Simosthenurus occidentalis

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Simosthenurus occidentalis
Simosthenurus BW.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Marsupialia
Order: Diprotodontia
Family: Macropodidae
Genus: Simosthenurus
Species:
S. occidentalis
Binomial name
Simosthenurus occidentalis
Glauert 1910 [1]

Simosthenurus occidentalis is a species of sthenurine marsupial ("short-faced kangaroo") that existed in Australia during the Pliocene, becoming extinct in the Pleistocene epoch around 42,000 years ago. It was a large herbivorous biped that resembles large kangaroos, but with a heavier body than modern kangaroos. The structure of the skull and teeth - resembling koalas and panda bears - indicates that it consumed tough vegetation.

Taxonomy

The type specimen was collected by E. A. Le Souef and noted in a report to the state's Caves Board, then revised and published by Ludwig Glauert as Sthenurus occidentalis in 1910. This holotype is fossil material preserving the left and right dentary of an adult found at Mammoth Cave in Southwest Australia. [1] The specific epithet occidentalis, meaning "of the west", refers to the discovery of this species in Western Australia. [2]

Description

Reconstruction Simosthenurus occidentalis.JPG
Reconstruction

A mid-sized species of Simosthenurus , known as 'short-faced' kangaroos, one of several genera in a macropodid lineage that diversified in Pliocene Australia. The mass of adults is estimated to have around 118 kilograms, standing at a similar height to larger modern roos but with a heavier and more robust body. The dentition contains molars set close to the jaw, which combined with the cranial structure, strongly indicates an animal capable of consuming tough vegetation. [2] The short jaw and facial anatomy clearly distinguishes their appearance from the surviving lineage of kangaroos, which usually graze on grasses rather than a browsing diet, and more closely resembles the habits of the modern koala Phascolarctos cinereus . [3]

A set of powerful molars located close to the hinge of jaw, similar to but further back than a koala, improved the mechanical advantage and a potential enlargement of the zygomaticomandibularis muscles would have reduced the hazard of dislocation while biting with the rear molars. [3] Modelling of the bite force and the skulls resistance to torsional forces suggests that the species was able to consume tough material, such as leaves, bark and branches, that could be torn with the claws of the long forelimbs. The diamond-like shape of the forehead and reinforced cheek bones was able to distribute the forces applied by large muscles to the rear molars at one side of the jaw, strongly suggesting that browsing habit included an ability to tear and chew hardy and fibrous material. The morphology of the skull and dentition was found to be most comparable to Asian mammal Ailuropoda melanoleuca (panda), an animal that primarily feeds as a browser of tough plant material. [4]

Distribution and habitat

The species is represented in the records of many southern fossil sites in Australia, including Tasmania. The type locality is located in Southwest Australia, at Mammoth Cave, and evidence from another site in the southwest indicates it existed until about 42,000 years ago. [3] Fossils assigned to the species have been obtained at sites in the Southwest, southern regions of Central Australia and the southeast of the continent. In Tasmania S. occidentalis has been identified at fossil sites in the northeast and central regions, the species has also been found at the nearby King Island. [2]

Their distribution in the Pleistocene is dated to late-middle to late deposits. [2]

Related Research Articles

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Diprotodon is an extinct genus of marsupial from the Pleistocene of Australia containing one species, D. optatum. The earliest finds date to 1.77 million to 780,000 years ago but most specimens are dated to after 110,000 years ago. Its remains were first unearthed in 1830 in Wellington Caves, New South Wales, and contemporaneous paleontologists guessed they belonged to rhinos, elephants, hippos or dugongs. Diprotodon was formally described by English naturalist Richard Owen in 1838, and was the first named Australian fossil mammal, and led Owen to become the foremost authority of his time on other marsupials and Australian megafauna, which were enigmatic to European science.

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<i>Thylacoleo</i> Extinct genus of marsupials

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Procoptodon is an extinct genus of giant short-faced (sthenurine) kangaroos that lived in Australia during the Pleistocene Epoch. P. goliah, the largest known kangaroo species that ever existed, stood at about 2 m (6.6 ft). They weighed about 200–240 kg (440–530 lb). Other members of the genus were smaller, however; Procoptodon gilli was the smallest of all of the sthenurine kangaroos, standing approximately 1 m tall.

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<i>Hipposideros besaoka</i> Extinct species of bat

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<i>Congruus</i> Extinct genus of macropod

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References

  1. 1 2 Glauert, L. (1910). "The Mammoth Cave". Records of the Western Australian Museum. 1. Western Australian Museum: 11–36.
  2. 1 2 3 4 Prideaux, G.J. (2004). Systematics and evolution of the sthenurine kangaroo. University of California Press. ISBN   0-520-09845-5.
  3. 1 2 3 Mitchell, D.R.; Evans, A.R. (11 September 2019). "The anatomy of a crushing bite: The specialised cranial mechanics of a giant extinct kangaroo". PLOS ONE. 14 (9): e0221287. Bibcode:2019PLoSO..1421287M. doi: 10.1371/journal.pone.0221287 . PMC   6738596 . PMID   31509570.
  4. Lyons, Suzannah (12 September 2019). "A jaw made to gnaw: Ancient kangaroos had strong skulls and a serious bite". ABC News.