Mesozoic

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Mesozoic Era
251.902–66 million years ago
Key events in the Mesozoic
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Phanerozoic
An approximate timescale of key Mesozoic events.
Axis scale: millions of years ago.

The Mesozoic Era ( /ˌmɛz.əˈz.ɪk,ˌmɛz.-,ˌmɛs-,ˌm.zə-,-z-,ˌm.sə-,-s-/ mez-ə-ZOH-ik, mez-oh-, mess-, mee-zə-, -zoh-, mee-sə-, -soh-) [1] [2] is an interval of geological time from about 252 to 66 million years ago. It is also called the Age of Reptiles and the Age of Conifers. [3]

Contents

The Mesozoic ("middle life") is one of three geologic eras of the Phanerozoic Eon, preceded by the Paleozoic ("ancient life") and succeeded by the Cenozoic ("new life"). The era is subdivided into three major periods: the Triassic, Jurassic, and Cretaceous, which are further subdivided into a number of epochs and stages.

The era began in the wake of the Permian–Triassic extinction event, the largest well-documented mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event, another mass extinction whose victims included the non-avian dinosaurs. The Mesozoic was a time of significant tectonic, climate, and evolutionary activity. The era witnessed the gradual rifting of the supercontinent Pangaea into separate landmasses that would move into their current positions during the next era. The climate of the Mesozoic was varied, alternating between warming and cooling periods. Overall, however, the Earth was hotter than it is today. Dinosaurs first appeared in the Mid-Triassic, and became the dominant terrestrial vertebrates in the Late Triassic or Early Jurassic, occupying this position for about 150 or 135 million years until their demise at the end of the Cretaceous. Birds first appeared in the Jurassic (however, true toothless birds appeared first in the Cretaceous), having evolved from a branch of theropod dinosaurs. The first mammals also appeared during the Mesozoic, but would remain small—less than 15 kg (33 lb)—until the Cenozoic. The flowering plants (angiosperms) arose in the Triassic or Jurassic and came to prominence in the late Cretaceous when they replaced the conifers and other gymnosperms as the dominant trees.

Naming

The phrase "Age of Reptiles" was introduced by the 19th century paleontologist Gideon Mantell who viewed it as dominated by diapsids such as Iguanodon , Megalosaurus , Plesiosaurus , and Pterodactylus .

Mesozoic means "middle life", deriving from the Greek prefix meso-/μεσο- for "between" and zōon/ζῷον meaning "animal" or "living being". The name "Mesozoic" was proposed in 1840 by the British geologist John Phillips (1800–1874). [4] [5]

The Mesozoic era was originally described as the "secondary" era, following the primary or Paleozoic, and preceding the Tertiary. [6]

Geologic periods

Following the Paleozoic, the Mesozoic extended roughly 186 million years, from 251.902 to 66 million years ago when the Cenozoic Era began. This time frame is separated into three geologic periods. From oldest to youngest:

The lower boundary of the Mesozoic is set by the Permian–Triassic extinction event, during which approximately 90% to 96% of marine species and 70% of terrestrial vertebrates became extinct. [7] It is also known as the "Great Dying" because it is considered the largest mass extinction in the Earth's history. The upper boundary of the Mesozoic is set at the Cretaceous–Paleogene extinction event (or K–Pg extinction event [8] ), which may have been caused by an asteroid impactor that created Chicxulub Crater on the Yucatán Peninsula. Towards the Late Cretaceous, large volcanic eruptions are also believed to have contributed to the Cretaceous–Paleogene extinction event. Approximately 50% of all genera became extinct, including all of the non-avian dinosaurs.

Triassic

The Triassic ranges roughly from 252 million to 201 million years ago, preceding the Jurassic Period. The period is bracketed between the Permian–Triassic extinction event and the Triassic–Jurassic extinction event, two of the "big five", and it is divided into three major epochs: Early, Middle, and Late Triassic. [9]

The Early Triassic, about 252 to 247 million years ago, was dominated by deserts in the interior of the Pangaea supercontinent. The Earth had just witnessed a massive die-off in which 95% of all life became extinct, and the most common vertebrate life on land were Lystrosaurus , labyrinthodonts, and Euparkeria along with many other creatures that managed to survive the Permian extinction. Temnospondyls evolved during this time and would be the dominant predator for much of the Triassic. [10]

Plateosaurus (a prosauropod) Sellosaurus.jpg
Plateosaurus (a prosauropod)

The Middle Triassic, from 247 to 237 million years ago, featured the beginnings of the breakup of Pangaea and the opening of the Tethys Ocean. Ecosystems had recovered from the Permian extinction. Algae, sponge, corals, and crustaceans all had recovered, and new aquatic reptiles evolved, such as ichthyosaurs and nothosaurs. On land, pine forests flourished, as did groups of insects like mosquitoes and fruit flies. Reptiles began to get bigger and bigger, and the first crocodilians and dinosaurs evolved, which sparked competition with the large amphibians that had previously ruled the freshwater world, respectively mammal-like reptiles on land. [11]

Following the bloom of the Middle Triassic, the Late Triassic, from 237 to 201 million years ago, featured frequent heat spells and moderate precipitation (10–20 inches per year). The recent warming led to a boom of dinosaurian evolution on land as those one began to separate from each other (Nyasasaurus from 243 to 210 million years ago, approximately 235–30 ma, some of them separated into Sauropodomorphs, Theropods and Herrerasaurids), as well as first pterosaurs. During the Late Triassic, some advanced cynodonts gave rise to the first Mammaliaformes. All this climatic change, however, resulted in a large die-out known as the Triassic–Jurassic extinction event, in which many archosaurs (excluding pterosaurs, dinosaurs and crocodylomorphs), most synapsids, and almost all large amphibians became extinct, as well as 34% of marine life, in the Earth's fourth mass extinction event. The cause is debatable; [12] [13] flood basalt eruptions at the Central Atlantic magmatic province is cited as one possible cause.

Jurassic

Sericipterus Sericipterus NT.jpg
Sericipterus

The Jurassic ranges from 200 million years to 145 million years ago and features three major epochs: The Early Jurassic, the Middle Jurassic, and the Late Jurassic. [14]

The Early Jurassic spans from 200 to 175 million years ago. [14] The climate was tropical, much more humid than the Triassic. In the oceans, plesiosaurs, ichthyosaurs and ammonites were abundant. On land, dinosaurs and other archosaurs staked their claim as the dominant race, with theropods such as Dilophosaurus at the top of the food chain. The first true crocodiles evolved, pushing the large amphibians to near extinction. All-in-all, archosaurs rose to rule the world. Meanwhile, the first true mammals evolved, remaining relatively small but spreading widely; the Jurassic Castorocauda , for example, had adaptations for swimming, digging and catching fish. Fruitafossor , from the late Jurassic period about 150 million years ago, was about the size of a chipmunk, and its teeth, forelimbs and back suggest that it dug open the nests of social insects (probably termites, as ants had not yet appeared). The first multituberculates like Rugosodon evolved, while volaticotherians took to the skies.

The Middle Jurassic spans from 175 to 163 million years ago. [14] During this epoch, dinosaurs flourished as huge herds of sauropods, such as Brachiosaurus and Diplodocus , filled the fern prairies, chased by many new predators such as Allosaurus . Conifer forests made up a large portion of the forests. In the oceans, plesiosaurs were quite common, and ichthyosaurs flourished. This epoch was the peak of the reptiles. [15]

Stegosaurus Stegosaurus BW.jpg
Stegosaurus

The Late Jurassic spans from 163 to 145 million years ago. [14] During this epoch, the first avialans, like Archaeopteryx , evolved from small coelurosaurian dinosaurs. The increase in sea levels opened up the Atlantic seaway, which has grown continually larger until today. The divided landmasses gave opportunity for the diversification of new dinosaurs.

Cretaceous

The Cretaceous is the longest period of the Mesozoic, but has only two epochs: Early and Late Cretaceous. [16]

Tylosaurus (a mosasaur) hunting Xiphactinus Knight Tylosaurus.jpg
Tylosaurus (a mosasaur) hunting Xiphactinus

The Early Cretaceous spans from 145 to 100 million years ago. [16] The Early Cretaceous saw the expansion of seaways, and as a result, the decline and extinction of sauropods (except in South America). Some island-hopping dinosaurs, like Eustreptospondylus , evolved to cope with the coastal shallows and small islands of ancient Europe. Other dinosaurs rose up to fill the empty space that the Jurassic-Cretaceous extinction left behind, such as Carcharodontosaurus and Spinosaurus . Of the most successful was the Iguanodon , which spread to every continent. Seasons came back into effect and the poles got seasonally colder, but some dinosaurs still inhabited the polar forests year round, such as Leaellynasaura and Muttaburrasaurus . The poles were too cold for crocodiles, and became the last stronghold for large amphibians like Koolasuchus . Pterosaurs got larger as genera like Tapejara and Ornithocheirus evolved. Mammals continued to expand their range: eutriconodonts produced fairly large, wolverine-like predators like Repenomamus and Gobiconodon , early therians began to expand into metatherians and eutherians, and cimolodont multituberculates went on to become common in the fossil record.

The Late Cretaceous spans from 100 to 66 million years ago. The Late Cretaceous featured a cooling trend that would continue in the Cenozoic era. Eventually, tropics were restricted to the equator and areas beyond the tropic lines experienced extreme seasonal changes in weather. Dinosaurs still thrived, as new taxa such as Tyrannosaurus , Ankylosaurus , Triceratops and hadrosaurs dominated the food web. In the oceans, mosasaurs ruled, filling the role of the ichthyosaurs, which, after declining, had disappeared in the Cenomanian-Turonian boundary event. Though pliosaurs had gone extinct in the same event, long-necked plesiosaurs such as Elasmosaurus continued to thrive. Flowering plants, possibly appearing as far back as the Triassic, became truly dominant for the first time. Pterosaurs in the Late Cretaceous declined for poorly understood reasons, though this might be due to tendencies of the fossil record, as their diversity seems to be much higher than previously thought. Birds became increasingly common and diversified into a variety of enantiornithe and ornithurine forms. Though mostly small, marine hesperornithes became relatively large and flightless, adapted to life in the open sea. Metatherians and primitive eutherian also became common and even produced large and specialised genera like Didelphodon and Schowalteria . Still, the dominant mammals were multituberculates, cimolodonts in the north and gondwanatheres in the south. At the end of the Cretaceous, the Deccan traps and other volcanic eruptions were poisoning the atmosphere. As this continued, it is thought that a large meteor smashed into earth 66 million years ago, creating the Chicxulub Crater in an event known as the K-Pg Extinction (formerly K-T), the fifth and most recent mass extinction event, in which 75% of life became extinct, including all non-avian dinosaurs. [17] Everything over 10 kilograms became extinct. The age of the dinosaurs was over. [18] [19]

Paleogeography and tectonics

Breakup of Pangaea Pangea animation 03.gif
Breakup of Pangaea

Compared to the vigorous convergent plate mountain-building of the late Paleozoic, Mesozoic tectonic deformation was comparatively mild. The sole major Mesozoic orogeny occurred in what is now the Arctic, creating the Innuitian orogeny, the Brooks Range, the Verkhoyansk and Cherskiy Ranges in Siberia, and the Khingan Mountains in Manchuria.

This orogeny was related to the opening of the Arctic Ocean and subduction of the North China and Siberian cratons under the Pacific Ocean. [20] In contrast, the era featured the dramatic rifting of the supercontinent Pangaea, which gradually split into a northern continent, Laurasia, and a southern continent, Gondwana. This created the passive continental margin that characterizes most of the Atlantic coastline (such as along the U.S. East Coast) today. [21]

By the end of the era, the continents had rifted into nearly their present forms, though not their present positions. Laurasia became North America and Eurasia, while Gondwana split into South America, Africa, Australia, Antarctica and the Indian subcontinent, which collided with the Asian plate during the Cenozoic, giving rise to the Himalayas.

Climate

The Triassic was generally dry, a trend that began in the late Carboniferous, and highly seasonal, especially in the interior of Pangaea. Low sea levels may have also exacerbated temperature extremes. With its high specific heat capacity, water acts as a temperature-stabilizing heat reservoir, and land areas near large bodies of waterespecially oceansexperience less variation in temperature. Because much of Pangaea's land was distant from its shores, temperatures fluctuated greatly, and the interior probably included expansive deserts. Abundant red beds and evaporites such as halite support these conclusions, but some evidence suggests the generally dry climate of was punctuated by episodes of increased rainfall. [22] The most important humid episodes were the Carnian Pluvial Event and one in the Rhaetian, a few million years before the Triassic–Jurassic extinction event.

Sea levels began to rise during the Jurassic, probably caused by an increase in seafloor spreading. The formation of new crust beneath the surface displaced ocean waters by as much as 200 m (656 ft) above today's sea level, flooding coastal areas. Furthermore, Pangaea began to rift into smaller divisions, creating new shoreline around the Tethys Ocean. Temperatures continued to increase, then began to stabilize. Humidity also increased with the proximity of water, and deserts retreated.

The climate of the Cretaceous is less certain and more widely disputed. Probably, higher levels of carbon dioxide in the atmosphere are thought to have almost eliminated the north-south temperature gradient: temperatures were about the same across the planet, and about 10°C higher than today. The circulation of oxygen to the deep ocean may also have been disrupted, [16] [ dubious ] preventing the decomposition of large volumes of organic matter, which was eventually deposited as "black shale".

Not all data support these hypotheses, however. Even with the overall warmth, temperature fluctuations should have been sufficient for the presence of polar ice caps and glaciers, but there is no evidence of either. Quantitative models have also been unable to recreate the flatness of the Cretaceous temperature gradient.[ citation needed ]

Different studies have come to different conclusions about the amount of oxygen in the atmosphere during different parts of the Mesozoic, with some concluding oxygen levels were lower than the current level (about 21%) throughout the Mesozoic, [23] [24] some concluding they were lower in the Triassic and part of the Jurassic but higher in the Cretaceous, [25] [26] [27] and some concluding they were higher throughout most or all of the Triassic, Jurassic and Cretaceous. [28] [29]

Life

Flora

Conifers were the dominant terrestrial plants for most of the Mesozoic, with grass becoming widespread in the Late Cretaceous. Flowering plants appeared late in the era but did not become widespread until the Cenozoic. ProspectPemulwuypine.jpg
Conifers were the dominant terrestrial plants for most of the Mesozoic, with grass becoming widespread in the Late Cretaceous. Flowering plants appeared late in the era but did not become widespread until the Cenozoic.

The dominant land plant species of the time were gymnosperms, which are vascular, cone-bearing, non-flowering plants such as conifers that produce seeds without a coating. This is opposed to the earth's current flora, in which the dominant land plants in terms of number of species are angiosperms. One particular plant genus, Ginkgo , is thought to have evolved at this time and is represented today by a single species, Ginkgo biloba . As well, the extant genus Sequoia is believed to have evolved in the Mesozoic. [30]

Flowering plants radiated sometime in the early Cretaceous, first in the tropics, but the even temperature gradient allowed them to spread toward the poles throughout the period. By the end of the Cretaceous, angiosperms dominated tree floras in many areas, although some evidence suggests that biomass was still dominated by cycads and ferns until after the Cretaceous–Paleogene extinction. Some plant species had distributions that were markedly different from succeeding periods; for example, the Schizeales, a fern order, were skewed to the Northern Hemisphere in the Mesozoic, but are now better represented in the Southern Hemisphere. [31]

Fauna

Dinosaurs were the dominant terrestrial vertebrates throughout much of the Mesozoic. Europasaurus holgeri Scene 2.jpg
Dinosaurs were the dominant terrestrial vertebrates throughout much of the Mesozoic.

The extinction of nearly all animal species at the end of the Permian Period allowed for the radiation of many new lifeforms. In particular, the extinction of the large herbivorous pareiasaurs and carnivorous gorgonopsians left those ecological niches empty. Some were filled by the surviving cynodonts and dicynodonts, the latter of which subsequently became extinct.

Recent research indicates that it took much longer for the reestablishment of complex ecosystems with high biodiversity, complex food webs, and specialized animals in a variety of niches, beginning in the mid-Triassic 4M to 6M years after the extinction, [32] and not fully proliferated until 30M years after the extinction. [33] Animal life was then dominated by various archosaurs: dinosaurs, pterosaurs, and aquatic reptiles such as ichthyosaurs, plesiosaurs, and mosasaurs.

The climatic changes of the late Jurassic and Cretaceous favored further adaptive radiation. The Jurassic was the height of archosaur diversity, and the first birds and eutherian mammals also appeared. Some have argued that insects diversified in symbiosis with angiosperms, because insect anatomy, especially the mouth parts, seems particularly well-suited for flowering plants. However, all major insect mouth parts preceded angiosperms, and insect diversification actually slowed when they arrived, so their anatomy originally must have been suited for some other purpose.

See also

Related Research Articles

The Cretaceous is a geological period that lasted from about 145 to 66 million years ago (mya). It is the third and final period of the Mesozoic Era. The name is derived from the Latin creta. It is usually abbreviated K, for its German translation Kreide.

Cenozoic Third era of the Phanerozoic Eon 66 million years ago to present

The Cenozoic Era meaning "new life" is the current and most recent of the three geological eras of the Phanerozoic Eon. It follows the Mesozoic Era and extends from 66 million years ago to the present day. It is generally believed to have started on the first day of the Cretaceous–Paleogene extinction event when an asteroid hit the earth.

Extinction event Widespread and rapid decrease in the biodiversity on Earth

An extinction event is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from the threshold chosen for describing an extinction event as "major", and the data chosen to measure past diversity.

The Jurassic is a geologic period and system that spanned 56 million years from the end of the Triassic Period 201.3 million years ago (Mya) to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles. The start of the period was marked by the major Triassic–Jurassic extinction event. Two other extinction events occurred during the period: the Pliensbachian-Toarcian extinction in the Early Jurassic, and the Tithonian event at the end; neither event ranks among the "Big Five" mass extinctions, however.

The Permian is a geologic period and system which spans 47 million years from the end of the Carboniferous period 298.9 million years ago (Mya), to the beginning of the Triassic period 251.902 Mya. It is the last period of the Paleozoic era; the following Triassic period belongs to the Mesozoic era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.

The PaleozoicEra is the earliest of three geologic eras of the Phanerozoic Eon. It is the longest of the Phanerozoic eras, lasting from 541 to 251.902 million years ago, and is subdivided into six geologic periods : the Cambrian, Ordovician, Silurian, Devonian, Carboniferous, and Permian. The Paleozoic comes after the Neoproterozoic Era of the Proterozoic Eon and is followed by the Mesozoic Era.

The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 541 million years to the present, and began with the Cambrian Period when animals first developed hard shells preserved in the fossil record. The time before the Phanerozoic, called the Precambrian, is now divided into the Hadean, Archaean and Proterozoic eons.

Permian–Triassic extinction event Most severe extinction event of Earths chronology

The Permian–Triassic extinction event, also known as the P–Tr extinction, the P–T extinction, the End-Permian Extinction, and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as well as between the Paleozoic and Mesozoic eras, approximately 252 million years ago. It is the Earth's most severe known extinction event, with up to 96% of all marine species and 70% of terrestrial vertebrate species becoming extinct. It was the largest known mass extinction of insects. Some 57% of all biological families and 83% of all genera became extinct.

Supercontinent Landmass comprising more than one continental core, or craton

In geology, a supercontinent is the assembly of most or all of Earth's continental blocks or cratons to form a single large landmass. However, many earth scientists use a different definition: "a clustering of nearly all continents", which leaves room for interpretation and is easier to apply to Precambrian times.

The Triassic is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.9 million years ago (Mya), to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.

Triassic–Jurassic extinction event

The Triassic–Jurassic extinction event marks the boundary between the Triassic and Jurassic periods, 201.3 million years ago, and is one of the major extinction events of the Phanerozoic eon, profoundly affecting life on land and in the oceans. In the seas, a whole class (conodonts) and 23–34% of marine genera disappeared. On land, all archosaurs other than crocodylomorphs and Avemetatarsalia, some remaining therapsids, and many of the large amphibians became extinct. Statistical analysis of marine losses at this time suggests that the decrease in diversity was caused more by a decrease in speciation than by an increase in extinctions.

Early Triassic first of three epochs of the Triassic period of the geologic timescale

The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.902 Ma and 247.2 Ma. Rocks from this epoch are collectively known as the Lower Triassic series, which is a unit in chronostratigraphy.

Olenekian

In the geologic timescale, the Olenekian is an age in the Early Triassic epoch; in chronostratigraphy, it is a stage in the Lower Triassic series. It spans the time between 251.2 Ma and 247.2 Ma. The Olenekian is sometimes divided into the Smithian and the Spathian subages or substages. The Olenekian follows the Induan and is followed by the Anisian.

Late Triassic

The Late Triassic is the third and final of three epochs of the Triassic Period in the geologic timescale. The Triassic-Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. The corresponding series is known as the Upper Triassic. In Europe the epoch was called the Keuper, after a German lithostratigraphic group that has a roughly corresponding age. The Late Triassic spans the time between 237 Ma and 201.3 Ma. It is preceded by the Middle Triassic epoch and is followed by the Early Jurassic epoch. The Late Triassic is divided into the Carnian, Norian and Rhaetian ages.

The most recent understanding of the evolution of insects is based on studies of the following branches of science: molecular biology, insect morphology, paleontology, insect taxonomy, evolution, embryology, bioinformatics and scientific computing. It is estimated that the class of insects originated on Earth about 480 million years ago, in the Ordovician, at about the same time terrestrial plants appeared. Insects evolved from a group of crustaceans. The first insects were land bound, but about 400 million years ago in the Devonian period one lineage of insects evolved flight, the first animals to do so. The oldest insect fossil has been proposed to be Rhyniognatha hirsti, estimated to be 400 million years old, but the insect identity of the fossil has been contested. Global climate conditions changed several times during the history of Earth, and along with it the diversity of insects. The Pterygotes underwent a major radiation in the Carboniferous while the Endopterygota underwent another major radiation in the Permian.

Geological history of Earth The sequence of major geological events in Earths past

The geological history of Earth follows the major events in Earth's past based on the geological time scale, a system of chronological measurement based on the study of the planet's rock layers (stratigraphy). Earth formed about 4.54 billion years ago by accretion from the solar nebula, a disk-shaped mass of dust and gas left over from the formation of the Sun, which also created the rest of the Solar System.

Gondwana Neoproterozoic to Carboniferous supercontinent

Gondwana or Gondwanaland was a supercontinent that existed from the Neoproterozoic until the Jurassic.

Pangaea Supercontinent from the late Paleozoic to early Mesozoic eras

Pangaea or Pangea was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from earlier continental units approximately 335 million years ago, and it began to break apart about 175 million years ago. In contrast to the present Earth and its distribution of continental mass, Pangaea was centred on the Equator and surrounded by the superocean Panthalassa. Pangaea is the most recent supercontinent to have existed and the first to be reconstructed by geologists.

Prehistory of the United States

The prehistory of the United States comprises the occurrences within regions now part of the United States of America during the interval of time spanning from the formation of the Earth to the documentation of local history in written form. At the start of the Paleozoic era, what is now "North" America was actually in the southern hemisphere. Marine life flourished in the country's many seas, although terrestrial life had not yet evolved. During the latter part of the Paleozoic, seas were largely replaced by swamps home to amphibians and early reptiles. When the continents had assembled into Pangaea drier conditions prevailed. The evolutionary precursors to mammals dominated the country until a mass extinction event ended their reign.

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