485.4–443.8 million years ago
|Mean atmospheric O|
2 content over period duration
|c. 13.5 vol %|
(68 % of modern level)
|Mean atmospheric CO|
2 content over period duration
|c. 4200 ppm |
(15 times pre-industrial level)
|Mean surface temperature over period duration||c. 16 °C|
(2 °C above modern level)
|Sea level (above present day)||180 m; rising to 220 m in Caradoc and falling sharply to 140 m in end-Ordovician glaciations|
The Ordovician ( /,- -,- / or-də-VISH-ee-ən, -doh-, -VISH-ən) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.
The Ordovician, named after the Celtic tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in northern Wales into the Cambrian and Silurian systems, respectively.Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress.
Life continued to flourish during the Ordovician as it did in the earlier Cambrian period, although the end of the period was marked by the Ordovician–Silurian extinction events. Invertebrates, namely molluscs and arthropods, dominated the oceans. The Great Ordovician Biodiversification Event considerably increased the diversity of life. Fish, the world's first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. Life had yet to diversify on land. About 100 times as many meteorites struck the Earth per year during the Ordovician compared with today.
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The Ordovician Period began with a major extinction called the Cambrian–Ordovician extinction event, about 485.4 Mya (million years ago). It lasted for about 42 million years and ended with the Ordovician–Silurian extinction events, about 443.8 Mya (ICS, 2004) which wiped out 60% of marine genera. The dates given are recent radiometric dates and vary slightly from those found in other sources. This second period of the Paleozoic era created abundant fossils that became major petroleum and gas reservoirs.
The boundary chosen for the beginning of both the Ordovician Period and the Tremadocian stage is highly significant. It correlates well with the occurrence of widespread graptolite, conodont, and trilobite species. The base (start) of the Tremadocian allows scientists to relate these species not only to each other, but also to species that occur with them in other areas. This makes it easier to place many more species in time relative to the beginning of the Ordovician Period.
A number of regional terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected a formal international system of subdivisions.There exist Baltoscandic, British, Siberian, North American, Australian, Chinese Mediterranean and North-Gondwanan regional stratigraphic schemes.
|ICS Epoch||ICS stage||British epoch||British stage||North American epoch||North American stage||Australian epoch||Australian stage||Chinese epoch||Chinese stage|
|Late Ordovician||Hirnantian stage||Ashgill stage||Hirnantian stage||Cincinnati series||Gamach stage||Late Ordovician||Bolinda stage||Late Ordovician||Hirnantian stage|
|Katian stage||Rawthey stage||Richmond stage||Chientangkiang stage|
|Cautley stage||Maysville stage||Easton stage||Neichiashan stage|
|Pusgill stage||Eden stage|
|Caradoc series||Strefford stage||Mohawk stage||Chatfield stage|
|Sandbian stage||Burrell stage||Turin stage||Gisborne stage|
|Aureluc stage||Whiterock stage||Chazy stage|
|Middle Ordovician||Darriwilian stage||Llanvirn series||Llandeilo stage||Middle Ordovician||Darriwilian stage||Middle Ordovician||Darriwilian stage|
|Abereiddy stage||Not defined|
|Dapingian stage||Arenig series||Fenn stage||Early Ordovician||Yapeen stage||Dapingian stage|
|Whitland stage||Ranger stage||Castlemaine stage|
|Ibex series||Black Hills stage||Chewton stage|
|Early Ordovician||Floian stage||Moridun stage||Tule stage||Lancefield stage||Early Ordovician||Floian stage|
|Tremadocian stage||Tremadoc series||Migneint stage||Stairs stage||Tremadocian stage|
|Cressage stage||Skullrock stage|
The Ordovician Period in Britain was traditionally broken into Early (Tremadocian and Arenig), Middle (Llanvirn (subdivided into Abereiddian and Llandeilian) and Llandeilo) and Late (Caradoc and Ashgill) epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, or Upper part of the column. The faunal stages (subdivisions of epochs) from youngest to oldest are:
The Tremadoc corresponds to the (modern) Tremadocian. The Floian corresponds to the lower Arenig; the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc; the Caradoc ends in the mid-Katian, and the Ashgill represents the last half of the Katian, plus the Hirnantian. [ This would be clearer as a diagram. ]
During the Ordovician, the southern continents were collected into Gondwana. Gondwana started the period in equatorial latitudes and, as the period progressed, drifted toward the South Pole.
Early in the Ordovician, the continents of Laurentia (in present-day North America), Siberia, and Baltica (present-day northern Europe) were still independent continents (since the break-up of the supercontinent Pannotia earlier), but Baltica began to move towards Laurentia later in the period, causing the Iapetus Ocean between them to shrink. The small continent Avalonia separated from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia.
The Taconic orogeny, a major mountain-building episode, was well under way in Cambrian times. In the early and middle Ordovician, temperatures were mild, but at the beginning of the Late Ordovician, from 460 to 450 Ma, volcanoes along the margin of the Iapetus Ocean spewed massive amounts of carbon dioxide, a greenhouse gas, into the atmosphere, turning the planet into a hothouse.
Initially, sea levels were high, but as Gondwana moved south, ice accumulated into glaciers and sea levels dropped. At first, low-lying sea beds increased diversity, but later glaciation led to mass extinctions as the seas drained and continental shelves became dry land. During the Ordovician, in fact during the Tremadocian, marine transgressions worldwide were the greatest for which evidence is preserved.
These volcanic island arcs eventually collided with proto North America to form the Appalachian mountains. By the end of the Late Ordovician the volcanic emissions had stopped. Gondwana had by that time neared the South Pole and was largely glaciated.
The Ordovician meteor event is a proposed shower of meteors that occurred during the Middle Ordovician period, about 467.5±0.28 million years ago.It is not associated with any major extinction event.
The Ordovician was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, and invertebrate faunas with dominantly calcitic skeletons. Biogenic aragonite, like that composing the shells of most molluscs, dissolved rapidly on the sea floor after death.
Unlike Cambrian times, when calcite production was dominated by microbial and non-biological processes, animals (and macroalgae) became a dominant source of calcareous material in Ordovician deposits.
The Ordovician saw the highest sea levels of the Paleozoic, and the low relief of the continents led to many shelf deposits being formed under hundreds of metres of water.The sea level rose more or less continuously throughout the Early Ordovician, leveling off somewhat during the middle of the period. Locally, some regressions occurred, but sea level rise continued in the beginning of the Late Ordovician. Sea levels fell steadily in accord with the cooling temperatures for ~30 million years leading up to the Hirnantian glaciation. During this icy stage, sea level seems to have risen and dropped somewhat, but despite much study the details remain unresolved.
As with North America and Europe, Gondwana was largely covered with shallow seas during the Ordovician. Shallow clear waters over continental shelves encouraged the growth of organisms that deposit calcium carbonates in their shells and hard parts. The Panthalassic Ocean covered much of the northern hemisphere, and other minor oceans included Proto-Tethys, Paleo-Tethys, Khanty Ocean, which was closed off by the Late Ordovician, Iapetus Ocean, and the new Rheic Ocean.
As the Ordovician progressed, we see evidence of glaciers on the land we now know as Africa and South America, which were near the South Pole at the time, and covered by ice caps.
For most of the Late Ordovician life continued to flourish, but at and near the end of the period there were mass-extinction events that seriously affected planktonic forms like conodonts and graptolites. The trilobites Agnostida and Ptychopariida completely died out, and the Asaphida were much reduced. Brachiopods, bryozoans and echinoderms were also heavily affected, and the endocerid cephalopods died out completely, except for possible rare Silurian forms. The Ordovician–Silurian extinction events may have been caused by an ice age that occurred at the end of the Ordovician period, due to the expansion of the first terrestrial plants,as the end of the Late Ordovician was one of the coldest times in the last 600 million years of Earth's history.
On the whole, the fauna that emerged in the Ordovician were the template for the remainder of the Palaeozoic.The fauna was dominated by tiered communities of suspension feeders, mainly with short food chains. The ecological system reached a new grade of complexity far beyond that of the Cambrian fauna, which has persisted until the present day.
Though less famous than the Cambrian explosion, the Ordovician radiation was no less remarkable; marine faunal genera increased fourfold, resulting in 12% of all known Phanerozoic marine fauna.Another change in the fauna was the strong increase in filter-feeding organisms. The trilobite, inarticulate brachiopod, archaeocyathid, and eocrinoid faunas of the Cambrian were succeeded by those that dominated the rest of the Paleozoic, such as articulate brachiopods, cephalopods, and crinoids. Articulate brachiopods, in particular, largely replaced trilobites in shelf communities. Their success epitomizes the greatly increased diversity of carbonate shell-secreting organisms in the Ordovician compared to the Cambrian.
In North America and Europe, the Ordovician was a time of shallow continental seas rich in life. Trilobites and brachiopods in particular were rich and diverse. Although solitary corals date back to at least the Cambrian, reef-forming corals appeared in the early Ordovician, corresponding to an increase in the stability of carbonate and thus a new abundance of calcifying animals.
Molluscs, which appeared during the Cambrian or even the Ediacaran, became common and varied, especially bivalves, gastropods, and nautiloid cephalopods.
Now-extinct marine animals called graptolites thrived in the oceans. Some new cystoids and crinoids appeared.
It was long thought that the first true vertebrates (fish — Ostracoderms) appeared in the Ordovician, but recent discoveries in China reveal that they probably originated in the Early Cambrian.[ citation needed ] The very first gnathostome (jawed fish) appeared in the Late Ordovician epoch.
During the Middle Ordovician there was a large increase in the intensity and diversity of bioeroding organisms. This is known as the Ordovician Bioerosion Revolution.It is marked by a sudden abundance of hard substrate trace fossils such as Trypanites , Palaeosabella, Petroxestes and Osprioneides . Several groups of endobiotic symbionts appeared in the Ordovician.
In the Early Ordovician, trilobites were joined by many new types of organisms, including tabulate corals, strophomenid, rhynchonellid, and many new orthid brachiopods, bryozoans, planktonic graptolites and conodonts, and many types of molluscs and echinoderms, including the ophiuroids ("brittle stars") and the first sea stars. Nevertheless, the arthropods remained abundant, all the Late Cambrian orders continued, and were joined by the new group Phacopida. The first evidence of land plants also appeared (see evolutionary history of life).
In the Middle Ordovician, the trilobite-dominated Early Ordovician communities were replaced by generally more mixed ecosystems, in which brachiopods, bryozoans, molluscs, cornulitids, tentaculitids and echinoderms all flourished, tabulate corals diversified and the first rugose corals appeared. The planktonic graptolites remained diverse, with the Diplograptina making their appearance. Bioerosion became an important process, particularly in the thick calcitic skeletons of corals, bryozoans and brachiopods, and on the extensive carbonate hardgrounds that appear in abundance at this time. One of the earliest known armoured agnathan ("ostracoderm") vertebrate, Arandaspis , dates from the Middle Ordovician.
Trilobites in the Ordovician were very different from their predecessors in the Cambrian. Many trilobites developed bizarre spines and nodules to defend against predators such as primitive eurypterids and nautiloids while other trilobites such as Aeglina prisca evolved to become swimming forms. Some trilobites even developed shovel-like snouts for ploughing through muddy sea bottoms. Another unusual clade of trilobites known as the trinucleids developed a broad pitted margin around their head shields.Some trilobites such as Asaphus kowalewski evolved long eyestalks to assist in detecting predators whereas other trilobite eyes in contrast disappeared completely. Molecular clock analyses suggest that early arachnids started living on land by the end of the Ordovician.
The earliest-known octocorals date from the Ordovician.
Green algae were common in the Late Cambrian (perhaps earlier) and in the Ordovician. Terrestrial plants probably evolved from green algae, first appearing as tiny non-vascular forms resembling liverworts. Fossil spores from land plants have been identified in uppermost Ordovician sediments.
Among the first land fungi may have been arbuscular mycorrhiza fungi (Glomerales), playing a crucial role in facilitating the colonization of land by plants through mycorrhizal symbiosis, which makes mineral nutrients available to plant cells; such fossilized fungal hyphae and spores from the Ordovician of Wisconsin have been found with an age of about 460 million years ago, a time when the land flora most likely only consisted of plants similar to non-vascular bryophytes.
The Ordovician came to a close in a series of extinction events that, taken together, comprise the second largest of the five major extinction events in Earth's history in terms of percentage of genera that became extinct. The only larger one was the Permian–Triassic extinction event.
The extinctions occurred approximately 447–444 million years ago and mark the boundary between the Ordovician and the following Silurian Period. At that time all complex multicellular organisms lived in the sea, and about 49% of genera of fauna disappeared forever; brachiopods and bryozoans were greatly reduced, along with many trilobite, conodont and graptolite families.
The most commonly accepted theory is that these events were triggered by the onset of cold conditions in the late Katian, followed by an ice age, in the Hirnantian faunal stage, that ended the long, stable greenhouse conditions typical of the Ordovician.
The ice age was possibly not long-lasting. Oxygen isotopes in fossil brachiopods show its duration may have been only 0.5 to 1.5 million years.Other researchers (Page et al.) estimate more temperate conditions did not return until the late Silurian.
The late Ordovician glaciation event was preceded by a fall in atmospheric carbon dioxide (from 7000 ppm to 4400 ppm).The dip was triggered by a burst of volcanic activity that deposited new silicate rocks, which draw CO2 out of the air as they erode. This selectively affected the shallow seas where most organisms lived. As the southern supercontinent Gondwana drifted over the South Pole, ice caps formed on it, which have been detected in Upper Ordovician rock strata of North Africa and then-adjacent northeastern South America, which were south-polar locations at the time.
As glaciers grew, the sea level dropped, and the vast shallow intra-continental Ordovician seas withdrew, which eliminated many ecological niches. When they returned, they carried diminished founder populations that lacked many whole families of organisms. They then withdrew again with the next pulse of glaciation, eliminating biological diversity with each change.Species limited to a single epicontinental sea on a given landmass were severely affected. Tropical lifeforms were hit particularly hard in the first wave of extinction, while cool-water species were hit worst in the second pulse.
Those species able to adapt to the changing conditions survived to fill the ecological niches left by the extinctions.
At the end of the second event, melting glaciers caused the sea level to rise and stabilise once more. The rebound of life's diversity with the permanent re-flooding of continental shelves at the onset of the Silurian saw increased biodiversity within the surviving Orders.
An alternate extinction hypothesis suggested that a ten-second gamma-ray burst could have destroyed the ozone layer and exposed terrestrial and marine surface-dwelling life to deadly ultraviolet radiation and initiated global cooling.
Recent work considering the sequence stratigraphy of the Late Ordovician argues that the mass extinction was a single protracted episode lasting several hundred thousand years, with abrupt changes in water depth and sedimentation rate producing two pulses of last occurrences of species.
The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian, 419.2 million years ago (Mya), to the beginning of the Carboniferous, 358.9 Mya. It is named after Devon, England, where rocks from this period were first studied.
The PaleozoicEra is the earliest of three geologic eras of the Phanerozoic Eon. It is the longest of the Phanerozoic eras, lasting from, and is subdivided into six geologic periods : the Cambrian, Ordovician, Silurian, Devonian, Carboniferous, and Permian. The Paleozoic comes after the Neoproterozoic Era of the Proterozoic Eon and is followed by the Mesozoic Era.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by several million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
The Ordovician–Silurian extinction events, when combined, are the second-largest of the five major extinction events in Earth's history in terms of percentage of genera that became extinct. This event greatly affected marine communities, which caused the disappearance of one third of all brachiopod and bryozoan families, as well as numerous groups of conodonts, trilobites, and graptolites. The Ordovician–Silurian extinction occurred during the Hirnantian stage of the Ordovician Period and the subsequent Rhuddanian stage of the Silurian Period. The last event is dated in the interval of, lasting from the Middle Ordovician to Early Silurian, thus including the extinction period. This event was the first of the big five Phanerozoic events and was the first to significantly affect animal-based communities.
The Hirnantian is the seventh and final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 1.4 million years, from 445.2 to 443.8 Ma. The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation.
The Tremadocian is the lowest stage of Ordovician. Together with the later Floian stage it forms the Lower Ordovician epoch. The Tremadocian lasted from 485.4 to 477.7 million years ago. The base of the Tremadocian is defined as the first appearance of the conodont species Iapetognathus fluctivagus at the Global Boundary Stratotype Section and Point (GSSP) section on Newfoundland.
The history of invertebrate paleozoology differs from the history of paleontology in that the former usually emphasizes paleobiology and the paleoecology of extinct marine invertebrates, while the latter typically emphasizes the earth sciences and the sedimentary rock remains of terrestrial vertebrates.
The Kirengellids are a group of problematic Cambrian fossil shells of marine organisms. The shells bear a number of paired muscle scars on the inner surface of the valve.
Fossils of many types of water-dwelling animals from the Devonian period are found in deposits in the U.S. state of Michigan. Among the more commonly occurring specimens are bryozoans, corals, crinoids, and brachiopods. Also found, but not so commonly, are armored fish called placoderms, snails, sharks, stromatolites, trilobites and blastoids.
Paleontology in Ohio refers to paleontological research occurring within or conducted by people from the U.S. state of Ohio. Ohio is well known for having a great quantity and diversity of fossils preserved in its rocks. The state's fossil record begins early in the Paleozoic era, during the Cambrian period. Ohio was generally covered by seawater from that time on through the rest of the early Paleozoic. Local invertebrates included brachiopods, cephalopods, coral, graptolites, and trilobites. Vertebrates included bony fishes and sharks. The first land plants in the state grew during the Devonian. During the Carboniferous, Ohio became a more terrestrial environment with an increased diversity of plants that formed expansive swampy deltas. Amphibians and reptiles began to inhabit the state at this time, and remained present into the ensuing Permian. A gap in the local rock record spans from this point until the start of the Pleistocene. During the Ice Age, Ohio was home to giant beavers, humans, mammoths, and mastodons. Paleo-Indians collected fossils that were later incorporated into their mounds. Ohio has been the birthplace of many world famous paleontologists, like Charles Schuchert. Many significant fossils curated by museums in Europe and the United States were found in Ohio. Major local fossil discoveries include the 1965 discovery of more than 50,000 Devonian fish fossils in Cuyahoga County. The Ordovician trilobite Isotelus maximus is the Ohio state fossil.
Paleontology in Illinois refers to paleontological research occurring within or conducted by people from the U.S. state of Illinois. Scientists have found that Illinois was covered by a sea during the Paleozoic Era. Over time this sea was inhabited by animals including brachiopods, clams, corals, crinoids, sea snails, sponges, and trilobites.
Paleontology in Virginia refers to paleontological research occurring within or conducted by people from the U.S. state of Virginia. The geologic column in Virginia spans from the Cambrian to the Quaternary. During the early part of the Paleozoic, Virginia was covered by a warm shallow sea. This sea would come to be inhabited by creatures like brachiopods, bryozoans, corals, and nautiloids. The state was briefly out of the sea during the Ordovician, but by the Silurian it was once again submerged. During this second period of inundation the state was home to brachiopods, trilobites and entire reef systems. During the mid-to-late Carboniferous the state gradually became a swampy environment.
Paleontology in Tennessee refers to paleontological research occurring within or conducted by people from the U.S. state of Tennessee. During the early part of the Paleozoic era, Tennessee was covered by a warm, shallow sea. This sea was home to brachiopods, bryozoans, cephalopods, corals, and trilobites. Tennessee is one of the best sources of Early Devonian fossils in North America. During the mid-to-late Carboniferous, the state became a swampy environment, home to a rich variety of plants including ferns and scale trees. A gap in the local rock record spans from the Permian through the Jurassic. During the Cretaceous, the western part of the state was submerged by seawater. The local waters were home to more fossil gastropods than are known from anywhere else in the world. Mosasaurs and sea turtles also inhabited these waters. On land the state was home to dinosaurs. Western Tennessee was still under the sea during the early part of the Cenozoic. Terrestrial portions of the state were swampy. Climate cooled until the Ice Age, when the state was home to Camelops, horses, mammoths, mastodons, and giant ground sloths. The local Yuchi people told myths of giant lizard monsters that may have been inspired by fossils either local or encountered elsewhere. In 1920, after local fossils became a subject of formal scientific study, a significant discovery of a variety of Pleistocene creatures was made near Nashville. The Cretaceous bivalve Pterotrigonia thoracica is the Tennessee state fossil.
Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.
Paleontology in Missouri refers to paleontological research occurring within or conducted by people from the U.S. state of Missouri. The geologic column of Missouri spans all of geologic history from the Precambrian to present with the exception of the Permian, Triassic, and Jurassic. Brachiopods are probably the most common fossils in Missouri.
Paleontology in Minnesota refers to paleontological research occurring within or conducted by people from the U.S. state of Minnesota. The geologic record of Minnesota spans from Precambrian to recent with the exceptions of major gaps including the Silurian period, the interval from the Middle to Upper Devonian to the Cretaceous, and the Cenozoic. During the Precambrian, Minnesota was covered by an ocean where local bacteria ended up forming banded iron formations and stromatolites. During the early part of the Paleozoic era southern Minnesota was covered by a shallow tropical sea that would come to be home to creatures like brachiopods, bryozoans, massive cephalopods, corals, crinoids, graptolites, and trilobites. The sea withdrew from the state during the Silurian, but returned during the Devonian. However, the rest of the Paleozoic is missing from the local rock record. The Triassic is also missing from the local rock record and Jurassic deposits, while present, lack fossils. Another sea entered the state during the Cretaceous period, this one inhabited by creatures like ammonites and sawfish. Duckbilled dinosaurs roamed the land. The Cenozoic period of the ensuing Cenozoic era is also missing from the local rock record, but during the Ice Age evidence points to glacial activity in the state. Woolly mammoths, mastodons, and musk oxen inhabited Minnesota at the time. Local Native Americans interpreted such remains as the bones of the water monster Unktehi. They also told myths about thunder birds that may have been based on Ice Age bird fossils. By the early 19th century, the state's fossil had already attracted the attention of formally trained scientists. Early research included the Cretaceous plant discoveries made by Leo Lesquereux.
Paleontology in Oklahoma refers to paleontological research occurring within or conducted by people from the U.S. state of Oklahoma. Oklahoma has a rich fossil record spanning all three eras of the Phanerozoic Eon. Oklahoma is the best source of Pennsylvanian fossils in the United States due to having an exceptionally complete geologic record of the epoch. From the Cambrian to the Devonian, all of Oklahoma was covered by a sea that would come to be home to creatures like brachiopods, bryozoans, graptolites and trilobites. During the Carboniferous, an expanse of coastal deltaic swamps formed in areas of the state where early tetrapods would leave behind footprints that would later fossilize. The sea withdrew altogether during the Permian period. Oklahoma was home a variety of insects as well as early amphibians and reptiles. Oklahoma stayed dry for most of the Mesozoic. During the Late Triassic, carnivorous dinosaurs left behind footprints that would later fossilize. During the Cretaceous, however, the state was mostly covered by the Western Interior Seaway, which was home to huge ammonites and other marine invertebrates. During the Cenozoic, Oklahoma became home to creatures like bison, camels, creodonts, and horses. During the Ice Age, the state was home to mammoths and mastodons. Local Native Americans are known to have used fossils for medicinal purposes. The Jurassic dinosaur Saurophaganax maximus is the Oklahoma state fossil.
Trinodus is a very small to small blind trilobite, a well known group of extinct marine arthropods, which lived during the Ordovician, in what are now the Yukon Territories, Virginia, Italy, Czech Republic, Poland, Denmark, Sweden, Svalbard, Ireland, Scotland, Wales, Iran, Kazakhstan and China. It is one of the last of the Agnostida order to survive.
The Precordillera terrane of western Argentina is a large mountain range located southeast of the main Andes mountain range. The evolution of the Precordillera is noted for its unique formation history compared to the region nearby. The Cambrian-Ordovian sedimentology in the Precordillera terrane has its source neither from old Andes nor nearby country rock, but shares similar characteristics with the Grenville orogeny of eastern North America. This indicates a rift-drift history of the Precordillera in the early Paleozoic. The Precordillera is a moving micro-continent which started from the southeast part of the ancient continent Laurentia. The separation of the Precordillera started around the early Cambrian. The mass collided with Gondwana around Late Ordovician period. Different models and thinking of rift-drift process and the time of occurrence have been proposed. This page focuses on the evidence of drifting found in the stratigraphical record of the Precordillera, as well as exhibiting models of how the Precordillera drifted to Gondwana.
The geology of Morocco formed beginning up to two billion years ago, in the Paleoproterozoic and potentially even earlier. It was affected by the Pan-African orogeny, although the later Hercynian orogeny produced fewer changes and left the Maseta Domain, a large area of remnant Paleozoic massifs. During the Paleozoic, extensive sedimentary deposits preserved marine fossils. Throughout the Mesozoic, the rifting apart of Pangaea to form the Atlantic Ocean created basins and fault blocks, which were blanketed in terrestrial and marine sediments—particularly as a major marine transgression flooded much of the region. In the Cenozoic, a microcontinent covered in sedimentary rocks from the Triassic and Cretaceous collided with northern Morocco, forming the Rif region. Morocco has extensive phosphate and salt reserves, as well as resources such as lead, zinc, copper and silver.
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