Paleobiota of the Liushu Formation

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An artist's illustration of the Paleobiota and environment of the upper part of the Liushu formation Lishu Formation ecosystem.png
An artist's illustration of the Paleobiota and environment of the upper part of the Liushu formation

The Liushu formation is a geological formation in the Gansu province of China that spans up to 100 m thick and is widely distributed within the Linxia Basin, with a paleomagnetic age of between 11 and 6.4 Mya (Upper/Late Miocene).

Contents

The formation is divided into three parts which represent three different ages, generally referred to as the "upper part" (youngest, at 6-7 mya), the "middle part" (intermediate, at 9 to 7.6 mya. Also known as the Dashengou fauna, and the "lower part" (oldest). [1] [2]

Mammals

Euungulata

Perissodactyla

The Liushu formation displays a lineage of Elasmotheriine rhinoceros throughout its stratigraphy. Iranotherium is restricted to the middle part of the formation but remains become absent in the overlying layers, the only Elasmotheriine that can be found in the upper layers is Sinotherium .

GenusSpeciesLocalityMaterialNotesImages
Upper part
Ancylotherium sp. [3] Guanghe County [3] A Shizotheriine Chalicothere
HipparionH. coelophyes [3] Guanghe County [3] An ancient horse

Sinotherium

S. lagrelii [3]

Guanghe County [3]

A partial Skull

An Elasmotheriine, ancestral to

Elasmotherium . Displays the important

transition from nose-horned to forehead

horned Rhinos.

Middle part
Acerorhinus hezhengensisA acerorhine rhino

Chilotherium

C. wimani

Guanghe County [3]

A tusked rhino

Hipparion H. chiaiAn ancient horse
HipparionH. weihoensisAn ancient horse

Iranotherium

I. morgani [2]

Guanghe County [2]

one female skull, one

male skull, and one

isolated mandible

An elasmotheriine with unique sexual

dimorphism. Ancestral to Sinotherium

Lower part
HipparionH. dongxiangense [4]
Ningxiatherium sp.
Parelasmotherium P. linxiaense [4] Dongxiang County [4] A mostly complete

skull

A grazing Elasmotheriine that coexisted

with another browsing Elasmotheriine

( Ningxiatherium ).

Parelasmotherium P. simplum [4]

Artiodactyla

GenusSpeciesCountyMaterialNotesImages
Upper part
Dicrocerus sp. [3] Guanghe County [3] an ancient deer
Paleotragus microdon [3] Guanghe County [3] an ancient giraffid
Protoryx sp. [3] Guanghe County [3] an ancient bovid
Sinotragus wimani [3] Guanghe County [3] an ancient bovid
Middle part
Chleuastochoerus stehlinia prehistoric pig
Dicrocerus sp. [3] Guanghe County [3] an ancient deer
Gazella sp.
Hezhengiabohlinia bovid
Honanotherium schlosserian ancient giraffid
Miotragocerus sp.an extinct antelope
Samotherium sp.an ancient giraffid
Lower part
Shaanxispira sp. [4] an ancient bovid

Carnivora

Caniformia

GenusSpeciesCountyMemberMaterialNotesImages
Melodon majoriMiddle parta musteloid
Parataxidea [3] sinensis [3] Guanghe County [3] Upper part [3] a badger-like mustelid
Promephitis hootoni
Promephitis sp.Middle parta musteloid
Sinictis sp.Middle parta musteloid
Ursavus [3] sp. [3] Guanghe County [3] Upper part [3] an early bear

Feliformia

GenusSpeciesCountyMemberMaterialNotesImages
Amphimachairodus [3] giganteus [3] Guanghe County [3] Upper part [3] to Middle parta homotheriine sabercat
Dinocrocuta giganteaMiddle parta gigantic hyena
Felis [3] sp. [3] Guanghe County [3] Upper part [3] to Middle part
Hyaenictitherium [3] wongii [3] Guanghe County [3] Upper part to Middle part [3] a Hyaenid
Hyaenictitheriumhyaenoides [3] Upper part to Middle part
Ictitherium [3] sp. [3] Guanghe County [3] Upper part [3] to Middle parta Hyaenid
Metailurus [3] major [3] Guanghe County [3] Upper part [3] a metalurin sabercat

Glires

GenusSpeciesCountyMemberMaterialNotesImages
Pararhizomys hipparionumMiddle partA rodent.

Proboscidea

GenusSpeciesCountyMemberMaterialNotesImages
Tetralophodon exoletusMiddle part
Konobelodon robustusAn amebelodont

Birds

GenusSpeciesLocalityMemberMaterialNotesImages
Linxiavis [5] inaquosus [5] Tibetan plateau, Gansu provinceMiddle part - Upper part [5] articulated and associated elements of the

wings, shoulder girdle, vertebrae, and hind limb

An extinct Sandgrouse
Vulture
Ostrich
Pheasant
Falcon
eogruid

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References

  1. "Vertebrata PalAsiatica, Vol. 42, 2004----Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences". english.ivpp.cas.cn. Retrieved 2020-12-21.
  2. 1 2 3 Deng, Tao (2005-06-27). "New discovery of Iranotherium morgani (Perissodactyla, Rhinocerotidae) from the late Miocene of the Linxia Basin in Gansu, China, and its sexual dimorphism". Journal of Vertebrate Paleontology. 25 (2): 442–450. doi:10.1671/0272-4634(2005)025[0442:NDOIMP]2.0.CO;2. ISSN   0272-4634. S2CID   85820005.
  3. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 Deng, Tao; Wang, ShiQi; Hou, SuKuan (2013-05-01). "A bizarre tandem-horned elasmothere rhino from the Late Miocene of northwestern China and origin of the true elasmothere". Chinese Science Bulletin. 58 (15): 1811–1817. Bibcode:2013ChSBu..58.1811D. doi: 10.1007/s11434-012-5574-4 . ISSN   1861-9541.
  4. 1 2 3 4 5 Deng, Tao (2007-06-12). "Skull of Parelasmotherium (Perissodactyla, Rhinocerotidae) from the upper miocene in the Linxia Basin (Gansu, China)". Journal of Vertebrate Paleontology. 27 (2): 467–475. doi:10.1671/0272-4634(2007)27[467:SOPPRF]2.0.CO;2. ISSN   0272-4634. S2CID   130213090.
  5. 1 2 3 Li, Zhiheng; Stidham, Thomas A.; Deng, Tao; Zhou, Zhonghe (2020-03-31). "Evidence of Late Miocene Peri-Tibetan Aridification From the Oldest Asian Species of Sandgrouse (Aves: Pteroclidae)". Frontiers in Ecology and Evolution. 8: 59. doi: 10.3389/fevo.2020.00059 . ISSN   2296-701X.