Coelophysidae

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Coelophysids
Temporal range: Late Triassic-Early Jurassic, 228–183  Ma
Coelophysidae.png
Skull comparison of Coelophysidae ( Megapnosaurus rhodesiensis at the top left, Panguraptor lufengensis at the top right, Coelophysis bauri at the bottom left, "Syntarsus" kayentakatae at the bottom right)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Superfamily: Coelophysoidea
Family: Coelophysidae
Nopcsa, 1923
Type species
Coelurus bauri
Cope, 1887
Genera
Synonyms
  • Procompsognathidae Nopcsa, 1923
  • Segisauridae Camp, 1936

Coelophysidae is a family of primitive carnivorous theropod dinosaurs. Most species were relatively small in size. The family flourished in the Late Triassic and Early Jurassic periods, and has been found on numerous continents. Many members of Coelophysidae are characterized by long, slender skulls and light skeletons built for speed. [2] One member genus, Coelophysis , displays the earliest known furcula in a dinosaur. [3]

Contents

Under cladistic analysis, Coelophysidae was first defined by Paul Sereno in 1998 as the most recent common ancestor of Coelophysis bauri and Procompsognathus triassicus , and all of that common ancestor's descendants. [2] However, Tykoski (2005) has advocated for the definition to change to include the additional taxa of "Syntarsus" kayentakatae and Segisaurus halli . [4] Coelophysidae is part of the superfamily Coelophysoidea, which in turn is a subset of the larger Neotheropoda clade. [2] As part of Coelophysoidea, Coelophysidae is often placed as sister to the Dilophosauridae family, however, the monophyly of this clade has often been disputed. [2] The older term "Podokesauridae", named 14 years prior to Coelophysidae (which would normally grant it priority), is now usually ignored, since its type specimen was destroyed in a fire and can no longer be compared to new finds. [5]

Classification

Characteristics

Coelophysids are characterized by slender, skinny builds and long, narrow skulls with large fenestrae to allow for a lighter skull. [6] They are fairly primitive theropods, and so have fairly basal characteristics, such as hollow air sacs in the cervical vertebrae and obligate bipedalism. [6] Their slender builds allowed them to be fast and agile runners. All known members of Coelophysidae are carnivores. One species, Coelophysis bauri has the oldest known furcula (wishbone) of any dinosaur. [3]

It has also been speculated that some species within Coelophysidae, namely Coelophysis bauri, displayed cannibalism, although the fossil evidence behind these claims has been heavily debated (Rinehart et al., 2009; Gay, 2002; Gay, 2010). [7] [8] [9]

Phylogenetics

Coelophysidae is part of the larger superfamily of Coelophysoidea, which contains Dilophosauridae, Liliensternus , and Zupaysaurus in addition to Coelophysidae. [2] [10] [11] Coelophysoidea, in turn, is part of the larger clade of Neotheropoda. [2]

The cladogram below was recovered in a study by Matthew T. Carrano, John R. Hutchinson and Scott D. Sampson, 2005. [12]

Procompsognathus

Segisaurus

Coelophysis

Megapnosaurus rhodesiensis

Megapnosaurus kayentakatae

The cladogram below follows the topology from a 2011 analysis by paleontologists Martin D. Ezcurra and Stephen L. Brusatte, modified with additional data by You Hai-Lu and colleagues in 2014. [1] [13]

"Syntarsus" kayentakatae

Panguraptor lufengensis

Coelophysis bauri

Coelophysis rhodesiensis

Camposaurus arizonensis

This cladogram follows the 2017 analysis by Ezcurra, [14] with added results of the Martinez & Apaldetti analysis from their description of Lucianovenator. [15]

"Syntarsus" kayentakatae

Panguraptor lufengensis

Powellvenator podocitus

Procompsognathus triassicus

Coelophysis bauri

Lepidus praecisio

Segisaurus halli

Megapnosaurus rhodesiensis

Lucianovenator bonoi

Camposaurus arizonensis

Biogeography

Fossils of members of Coelophysidae have been found across many continents, including North America, South America, Europe, Asia, and Africa. Powellvenator podocitus was discovered in Northwestern Argentina. [16] Procompsognathus triassicus was discovered in Germany, and Camposaurus arizonensis is from Arizona in North America. [17] [13] No coelophysid fossils were known from Asia until the discovery of Panguraptor lufengensis in 2014 in the Yunnan Province of China. [1] The genus Coelophysis has been found in North America, South Africa, and Zimbabwe. [18]

See also

Related Research Articles

<i>Coelophysis</i> Genus of theropod dinosaurs from the late Triassic

Coelophysis is a genus of coelophysid theropod dinosaur that lived approximately 215 to 208.5 million years ago during the Late Triassic period from the middle to late Norian age in what is now the southwestern United States. Megapnosaurus was once considered to be a species within this genus, but this interpretation has been challenged since 2017 and the genus Megapnosaurus is now considered valid.

<span class="mw-page-title-main">Coelophysoidea</span> Extinct superfamily of dinosaurs

Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the coelurosaurs, with which they were formerly classified, and some species had delicate cranial crests. Sizes range from about 1 to 6 m in length. It is unknown what kind of external covering coelophysoids had, and various artists have portrayed them as either scaly or feathered. Some species may have lived in packs, as inferred from sites where numerous individuals have been found together.

<i>Procompsognathus</i> Extinct genus of dinosaurs

Procompsognathus is an extinct genus of coelophysid theropod dinosaur that lived approximately 210 million years ago during the later part of the Triassic Period, in what is now Germany. Procompsognathus was a small-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 1 m (3.3 ft) long.

<i>Sarcosaurus</i> Extinct genus of dinosaurs

Sarcosaurus is a genus of basal neotheropod dinosaur, roughly 3.5 metres (11 ft) long. It lived in what is now England and maybe Ireland and Scotland during the Hettangian-Sinemurian stages of the Early Jurassic, about 199-196 million years ago. Sarcosaurus is one of the earliest known Jurassic theropods, and one of only a handful of theropod genera from this time period. Along with Dracoraptor hanigani it is one of the two described neotheropods from the lowermost Jurassic of the United Kingdom.

<i>Zupaysaurus</i> Extinct genus of dinosaurs

Zupaysaurus is an extinct genus of early theropod dinosaur living during the Norian stage of the Late Triassic in what is now Argentina. Fossils of the dinosaur were found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina. Although a full skeleton has not yet been discovered, Zupaysaurus can be considered a bipedal predator, up to 4 metres (13 ft) long. It may have had two parallel crests running the length of its snout.

<i>Camposaurus</i> Extinct genus of dinosaurs

Camposaurus is a coelophysid dinosaur genus from the Norian stage of the Late Triassic period of North America. The pertinent fossil remains date back to the early to middle Norian stage, and is widely regarded as the oldest known neotheropod.

Lophostropheus is an extinct genus of coelophysoid theropod dinosaur that lived approximately 205.6 to 196.5 million years ago during the boundary between the Late Triassic Period and the Early Jurassic Period, in what is now Normandy, France. Lophostropheus is one of the few dinosaurs that may have survived the Triassic–Jurassic extinction event.

<span class="mw-page-title-main">Los Colorados Formation</span>

The Los Colorados Formation is a sedimentary rock formation of the Ischigualasto-Villa Unión Basin, found in the provinces of San Juan and La Rioja in Argentina. The formation dates back to the Norian age of the Late Triassic.

The Quebrada del Barro Formation is a geological formation of the Marayes-El Carrizal Basin in San Juan Province, Argentina. This formation is the most fossiliferous portion of the Triassic Marayes Group, and is also the youngest unit of the group, overlying the El Carrizal Formation. An unconformity at the top of the Quebrada del Barro Formation separates it from the Cretaceous-age Los Riscos Formation of the El Gigante Group. Part of the formation may be made into a provincial park following the discovery of the fossils of Ingentia, a giant sauropodomorph dinosaur which helped elucidate the early evolution of sauropods.

<span class="mw-page-title-main">Neotheropoda</span> Clade of theropods

Neotheropoda is a clade that includes coelophysoids and more advanced theropod dinosaurs, and is the only group of theropods that survived the Triassic–Jurassic extinction event. All neotheropods became extinct by the end of the Early Jurassic except for Averostra.

<i>Tawa hallae</i> Extinct species of dinosaur

Tawa is a genus of possible basal theropod dinosaurs from the Late Triassic period. The fossil remains of Tawa hallae, the type and only species were found in the Hayden Quarry of Ghost Ranch, New Mexico, US. Its discovery alongside the relatives of Coelophysis and Herrerasaurus supports the hypothesis that the earliest dinosaurs arose in Gondwana during the early Late Triassic period in what is now South America, and radiated from there around the globe. The specific name honours Ruth Hall, founder of the Ghost Ranch Museum of Paleontology.

<i>Megapnosaurus</i> Extinct genus of dinosaur

Megapnosaurus is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago during the early part of the Jurassic Period in what is now Africa. The species was a small to medium-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 2.2 m (7.2 ft) long and weigh up to 13 kg (29 lb). It was originally given the genus name Syntarsus, but that name was later determined to be preoccupied by a beetle. The species was subsequently given a new genus name, Megapnosaurus, by Ivie, Ślipiński & Węgrzynowicz in 2001. Some studies have classified it as a species within the genus Coelophysis, but this interpretation has been challenged by more subsequent studies and the genus Megapnosaurus is now considered valid.

<i>Coelophysis</i>? <i>kayentakatae</i> Extinct species of dinosaur

Coelophysis? kayentakatae is an extinct species of neotheropod dinosaur that lived approximately 200–196 million years ago during the early part of the Jurassic Period in what is now the southwestern United States. It was originally named Syntarsus kayentakatae, but the genus Syntarus was found to be preoccupied by a Colydiine beetle, so it was moved to the genus Megapnosaurus, and then to Coelophysis. A recent reassessment suggests that this species may require a new genus name.

Panguraptor is a genus of coelophysid theropod dinosaur known from fossils discovered in Lower Jurassic rocks of southern China. The type and only known species is Panguraptor lufengensis. The generic name refers to the deity Pangu but also to the supercontinent Pangaea for which in a geological context the same characters are used: 盘古. Raptor means "seizer", "robber" in Latin. The specific name is a reference to the Lufeng Formation. The holotype specimen was recovered on 12 October 2007 from the Lufeng Formation of Yunnan, which is noted for sauropodomorph fossils. It was described in 2014 by You Hai-Lu and colleagues.

<span class="mw-page-title-main">Timeline of coelophysoid research</span>

This timeline of coelophysoid research is a chronological listing of events in the history of paleontology focused on the coelophysoids, a group of primitive theropod dinosaurs that were among Earth's dominant predators during the Late Triassic and Early Jurassic epochs. Although formally trained scientists didn't discover coelophysoid fossils until the late 19th century, Native Americans of the modern southwestern United States may have already encountered their fossils. Navajo creation mythology describes the early Earth as being inhabited by a variety of different kinds of monsters who hunted humans for food. These monsters were killed by storms and the heroic Monster Slayers, leaving behind their bones. As these tales were told in New Mexico not far from bonebeds of Coelophysis, this dinosaur's remains may have been among the fossil remains that inspired the story.

Lepidus is a genus of extinct theropod from the Upper Triassic of the United States. It lived in the Otis Chalk localities of the Dockum Group in Texas, around 223 million years ago.

<i>Lucianovenator</i> Extinct genus of dinosaurs

Lucianovenator is an extinct genus of coelophysid theropod dinosaur which lived in Argentina during the Triassic. The genus name Lucianovenator translates to "Luciano's hunter", in reference to Don Luciano Leyes, who first reported the remains. The species name bonoi refers to Tulio del Bono, a local scientific authority who collaborated on the describers' research. It is one of the few neotheropods known from South America.

Powellvenator is an extinct genus of coelophysoid theropod dinosaur that lived during the latter part of the Triassic Period in what is now northwestern Argentina. Fossils of the dinosaur were found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin. The type species, Powellvenator podocitus, was named by Martin Ezcurra in 2017.

Martín Dário Ezcurra is an Argentine palaeontologist naming many extinct genera such as Aerotitan, Lophostropheus and Powellvenator.

<i>Notatesseraeraptor</i> Genus of reptiles (fossil)

Notatesseraeraptor is a genus of carnivorous theropod dinosaur that lived during the Late Triassic of what is now Switzerland. It was found in the Gruhalde Member of the Klettgau Formation. It was an early member of Neotheropoda with affinities to Dilophosaurus and Averostra. The new genus and species Notatesseraeraptor frickensis was named by Marion Zahner and colleagues in 2019.

References

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