Kileskus Temporal range: Bathonian, | |
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Diagram showing fossil remains described in initial description | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Proceratosauridae |
Genus: | † Kileskus Averianov et al., 2010 |
Species: | †K. aristotocus |
Binomial name | |
†Kileskus aristotocus Averianov et al., 2010 | |
Kileskus (meaning lizard in the Khakas language) is a genus of tyrannosauroid dinosaur known from partial remains found in Middle Jurassic (Bathonian stage) Itat Formation of Sharypovsky District, Krasnoyarsk Krai (Russia). Fossils recovered include the holotype maxilla, a premaxilla, a surangular, and a few bones from the hand and foot. [1] Additional remains referred to the species include cervical and caudal vertebrae, as well as a fibula. [2] The skull bones are similar to those of Proceratosaurus . The type species is K. aristotocus. Kileskus was named in 2010 by Averianov and colleagues. Its size has been estimated at 5.2 meters (17 ft) in length and 700 kg (1,540 lbs) in weight. [3]
Kileskus has been included in two phylogenetic analyses and found to be a basal proceratosaurid both times. [1] [4]
Although it is unknown whether Kileskus sported a nasal crest, it can be assigned to Proceratosauridae due to a number of other features. These include elongated external nares, a short ventral margin of the premaxilla, and the area of the antorbital fossa directly below the antorbital fenestra being deeper than the maxilla directly below it. Kileskus also shares with Proceratosaurus nares inclined posterodorsally at a 40 degree angle to the skull. Kileskus is distinguished from other proceratosaurids by the anterior rim of its maxilla being confluent with the ascending process of the maxilla and gently sloping posterodorsally. [1]
Below is a cladogram published in 2013 by Loewen et al.. [4]
Cladogram published in 2018. [5]
Aviatyrannis is a genus of coelurosaurian theropod dinosaur, possibly a tyrannosauroid, from the Oxfordian-Tithonian stages of the Late Jurassic found in Portugal.
Coelurosauria is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Proceratosaurus is a genus of carnivorous theropod dinosaur from the Middle Jurassic (Bathonian) of England. It contains a single species. P. bradleyi, known from a mostly complete skull and lower jaws. Proceratosaurus was a small dinosaur, estimated to measure around 3 m (9.8 ft) in length. Its name refers to how it was originally thought to be an ancestor of Ceratosaurus, due to the partially preserved portion of the crest of Proceratosaurus superficially resembling the small crest of Ceratosaurus. Now, however, it is considered a coelurosaur, specifically a member of the family Proceratosauridae, and amongst the earliest known members of the clade Tyrannosauroidea.
Monolophosaurus is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China. It was named for the single crest on top of its skull. Monolophosaurus was a mid-sized theropod at about 5–5.5 metres (16–18 ft) long and weighed 475 kilograms (1,047 lb).
Stokesosaurus is a genus of small, carnivorous early tyrannosauroid theropod dinosaurs from the late Jurassic period of Utah, United States.
Tyrannosauroidea is a superfamily of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic Tyrannosaurus. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia.
Guanlong (冠龍) is a genus of extinct proceratosaurid tyrannosauroid dinosaur from the Late Jurassic of China. The taxon was first described in 2006 by Xu Xing et al., who found it to represent a new taxon related to Tyrannosaurus. The name is derived from Chinese, translating as "crown dragon". Two individuals are currently known, a partially complete adult and a nearly complete juvenile. These specimens come from the Oxfordian stage of the Chinese Shishugou Formation.
Bagaraatan is a genus of theropod dinosaur from the Late Cretaceous period. Its fossils were found in the Nemegt Formation of Mongolia. Bagaraatan may have been around 3 to 4 metres in length.
Shansisuchus is an extinct genus of archosauriform reptile belonging to the family Erythrosuchidae that lived during the Middle Triassic in what is now China. The first fossils of Shansisuchus were discovered from the Ermaying Formation of Shanxi (Shansi) province in 1964 by Chinese paleontologist Yang Zhongjian. Like other erythrosuchids, Shansisuchus was a large-bodied carnivore with a large, deep skull. Shansisuchus is unique among early archosauriforms in having a hole in its skull called a subnarial fenestra.
Turfanosuchus is a genus of archosauriform reptile, likely a gracilisuchid archosaur, which lived during the Middle Triassic (Anisian) of northwestern China. The type species, T. dabanensis, was described by C.C. Young in 1973, based on a partially complete but disarticulated fossil skeleton found in the Kelamayi Formation of the Turfan Basin.
Duriavenator is a genus of theropod dinosaur that lived in what is now England during the Middle Jurassic, about 168 million years ago. In 1882, upper and lower jaw bones of a dinosaur were collected near Sherborne in Dorset, and Richard Owen considered the fossils to belong to the species Megalosaurus bucklandii, the first named non-bird dinosaur. By 1964, the specimen was recognised as belonging to a different species, and in 1974 it was described as a new species of Megalosaurus, M. hesperis; the specific name means 'the West' or 'western'. Later researchers questioned whether the species belonged to Megalosaurus, in which many fragmentary theropods from around the world had historically been placed. After examining the taxonomic issues surrounding Megalosaurus, Roger B. J. Benson moved M. hesperis to its own genus in 2008, Duriavenator; this name means "Dorset hunter".
Xiongguanlong is an extinct genus of tyrannosauroid theropod from the Early Cretaceous period of what is now China. The type and only species is X. baimoensis. The generic name comes from Jiayuguan City and the Mandarin word "long" which means dragon. The specific epithet, "baimoensis" is a latinization of the Mandarin word for "white ghost" in reference to one of the geological features of the type locality.
Sinotyrannus is a genus of large basal tyrannosauroid dinosaur, known from a single incomplete fossil specimen including a partial skull, from the Early Cretaceous Jiufotang Formation of Liaoning, China. Specifically, it is a member of the Proceratosauridae, a family that originated in the Jurassic whose members are known from Europe and Asia. Though it is not significantly younger than primitive tyrannosauroids such as Dilong, it is similar in size to later forms such as Tyrannosaurus. It was much larger than contemporary tyrannosauroids, reaching a total estimated length of 9–10 m (30–33 ft), and was the largest known theropod from the Jiufotang Formation. The type species, S. kazuoensis, was described by Ji et al. in 2009.
Proceratosauridae is a family or clade of tyrannosauroid theropod dinosaurs from the Middle Jurassic to the Early Cretaceous.
Xixiasaurus is a genus of troodontid dinosaur that lived during the Late Cretaceous Period in what is now China. The only known specimen was discovered in Xixia County, Henan Province, in central China, and became the holotype of the new genus and species Xixiasaurus henanensis in 2010. The names refer to the areas of discovery, and can be translated as "Henan Xixia lizard". The specimen consists of an almost complete skull, part of the lower jaw, and teeth, as well as a partial right forelimb.
The Itat Formation is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from the formation, including the proceratosaurid Kileskus, as well as fish, amphibians, mammals and many other vertebrates. The formation is noted for bearing significant coal reserves, with large open pit coal mines extracting lignite from the unit currently in operation.
Sineleutherus is an extinct genus of euharamiyids which existed in Asia during the Jurassic period. The type species is Sineleutherus uyguricus, which was described by Thomas Martin, Alexander O. Averianov and Hans-Ulrich Pfretzschner in 2010; it lived in what is now China during the late Jurassic Qigu Formation. A second species, Sineleutherus issedonicus, was described by A. O. Averianov, A. V. Lopatin and S. A. Krasnolutskii in 2011. It lived in what is now Sharypovsky District during the middle Jurassic ; its fossils were collected from the upper part of the Itat Formation. However, this is now believed to represent several euharamiyid taxa not closey related to Sineleutherus.
Zuolong is an extinct genus of tetanuran theropod from the Late Jurassic period of China. The type and only species is Z. salleei. The generic name of Zuolong is in honor of General Zu Zōngtáng with the Chinese word "long" which means dragon. The specific epithet "salleei" is in honor of Hilmar Sallee, who funded the expedition which led to the specimen's discovery.
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Polymorphodon is an extinct genus of archosauriform reptile from the Middle Triassic of Germany. The only known species is Polymorphodon adorfi, discovered in Lower Keuper deposits at a quarry in Eschenau, Germany. Polymorphodon is notable for its heterodont dentition, with long and conical premaxillary teeth followed by thin maxillary teeth with large serrations. Maxillary teeth near the back of the mouth are short and leaf-shaped, similar to some living and extinct reptiles with a herbivorous or omnivorous diet. This may suggest that Polymorphodon had some reliance on plants in its diet, a rarity among basal archosauriforms, most of which are carnivores.