Coelophysoidea

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Coelophysoids
Temporal range: Late Triassic-Early Jurassic, 228–183  Ma
Coelophysis bauri mount.jpg
Mounted skeleton of Coelophysis bauri , Cleveland Museum of Natural History
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Neotheropoda
Superfamily: Coelophysoidea
Nopcsa, 1928
Type species
Coelurus bauri
Cope, 1887
Subgroups
Synonyms
  • Podokesauroidea Huene, 1914

Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the coelurosaurs, with which they were formerly classified, and some species had delicate cranial crests. Sizes range from about 1 to 6 m in length. It is unknown what kind of external covering coelophysoids had, and various artists have portrayed them as either scaly or feathered. Some species may have lived in packs, as inferred from sites where numerous individuals have been found together.

Contents

Examples of coelophysoids include Coelophysis , Procompsognathus and Liliensternus . Most dinosaurs formerly referred to as being in the dubious taxon "Podokesauridae" are now classified as coelophysoids.

Classification

Despite their very early occurrence in the fossil record, coelophysoids have a number of derived features that separate them from primitive (basal) theropods. Among the most prominent of these derived features (apomorphies) is the way the upper jaw bones are connected (the premaxilla-maxilla articulation), which is flexible with a deep gap between the teeth in the two bones. A major source of disagreement among theropod experts is whether or not coelophysoids shared a more recent common ancestor with Ceratosauria (sensu stricto) than the ceratosaurs did with other theropods. Most recent analyses indicate the latter, that Coelophysoidea does not form a natural group with the ceratosaurians. Similarly, while Dilophosaurus and similar theropods have traditionally been classified as coelophysoids, several studies published in the late 2000s suggested that they may actually be more closely related to the tetanurans. [1]

The cladogram below was recovered in a study by Matthew T. Carrano, John R. Hutchinson and Scott D. Sampson, 2005. [2]

Coelophysoidea

Dilophosaurus Dilophosaurus wetherilli (flipped).PNG

Sarcosaurus

Gojirasaurus

Zupaysaurus Zupaysaurus.jpg

Coelophysidae

Procompsognathus Procompsognathus.jpg

Segisaurus Segisaurus.jpg

Liliensternus

Coelophysis Coelophysis size flipped.jpg

Megapnosaurus

The cladogram below was recovered in a study by Martin D. Ezcurra and Gilles Cuny, 2007. [3]

Liliensternus

Lophostropheus

  Coelophysidae  

"Megapnosaurus" kayentakatae

Coelophysis Coelophysis size flipped.jpg

Coelophysis rhodesiensis

The cladogram below was recovered in a study by Ezcurra et al. (2020). [4]

Coelophysoidea

Liliensternus

Dracoraptor

'Syntarsus' kayentakatae

Panguraptor

Powellvenator

Lepidus

Coelophysis bauri

Camposaurus

Lucianovenator

Megapnosaurus rhodesiensis

Segisaurus

See also

Related Research Articles

<i>Coelophysis</i> Genus of theropod dinosaurs from the late Triassic

Coelophysis is a genus of coelophysid theropod dinosaur that lived approximately 215 to 208.5 million years ago during the Late Triassic period from the middle to late Norian age in what is now the southwestern United States. Megapnosaurus was once considered to be a species within this genus, but this interpretation has been challenged since 2017 and the genus Megapnosaurus is now considered valid.

Cryolophosaurus is a genus of large theropod dinosaur known from only a single species Cryolophosaurus ellioti, from the early Jurassic of Antarctica. It was one of the largest theropods of the Early Jurassic, with the subadult being estimated to have reached 6–7 metres (20–23 ft) long and weighed 350–465 kilograms (772–1,025 lb).

<span class="mw-page-title-main">Ceratosauria</span> Extinct clade of dinosaurs

Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with Ceratosaurus than with birds. The oldest known ceratosaur, Saltriovenator, dates to the earliest part of the Jurassic, around 199 million years ago. Ceratosauria includes three major clades: Ceratosauridae, Noasauridae, and Abelisauridae, found primarily in the Southern Hemisphere. Originally, Ceratosauria included the above dinosaurs plus the Late Triassic to Early Jurassic Coelophysoidea and Dilophosauridae, implying a much earlier divergence of ceratosaurs from other theropods. However, most recent studies have shown that coelophysoids and dilophosaurids do not form a natural group with other ceratosaurs, and are excluded from this group.

<span class="mw-page-title-main">Tetanurae</span> Clade containing most theropod dinosaurs

Tetanurae is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans. Tetanurans are defined as all theropods more closely related to modern birds than to Ceratosaurus and contain the majority of predatory dinosaur diversity. Tetanurae likely diverged from its sister group, Ceratosauria, during the late Triassic. Tetanurae first appeared in the fossil record by the Early Jurassic about 190 mya and by the Middle Jurassic had become globally distributed.

<i>Procompsognathus</i> Extinct genus of dinosaurs

Procompsognathus is an extinct genus of coelophysid theropod dinosaur that lived approximately 210 million years ago during the later part of the Triassic Period, in what is now Germany. Procompsognathus was a small-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 1 m (3.3 ft) long.

<i>Afrovenator</i> Extinct genus of dinosaurs

Afrovenator is a genus of megalosaurid theropod dinosaur from the Middle or Late Jurassic Period on the Tiourarén Formation and maybe the Irhazer II Formation of the Niger Sahara region in northern Africa. Afrovenator represents the only properly identified Gondwanan megalosaur, with proposed material of the group present in the Late Jurassic on Tacuarembó Formation of Uruguay and the Tendaguru Formation of Tanzania.

<i>Segisaurus</i> Extinct genus of dinosaurs

Segisaurus is a genus of small coelophysid theropod dinosaur, that measured approximately 1 metre in length. The only known specimen was discovered in early Jurassic strata in Tsegi Canyon, Arizona, for which it was named. Segisaurus is the only dinosaur to have ever been excavated from the area.

<i>Liliensternus</i> Extinct genus of dinosaurs

Liliensternus is an extinct genus of basal neotheropod dinosaur that lived approximately 210 million years ago during the latter part of the Triassic Period in what is now Germany. Liliensternus was a moderate-sized, bipedal, ground-dwelling carnivore, that could grow up to 5.15 m (16.9 ft) long. It is the best represented Triassic theropod from Europe and one of the largest known.

<i>Gojirasaurus</i> Extinct genus of dinosaurs

Gojirasaurus is a potentially dubious genus of coelophysoid theropod dinosaur named after the giant monster movie character Godzilla.

<span class="mw-page-title-main">Coelophysidae</span> Extinct family of dinosaurs

Coelophysidae is a family of primitive carnivorous theropod dinosaurs. Most species were relatively small in size. The family flourished in the Late Triassic and Early Jurassic periods, and has been found on numerous continents. Many members of Coelophysidae are characterized by long, slender skulls and light skeletons built for speed. One member genus, Coelophysis, displays the earliest known furcula in a dinosaur.

<i>Sarcosaurus</i> Extinct genus of dinosaurs

Sarcosaurus is a genus of basal neotheropod dinosaur, roughly 3.5 metres (11 ft) long. It lived in what is now England and maybe Ireland and Scotland during the Hettangian-Sinemurian stages of the Early Jurassic, about 199-196 million years ago. Sarcosaurus is one of the earliest known Jurassic theropods, and one of only a handful of theropod genera from this time period. Along with Dracoraptor hanigani it is one of the two described neotheropods from the lowermost Jurassic of the United Kingdom.

<i>Zupaysaurus</i> Extinct genus of dinosaurs

Zupaysaurus is an extinct genus of early theropod dinosaur living during the Norian stage of the Late Triassic in what is now Argentina. Fossils of the dinosaur were found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina. Although a full skeleton has not yet been discovered, Zupaysaurus can be considered a bipedal predator, up to 4 metres (13 ft) long. It may have had two parallel crests running the length of its snout.

<i>Lukousaurus</i> Extinct species of reptile

Lukousaurus is an archosauromorph based on most of a small skull's snout, displaying distinctive lachrymal horns, found in the Early Jurassic-age Lower Lufeng Formation, Yunnan, China and was described by Chung Chien Young in 1940. The generic name refers to the Lugou Bridge, lit. “crossroads”, near Beijing, where the Sino-Japanese War started. L. yini is tentatively classified as a theropod dinosaur by some allied to ceratosaurs, by others a coelurosaur. Its skull is rather robust for its size though the teeth were described by the author as typically theropodan. Whatever Lukousaurus was, it was definitely an archosauromorph.

Lophostropheus is an extinct genus of coelophysoid theropod dinosaur that lived approximately 205.6 to 196.5 million years ago during the boundary between the Late Triassic Period and the Early Jurassic Period, in what is now Normandy, France. Lophostropheus is one of the few dinosaurs that may have survived the Triassic–Jurassic extinction event.

<span class="mw-page-title-main">Neotheropoda</span> Clade of theropods

Neotheropoda is a clade that includes coelophysoids and more advanced theropod dinosaurs, and is the only group of theropods that survived the Triassic–Jurassic extinction event. All neotheropods became extinct by the end of the Early Jurassic except for Averostra.

<span class="mw-page-title-main">Averostra</span> Clade of dinosaurs

Averostra, or "bird snouts", is a clade that includes most theropod dinosaurs, namely Ceratosauria and Tetanurae, and represent the only group of post-Early Jurassic theropods. Both survived into the Cretaceous period. When the Cretaceous–Paleogene extinction event occurred, ceratosaurians, megaraptorans an incertae sedis group within Tetanurae, and two groups of tetanurans within the clade Coelurosauria, the Tyrannosauroidea and Maniraptoriformes, were still extant. Only one subgroup of Maniraptoriformes, Aves, survived the extinction event and persisted to the present day.

<i>Megapnosaurus</i> Extinct genus of dinosaur

Megapnosaurus is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago during the early part of the Jurassic Period in what is now Africa. The species was a small to medium-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 2.2 m (7.2 ft) long and weigh up to 13 kg (29 lb). It was originally given the genus name Syntarsus, but that name was later determined to be preoccupied by a beetle. The species was subsequently given a new genus name, Megapnosaurus, by Ivie, Ślipiński & Węgrzynowicz in 2001. Some studies have classified it as a species within the genus Coelophysis, but this interpretation has been challenged by more subsequent studies and the genus Megapnosaurus is now considered valid.

<i>Coelophysis</i>? <i>kayentakatae</i> Extinct species of dinosaur

Coelophysis? kayentakatae is an extinct species of neotheropod dinosaur that lived approximately 200–196 million years ago during the early part of the Jurassic Period in what is now the southwestern United States. It was originally named Syntarsus kayentakatae, but the genus Syntarus was found to be preoccupied by a Colydiine beetle, so it was moved to the genus Megapnosaurus, and then to Coelophysis. A recent reassessment suggests that this species may require a new genus name.

<span class="mw-page-title-main">Timeline of coelophysoid research</span>

This timeline of coelophysoid research is a chronological listing of events in the history of paleontology focused on the coelophysoids, a group of primitive theropod dinosaurs that were among Earth's dominant predators during the Late Triassic and Early Jurassic epochs. Although formally trained scientists didn't discover coelophysoid fossils until the late 19th century, Native Americans of the modern southwestern United States may have already encountered their fossils. Navajo creation mythology describes the early Earth as being inhabited by a variety of different kinds of monsters who hunted humans for food. These monsters were killed by storms and the heroic Monster Slayers, leaving behind their bones. As these tales were told in New Mexico not far from bonebeds of Coelophysis, this dinosaur's remains may have been among the fossil remains that inspired the story.

Martín Dário Ezcurra is an Argentine palaeontologist naming many extinct genera such as Aerotitan, Lophostropheus and Powellvenator.

References

  1. Smith, N.D., Makovicky, P.J., Pol, D., Hammer, W.R., and Currie, P.J. (2007). "The dinosaurs of the Early Jurassic Hanson Formation of the Central Transantarctic Mountains: Phylogenetic review and synthesis." In Cooper, A.K. and Raymond, C.R. et al. (eds.), Antarctica: A Keystone in a Changing World––Online Proceedings of the 10th ISAES, USGS Open-File Report 2007-1047, Short Research Paper 003, 5 p.; doi : 10.3133/of2007-1047.srp003.
  2. Carrano, M.T, Hutchinson, J.R, Sampson, S.D. (2005). "New information on Segisaurus halli, a small theropod dinosaur from the Early Jurassic of Arizona." Journal of Vertebrate Paleontology. 25(4):835-849.
  3. Ezcurra, Martin D.; Cuny, Gilles (2007). "The coelophysoid Lophostropheus airelensis, gen. nov.: a review of the systematics of "Liliensternus" airelensis from the Triassic-Jurassic boundary outcrops of Normandy (France)". Journal of Vertebrate Paleontology. 27 (1): 73–86. doi:10.1671/0272-4634(2007)27[73:TCLAGN]2.0.CO;2. S2CID   85751238.
  4. Ezcurra, Martín D; Butler, Richard J; Maidment, Susannah C R; Sansom, Ivan J; Meade, Luke E; Radley, Jonathan D (2021-01-01). "A revision of the early neotheropod genus Sarcosaurus from the Early Jurassic (Hettangian–Sinemurian) of central England". Zoological Journal of the Linnean Society. 191 (1): 113–149. doi:10.1093/zoolinnean/zlaa054. hdl: 11336/160038 . ISSN   0024-4082.

Sources