoracic segments. [1] Their exoskeletons often have a grainy texture or have wart or spine-like tubercles. Some species are extraordinarily spiny, having spiny thoracic segments that are as long or longer than the entire body, from cephalon (head) to pygidium (tail). The sections of the pygidia are leaf-like in shape and also typically end in spines.
The order is divided into two families, Lichidae, and Lichakephalidae. [1] Some experts group the families of the closely related order Odontopleurida within Lichida, too, whereupon the family is then divided into three superfamilies, Dameselloidea, containing the family Damesellidae, Lichoidea, containing the families Lichidae and Lichakephalidae, and Odontopleuroidea, containing the family Odontopleuridae.
Lichids are some of the largest trilobites, with the second and third largest trilobites ( Uralichas hispanicus , and Teratapsis grandis ) being in the order [3]
As mentioned earlier, the order Lichida is divided into two families.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
The pygidium is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.
Dalmanites is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian.
Paradoxides is a genus of large to very large trilobite found throughout the world during the Middle Cambrian period. One record-breaking specimen of Paradoxides davidis, described by John William Salter in 1863, is 37 cm (15 in). The cephalon was semicircular with free cheeks ending in long, narrow, recurved spines. Eyes were crescent shaped providing an almost 360° view, but only in the horizontal plane. Its elongate thorax was composed of 19–21 segments and adorned with longish, recurved pleural spines. Its pygidium was comparatively small. Paradoxides is a characteristic Middle-Cambrian trilobite of the 'Atlantic' (Avalonian) fauna. Avalonian rocks were deposited near a small continent called Avalonia in the Paleozoic Iapetus Ocean. Avalonian beds are now in a narrow strip along the East Coast of North America, and in Europe.
Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian of the East Gondwana supercontinent, in what are today South-Australia and Antarctica.
Balcoracania dailyi is a small trilobite of the family Emuellidae. Its fossils have been found in south Australia and Antarctica.
Olenellina is a suborder of the order Redlichiida of trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. Olenellina are arguably the earliest trilobites in the fossil record as members of Redlichiina, although Ptychopariida and Eodiscina follow soon after. The suborder died out when the Lower Cambrian passed into the Middle Cambrian, at the end of the stage called Toyonian. A feature uniting the Olenellina is the lack of rupture lines in the headshield, which in other trilobites assist the periodic moulting, associated with arthropod growth. Some derived trilobites have lost facial sutures again, but all of these are blind, while all Olenellina have eyes.
Megalograptus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Megalograptus have been recovered in deposits of Katian age in North America. The genus contains five species: M. alveolatus, M. ohioensis, M. shideleri, M. welchi and M. williamsae, all based on fossil material found in the United States. Fossils unassigned to any particular species have also been found in Canada. The generic name translates to "great writing" and originates from the mistaken original belief that Megalograptus was a type of graptolite, often given names ending with -graptus.
Terataspis is a comparatively huge, 60 centimetre long lichid trilobite genus from the Early Devonian, about 397 million years ago. It lived in a shallow sea in what is now New York State and Ontario. No whole specimens have been found, only disarticulated fragments of its exoskeleton, but enough fragments have been found to allow researchers to form reconstructions of the whole animal. The genus only contains one species, T. grandis.
Ronald Pearson Tripp FRSE was a British palaeontologist specializing in trilobites. He was self-taught in palaeontology and became an authority on the taxonomy of the trilobite order Lichida and the trilobite family Encrinuridae.
Acimetopus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi (1943), Order Agnostida Salter (1864). It lived during the Botomian stage. = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.
Nevadella is an extinct genus of trilobites, fossil marine arthropods, with species of average size. It lived during the late Atdabanian stage, which lasted from 530 to 524 million years ago during the early part of the Cambrian Period.
Fallotaspis is a genus of redlichiid trilobites found in Early Cambrian-aged strata of the United States and Morocco.
Odontopleurida is an order of very spinose trilobites closely related to the trilobites of the order Lichida. Some experts group the Odontopleurid families, Odontopleuridae and Damesellidae, within Lichida. Odontopleurids tend to have convex, bar-shaped cephalons, and lobed, knob-shaped glabella that extend to, or almost to the anterior margin. Many, if not almost all odontopleurids have long spines that are derived either from the margins of the exoskeleton, or from granular or tubercular ornamentation, or both. Many odontopleurids are so spinose so as to be described as having "spines on (their) spines." Odontopleurids have 8 to 13 thoracic segments, with Odontopleuridae odontopleurids having no more than 10, and Damesellidae odontopleurids having no more than 13. The pygidium tends to be very small, and invariably has long spines emanating from it in all known genera.
Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica and Western Laurentia, and also in Morocco.
Peachella is an extinct genus of trilobites, fossil marine arthropods, with species of average size. It lived during the Toyonian stage, 516 to 513 million years ago, in what is today the southwestern United States. It can easily be distinguished from other trilobites by its club-like genal spines.
Eopeachella is an extinct genus of trilobites, fossil marine arthropods, with species of probably small size, entire, articulate specimens have not yet been found. It lived during the Toyonian stage, in what is today the South-Western United States. Eopeachella angustispina is the only known species in this genus.
Emigrantia is an extinct genus of trilobites, fossil marine arthropods, of small to average size. It lived during the Toyonian stage, in what is today the South-Western United States. Emigrantia can easily be distinguished from other trilobites by the sturdy but not inflated genal spines, that are attached at midlength of the cephalon, in combination with effaced features of the raised axial area of the head shield.
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.
Cyclopygidae is a family of asaphid trilobites from the Ordovician. Cyclopygids had an extratropical distribution, and there is evidence that they lived in darker parts of the water column. Cyclopygids are characterized by enlarged eyes, with a wide angle of view, both horizontal and vertical, reminiscent of the eyes of dragonflies. These typically touch the glabella directly on the side. Cyclopygids all lack genal spines, but Symphysops carries a forward directed frontal spine on the glabella. It is presumed that at least the members of the genus Pricyclopyge swam upside down and had bioluminescent organs on the third thorax segment. Cyclopygids had between 7 and 5 thorax segments, a wide and stout axis, and short side lobes.