The Liushu formation is a geological formation in the Gansu province of China that spans up to 100 m thick and is widely distributed within the Linxia Basin, with a paleomagnetic age of between 11 and 6.4 Mya (Upper/Late Miocene).
The formation is divided into three parts which represent three different ages, generally referred to as the "upper part" (youngest, at 6-7 mya), the "middle part" (intermediate, at 9 to 7.6 mya. Also known as the Dashengou fauna, and the "lower part" (oldest). [1] [2]
The Liushu formation displays a lineage of Elasmotheriine rhinoceros throughout its stratigraphy. Iranotherium is restricted to the middle part of the formation but remains become absent in the overlying layers, the only Elasmotheriine that can be found in the upper layers is Sinotherium .
Genus | Species | Locality | Material | Notes | Images |
---|---|---|---|---|---|
Upper part | |||||
Ancylotherium | sp. [3] | Guanghe County [3] | A Shizotheriine Chalicothere | ||
Hipparion | H. coelophyes [3] | Guanghe County [3] | An ancient horse | ||
S. lagrelii [3] | Guanghe County [3] | A partial Skull | An Elasmotheriine, ancestral to Elasmotherium . Displays the important transition from nose-horned to forehead horned Rhinos. | ||
Middle part | |||||
Acerorhinus | hezhengensis | A acerorhine rhino | |||
C. wimani | Guanghe County [3] | ||||
Hipparion | H. chiai | An ancient horse | |||
Hipparion | H. weihoensis | An ancient horse | |||
I. morgani [2] | Guanghe County [2] | one female skull, one male skull, and one isolated mandible | An elasmotheriine with unique sexual dimorphism. Ancestral to Sinotherium | ||
Lower part | |||||
Hipparion | H. dongxiangense [4] | ||||
Ningxiatherium | sp. | ||||
Parelasmotherium | P. linxiaense [4] | Dongxiang County [4] | A mostly complete skull | A grazing Elasmotheriine that coexisted with another browsing Elasmotheriine ( Ningxiatherium ). | |
Parelasmotherium | P. simplum [4] | ||||
Genus | Species | County | Material | Notes | Images |
---|---|---|---|---|---|
Upper part | |||||
Dicrocerus | sp. [3] | Guanghe County [3] | an ancient deer | ||
Paleotragus | microdon [3] | Guanghe County [3] | an ancient giraffid | ||
Protoryx | sp. [3] | Guanghe County [3] | an ancient bovid | ||
Sinotragus | wimani [3] | Guanghe County [3] | an ancient bovid | ||
Middle part | |||||
Chleuastochoerus | stehlini | a prehistoric pig | |||
Dicrocerus | sp. [3] | Guanghe County [3] | an ancient deer | ||
Gazella | sp. | ||||
Hezhengia | bohlini | a bovid | |||
Honanotherium | schlosseri | an ancient giraffid | |||
Miotragocerus | sp. | an extinct antelope | |||
Samotherium | sp. | an ancient giraffid | |||
Lower part | |||||
Shaanxispira | sp. [4] | an ancient bovid |
Genus | Species | County | Member | Material | Notes | Images |
---|---|---|---|---|---|---|
Melodon | majori | Middle part | a musteloid | |||
Parataxidea [3] | sinensis [3] | Guanghe County [3] | Upper part [3] | a badger-like mustelid | ||
Promephitis | hootoni | |||||
Promephitis | sp. | Middle part | a musteloid | |||
Sinictis | sp. | Middle part | a musteloid | |||
Ursavus [3] | sp. [3] | Guanghe County [3] | Upper part [3] | an early bear |
Genus | Species | County | Member | Material | Notes | Images |
---|---|---|---|---|---|---|
Amphimachairodus [3] | giganteus [3] | Guanghe County [3] | Upper part [3] to Middle part | a homotheriine sabercat | ||
Dinocrocuta | gigantea | Middle part | a gigantic hyena | |||
Felis [3] | sp. [3] | Guanghe County [3] | Upper part [3] to Middle part | |||
Hyaenictitherium [3] | wongii [3] | Guanghe County [3] | Upper part to Middle part [3] | a Hyaenid | ||
Hyaenictitherium | hyaenoides [3] | Upper part to Middle part | ||||
Ictitherium [3] | sp. [3] | Guanghe County [3] | Upper part [3] to Middle part | a Hyaenid | ||
Metailurus [3] | major [3] | Guanghe County [3] | Upper part [3] | a metalurin sabercat |
Genus | Species | County | Member | Material | Notes | Images |
---|---|---|---|---|---|---|
Pararhizomys | hipparionum | Middle part | A rodent. |
Genus | Species | County | Member | Material | Notes | Images |
---|---|---|---|---|---|---|
Tetralophodon | exoletus | Middle part | ||||
Konobelodon | robustus | An amebelodont |
Genus | Species | Locality | Member | Material | Notes | Images |
---|---|---|---|---|---|---|
Linxiavis [5] | inaquosus [5] | Tibetan plateau, Gansu province | Middle part - Upper part [5] | articulated and associated elements of the wings, shoulder girdle, vertebrae, and hind limb | An extinct Sandgrouse | |
Vulture | ||||||
Ostrich | ||||||
Pheasant | ||||||
Falcon | ||||||
eogruid |
A rhinoceros, commonly abbreviated to rhino, is a member of any of the five extant species of odd-toed ungulates in the family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South and Southeast Asia.
Diceros is a genus of rhinoceros containing the extant black rhinoceros (Diceros bicornis) and several extinct species.
Ceratotherium is a genus of the family Rhinocerotidae, consisting of a single extant species, the white rhinoceros, and its extinct relative Ceratotherium mauritanicum, of which Ceratotherium efficax is considered a synonym. Another species known as Ceratotherium praecox is now considered a member of the related genus Diceros. The placement of Ceratotherium neumayri from the Late Miocene of Europe and Western Asia within the genus has been questioned, with other authors assigning it to the separate genus Miodiceros. The species 'Ceratotherium’ advenientis is known from the Late Miocene of Italy.
Chalicotheriidae is an extinct family of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Pleistocene, existing from 48.6 to 1.806 mya. They are often called chalicotheres, a term which is also applied to the broader grouping of Chalicotherioidea. They are noted for their unusual morphology compared to other ungulates, such as their elongated clawed forelimbs. They are thought to have been browsers.
Ancylotherium is an extinct genus of the family Chalicotheriidae, subfamily Schizotheriinae, endemic to Europe, Asia, and Africa during the Late Miocene-Early Pleistocene, existing for approximately 9.8 million years.
Hemicyon, also known as the "dog-bear", is an extinct genus of hemicyonine bear, which probably originated in Eurasia but was found in Europe, Asia and North America during the Miocene epoch, existing for approximately 16 to 13 mya. Hemicyon is the best-known genus in the Hemicyoninae, a subfamily intermediate between bears and their Caniform ancestors but most often classified as bears. Hemicyonid bears should not be confused with Amphicyonids (bear-dogs), which are their own separate family of carnivores.
Sinotherium is an extinct genus of single-horned elasmotheriine rhinocerotids that lived from the late Miocene to Early Pliocene. It was ancestral to Elasmotherium, demonstrating a very important evolutionary transition from nasal-horned elasmotheriines to frontal-horned elasmotheriines. Its fossils have been found in the Karabulak Formation of Kazakhstan, lower jaw and teeth have been found in Mongolia, and a partial skull is known from the upper part of the Liushu Formation of western China. Sinotherium diverged from the ancestral genus, Iranotherium, first found in Iran, during the early Pliocene. Some experts prefer to lump Sinotherium, and Iranotherium into Elasmotherium.
The John Day Formation is a series of rock strata exposed in the Picture Gorge district of the John Day River basin and elsewhere in north-central Oregon in the United States. The Picture Gorge exposure lies east of the Blue Mountain uplift, which cuts southwest–northeast through the Horse Heaven mining district northeast of Madras. Aside from the Picture Gorge district, which defines the type, the formation is visible on the surface in two other areas: another exposure is in the Warm Springs district west of the uplift, between it and the Cascade Range, and the third is along the south side of the Ochoco Mountains. All three exposures, consisting mainly of tuffaceous sediments and pyroclastic rock rich in silica, lie unconformably between the older rocks of the Clarno Formation below and Columbia River basalts above.
Iranotherium is an extinct genus of large elasmotheriine rhinocerotids, comparable in size to a modern white rhino. It is known from the Late Miocene (Tortonian) of Maragha, Iran and the middle part of the Liushu formation of northwestern China. It was a precursor to the related Sinotherium and may have been ultimately outcompeted by its descendant. This species is most well known for showing unique sexual dimorphism among rhinos.
Cormohipparion is an extinct genus of horse belonging to the tribe Hipparionini that lived in North America during the late Miocene to Pliocene. This ancient species of horse grew up to 3 feet long.
Chilotherium is an extinct genus of rhinocerotids endemic to Eurasia during the Miocene through Pliocene living for 13.7—3.4 mya, existing for approximately 10.3 million years.
Nestoritherium is an extinct genus of chalicothere; it has been dated to have lived from the late Miocene to the Early Pleistocene. This range makes Nestoritherium one of the most recently dated chalicotheres. It has been found in fossil sites in Myanmar and China.
Tetralophodon is an extinct genus of "tetralophodont gomphothere" belonging to the superfamily Elephantoidea, known from the Miocene of Afro-Eurasia.
Hispanotherium is an extinct genus of rhinocerotid of the tribe Elasmotheriini endemic to Europe and Asia during the Miocene living from 16—7.25 mya existing for approximately 8.75 million years.
Ceratotherium neumayri is a fossil species of rhinoceros from the Late Miocene (Vallesian-Turolian) of the Balkans and Western Asia, with remains known from Greece, Bulgaria, Iran, and Anatolia in Turkey.
Deng Tao is a Chinese palaeontologist at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP), Chinese Academy of Sciences, who has made important fossil discoveries on Cenozoic mammals. He is a professor of vertebrate palaeontology, deputy director of the Academic Committee, and deputy director of Key Laboratory of Evolutionary Systematics of Vertebrates at IVPP.
The St. Marys Formation is a geologic formation in Maryland and Virginia, United States. It preserves fossils dating back to the Miocene Epoch of the Neogene period. It is the youngest Miocene formation present in the Calvert Cliffs and is part of the Chesapeake Group.
The Astoria Formation is a geologic formation in Washington state & Oregon. It preserves fossils dating back to the early to middle Miocene.
Linxiavis inaquosus is an extinct species of sandgrouse known from a partial skeleton, found in the Late Miocene Liushu Formation at the edge of the Tibetan Plateau in Gansu Province of China. It is the most substantial and oldest record of crown Sandgrouse in China which adds to the rapidly growing avian fauna of the Liushu formation.
Miosurnia is an extinct genus of surniin bird from the Late Miocene Liushu Formation of Gansu Province, China. The genus contains a single species, Miosurnia diurna, known from a nearly complete, articulated skeleton.