Wiehenvenator Temporal range: Callovian ~ | |
---|---|
Remains of the skull | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Megalosauridae |
Subfamily: | † Megalosaurinae |
Genus: | † Wiehenvenator Rauhut et al, 2016 |
Type species | |
†Wiehenvenator albati Rauhut et al, 2016 |
Wiehenvenator is a genus of megalosaurid theropod dinosaur from the Middle Jurassic (Callovian) of north western Germany. The genus contains a single species, W. albati.
In 1998, geologist Friedrich Albat, prospecting for the Landschaftsverband Westfalen-Lippe Museum of Natural History, discovered the remains of a large theropod at the abandoned Pott quarry in the Wiehen Hills near Minden, Westphalia. The remains were discovered within the Ornatenton Formation, a geological formation composed mainly of mudstone, sandstones, and a horizon of carbonate concretions. The fragmentary theropod skeleton, found alongside abundant marine invertebrates and fossilized wood, was excavated between October 1998 and October 2001. At the time of their discovery, the bones were heavily weathered out of the surrounding sediments and are somewhat poorly preserved. The numerous breaks and cracks found in the material risked being destroyed upon removal from the matrix, and so led the excavation team to instead extract them into jackets that were then later prepared in the laboratories of the LWL Museum für Naturkunde. [1]
The find received extensive press attention and was informally known as Das Monster von Minden (the Minden Monster). Reports in the German edition of the National Geographic of a rib 50% larger than that of Allosaurus stirred speculations that it reached 15 metres (49 ft) in length. [2] Thomas Holtz estimated it at 12 metres (39 ft) in 2012. [3] Other researchers concluded to smaller dimensions: Mickey Mortimer in 2003 estimated the animal to be 7 to 8 metres (23 to 26 ft) in length and 0.75 to 1.2 metric tons (0.83 to 1.32 short tons) in weight. [4]
In 2015, it was announced that the find had been identified as a new species of megalosaurid. [5] In 2016, the fossils were named and described as the type species Wiehenvenator albati by Oliver W. M. Rauhut, Tom R. Hübner and Klaus-Peter Lanser. The generic name combines a reference to the Wiehengebirge, the German name of the Wiehen Hills, with a Latin venator ("hunter"). The specific name honours Friedrich Albat, the discoverer. As the name was published in an electronic publication, Life Science Identifiers were needed for its validity. These were 95638CFF-5618-4D31-9086-D821F6EE6B39 for the genus and 262FA776-9ABC-4565-9A17-931CB4BEFBFC for the species. [1]
A megalosauroid pedal phalanx ((WMNM P27698, P27693) from the Ornatenton Formation discovered in two parts during October 1999 and April 2000 may have belonged to the holotype of W. albati. [6]
The type specimen of Wiehenvenator consists of an assortment of bones found in the Ornatenton Formation dating from the middle Callovian. They include parts of the skull (right premaxilla, right maxilla, right lacrimal bone, right postorbital and possible front branch of the right quadratojugal), the anterior parts of a right lower jaw (dentary), six teeth, three tail vertebrae, a pair of fused median segments of rear gastralia, one complete rib and four rib fragments, a finger phalanx, both fibulae, a right astragalus and a right calcaneum. All these bones were seen as belonging to a single individual. Two additional tail vertebrae may also belong to it. Histological analysis of its fibulae suggests that the holotype of Wiehenvenator albati was at least in its ninth year of life, however, the age at death might have been well over ten years. The remains indicated that the animal was actively growing, but narrow growth zones indicated that the skeletal growth rate was slowing down. From this it can be determined that the growth state of Wiehenvenator was that of a large subadult individual. [1]
The length of Wiehenvenator can be estimated by extrapolating from its maxilla, which has 82 percent of the length of the maxilla of Torvosaurus gurneyi , itself estimated at 10 metres (33 ft). Another estimate can be gleaned from the fact that its tail vertebrae and fibulae are roughly equal in length to those of Torvosaurus tanneri, which had been estimated at 9 metres (30 ft), thus making Wiehenvenator one of the largest known European theropods. [1]
In 2016, Wiehenvenator was placed in the Megalosauridae as a sister taxon to Torvosaurus . The following is a cladogram based on the phylogenetic analysis conducted by Rauhut et al., showing the relationships of Wiehenvenator. [1]
After discovering the initial remains of Wiehenvenator, members of the excavation team returned to the site and continued to search the surroundings for further material. After searching 35m both east and west of the Ornatenton Formation, some weathered vertebral centra and teeth of Liopleurodon were found. [1] One year later, in mid-October 1999, the remains consisting of a maxillary fragment, bone fragments, and a tooth, of a second theropod were found 28.5 m north-west of the first locality. [1] On October 3, 2014, in an overgrown quarry to the west, the skull and lower jaws of the crocodylomorph Metriorhynchus were discovered by an honorary member of the LWL Museum für Naturkunde. These multiple discoveries imply a potential for more material to be found in the future. [1]
Allosaurus is an extinct genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period. The name "Allosaurus" means "different lizard", alluding to its unique concave vertebrae. It is derived from the Greek words ἄλλος and σαῦρος. The first fossil remains that could definitively be ascribed to this genus were described in 1877 by famed paleontologist Othniel Charles Marsh. As one of the first well-known theropod dinosaurs, it has long attracted attention outside of paleontological circles.
Ceratosaurus was a carnivorous theropod dinosaur that lived in the Late Jurassic period. The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is Ceratosaurus nasicornis.
Torvosaurus is a genus of large megalosaurine theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle and Late Jurassic period in what is now Colorado, Portugal, Germany, and possibly England, Spain, Tanzania, and Uruguay. It contains two currently recognized species, Torvosaurus tanneri and Torvosaurus gurneyi, plus a third unnamed species from Germany.
Afrovenator is a genus of megalosaurid theropod dinosaur from the Middle or Late Jurassic Period on the Tiourarén Formation and maybe the Irhazer II Formation of the Niger Sahara region in northern Africa. Afrovenator represents the only properly identified Gondwanan megalosaur, with proposed material of the group present in the Late Jurassic on Tacuarembó Formation of Uruguay and the Tendaguru Formation of Tanzania.
Eustreptospondylus is a genus of megalosaurid theropod dinosaur, from the Oxfordian stage of the Late Jurassic period in southern England, at a time when Europe was a series of scattered islands.
Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively primitive group of basal tetanurans containing two main subfamilies, Megalosaurinae and Afrovenatorinae, along with the basal genus Eustreptospondylus, an unresolved taxon which differs from both subfamilies.
Monolophosaurus is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China. It was named for the single crest on top of its skull. Monolophosaurus was a mid-sized theropod at about 5–5.5 metres (16–18 ft) long and weighed 475 kilograms (1,047 lb).
Sinosaurus is an extinct genus of theropod dinosaur which lived during the Early Jurassic Period. It was a bipedal carnivore approximately 5.5 metres (18 ft) in length and 300 kilograms (660 lb) in body mass. Fossils of the animal were found at the Lufeng Formation, in the Yunnan Province of China.
Chuandongocoelurus is a genus of carnivorous tetanuran theropod dinosaur from the Jurassic of China.
Dubreuillosaurus is a genus of carnivorous dinosaur from the middle Jurassic Period. It is a megalosaurid theropod. Its fossils were found in France. The only named species, Dubreuillosaurus valesdunensis, was originally described as a species of Poekilopleuron, Poekilopleuron? valesdunensis, which is still formally the type species of the genus. It was later renamed Dubreuillosaurus valesdunensis when, in 2005, Allain came to the conclusion that it was not part of the genus Poekilopleuron. Its type specimen, MNHN 1998-13, is only rivalled in the number of preserved elements in this group by that of Eustreptospondylus. Dubreuillosaurus is considered to be the sister species of Magnosaurus. It did not show signs of insular dwarfism even though it was uncovered on an island.
Genyodectes is a genus of ceratosaurian theropod dinosaur from the Lower Cretaceous (Aptian) of South America. The holotype material was collected from the Cerro Barcino Formation, Cañadón Grande, Departamento Paso de Indios in the Chubut Province of Argentina and consists of an incomplete snout, including the premaxillae, portions of both maxillas, the right and left dentary, many teeth, a fragment of the left splenial, and parts of the supradentaries. These elements are generally poorly preserved and some are in articulation. The premaxilla of Genyodectes possesses relatively large and protruding teeth, similar to those of Ceratosaurus. The specific name, serus, means "late". In 2016 it was estimated to be 6.25 meters in length and 790 kg in weight.
Kaijiangosaurus is a genus of carnivorous tetanuran theropod dinosaur from the Middle Jurassic of China. In 1984 He Xinlu named and described the type species Kaijiangosaurus lini. The generic name refers to the River (jiang) Kai. The specific name honours the paleontologist Lin Wenqiu.
Piveteausaurus is a genus of theropod dinosaur known from a partial skull discovered in the Middle Jurassic Marnes de Dives formation of Calvados, in northern France and lived about 164.7-161.2 million years ago. In 2012 Thomas Holtz gave a possible length of 11 meters.
Lourinhanosaurus was a genus of carnivorous theropod dinosaur that lived during the Late Jurassic Period (Kimmeridgian/Tithonian) in Portugal. It is one of many large predators discovered at the Lourinhã Formation and probably competed with coeval Torvosaurus gurneyi, Allosaurus europaeus, and Ceratosaurus.
Poekilopleuron is a genus of tetanuran dinosaur, which lived during the middle Bathonian of the Jurassic, about 168 to 166 million years ago. The genus has been used under many different spelling variants, although only one, Poekilopleuron, is valid. The type species is P. bucklandii, named after William Buckland, and many junior synonyms of it have also been erected. Little material is currently known, as the holotype was destroyed in World War II, although many casts of the material still exist.
Duriavenator is a genus of theropod dinosaur that lived in what is now England during the Middle Jurassic, about 168 million years ago. In 1882, upper and lower jaw bones of a dinosaur were collected near Sherborne in Dorset, and Richard Owen considered the fossils to belong to the species Megalosaurus bucklandii, the first named non-bird dinosaur. By 1964, the specimen was recognised as belonging to a different species, and in 1974 it was described as a new species of Megalosaurus, M. hesperis; the specific name means 'the West' or 'western'. Later researchers questioned whether the species belonged to Megalosaurus, in which many fragmentary theropods from around the world had historically been placed. After examining the taxonomic issues surrounding Megalosaurus, Roger B. J. Benson moved M. hesperis to its own genus in 2008, Duriavenator; this name means "Dorset hunter".
Leshansaurus is a genus of theropod dinosaur from the Late Jurassic Shaximiao Formation of what is now China. It was described in 2009 by a team of Chinese paleontologists. The type species is Leshansaurus qianweiensis. Fossils of Leshansaurus were discovered in strata from the Shangshaximiao Formation, a formation rich in dinosaur fossils. Li et al. referred this taxon to Sinraptoridae – a group of carnosaurian theropods, but it may belong to Megalosauridae instead.
Piatnitzkysauridae is an extinct family of megalosauroid or basal allosauroid dinosaurs. It only consists of three to four known dinosaur genera: Condorraptor, Marshosaurus, Piatnitzkysaurus and possibly Xuanhanosaurus. The most complete and well known member of this family is Piatnitzkysaurus, which also gives the family its name.
Aorun is a genus of carnivorous theropod dinosaur first discovered in 2006, with its scientific description published in 2013. It is one of the oldest known coelurosaurian dinosaurs and is estimated to have lived ~161.6 million years ago during the Late Jurassic Period. It is the fifth extinct theropod discovered from Wucaiwan, China.
The Ornatenton Formation is a Jurassic marine formation in Germany that is middle Callovian in age. The formation represents a shallow marine environment.
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