Pycnonemosaurus Temporal range: Late Cretaceous ~ | |
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Skeletal restoration showing known material | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Abelisauridae |
Clade: | † Furileusauria |
Genus: | † Pycnonemosaurus Kellner & Campos, 2002 |
Species: | †P. nevesi |
Binomial name | |
†Pycnonemosaurus nevesi Kellner & Campos, 2002 | |
Synonyms | |
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Pycnonemosaurus (meaning 'dense forest lizard') is a genus of carnivorous theropod dinosaur that belonged to the family Abelisauridae. It was found in the Upper Cretaceous red conglomerate sandstones of the Cachoeira do Bom Jardim Formation, Mato Grosso, Brazil, and it lived during the late Campanian to early Maastrichtian stage of the Late Cretaceous. [1]
The type species, Pycnonemosaurus nevesi, was scientifically described by Kellner and Campos in 2002. [2] [3] The only known specimen (DGM 859-R) was found in a red conglomeratic sandstone at the Jangada Roncador locality (then known as Fazenda Roncador), in Mato Grosso State that is exposed close to Paulo Creek, which is currently referred to the Cachoeira do Bom Jardim Formation. During 1952-1953, Llewellyn Ivor Price visited a farm named "Roncador" in the state of Mato Grosso and collected several dinosaur bones. These remains were found by the owner of the farm, Max de Barros Erhart, and his hired workers at the Paulo Creek site. The most important specimen found was the incomplete skeleton of a large abelisaurid theropod, which was found near several titanosaurid sauropod bones. [3] [4]
Pycnonemosaurus nevesi was named from the Greek word pycnós meaning dense, némos meaning pastures and woods, and saûrus meaning reptile or lizard. This naming was an allusion to Mato Grosso State, where the remains were found. The specific name was named after the late Dr. Iedo Batista Neves, who passed in 2000, who encouraged the pursuit of paleontological studies, particularly of Alexander Kellner. [3]
Thus far, the remains of Pycnonemosaurus have been fragmentary. No elements were well preserved, and the bone surface is well abraded that indicates the elements were partially exposed at the discovery location before being collected. [3] The type specimen, housed at the Earth Sciences Museum, Rio de Janeiro, consists of five incomplete teeth, parts of seven caudal vertebrae, the distal part of a right pubis, a right tibia, and the distal articulation of the right fibula. The small pubic foot and hatchet-shaped cnemial crest of the tibia distinguishes this species within the abelisaurs. The caudal vertebra has distinct abelisaurid features, such as a fan-shaped transverse process and a cranial projection. However, these awl-like projections are somewhat unlike related abelisaurids, such as Aucasaurus , in that they diminish more towards the distal caudals. All remains were found associated and are presently regarded as belonging to the same individual. [3]
The type specimen of Pycnonemosaurus belongs to an immature individual of uncertain ontogenetic stage, probably a subadult. [1] Initial size estimates of this specimen put the animal at 6–7 metres (20–23 ft) in length and 1.2 tonnes (1.3 short tons) in weight, [3] [2] but later analyses have found that it was likely larger, being about 8.9 metres (29.2 ft) long. [5] This new size estimate currently makes Pycnonemosaurus the largest formally described member of the Abelisauridae thus far. Another estimation gave a length of 9 metres (29.5 ft) and a weight of 3.6 tonnes (4.0 short tons). [6]
The following cladogram follows an analysis of 2021 during the description of Llukalkan . [7]
Pycnonemosaurus is the best known abelisaurid from Brazil, where most theropod material is presently rare besides preserved teeth and footprints. Even though only a few species are known from Brazil, it is one of the most informative countries concerning the Lower Cretaceous period. Initially, the deposits in which this dinosaur was found were assigned to the Bauru Group, and later more specifically to the Cambambe Formation, with a possible temporal range spanning from the Campanian to the Maastrichtian stages. [3] [8] [9] However, these deposits have since been reassigned to the Campanian-aged Cachoeira do Bom Jardim Formation. [10] This formation is believed to have been deposited in a semi-arid climate, representing the intermediate parts of alluvial fans. [4] [11]
Pycnonemosaurus was a predator, probably preying on titanosaurs found in the same locality. [4] Its teeth were small yet sharp, and were used to get a hold of struggling prey as the abelisaur shook and tore them apart. [3]
Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with Ceratosaurus than with birds. The oldest known ceratosaur, Saltriovenator, dates to the earliest part of the Jurassic, around 199 million years ago. Ceratosauria includes three major clades: Ceratosauridae, Noasauridae, and Abelisauridae, found primarily in the Southern Hemisphere. Originally, Ceratosauria included the above dinosaurs plus the Late Triassic to Early Jurassic Coelophysoidea and Dilophosauridae, implying a much earlier divergence of ceratosaurs from other theropods. However, most recent studies have shown that coelophysoids and dilophosaurids do not form a natural group with other ceratosaurs, and are excluded from this group.
Abelisaurus is a genus of predatory abelisaurid theropod dinosaur alive during the Late Cretaceous Period (Campanian) of what is now South America. It was a bipedal carnivore that probably reached about 7.4 metres in length, although this is uncertain as it is known from only one partial skull.
Xenotarsosaurus is a genus of abelisaurid theropod dinosaur that lived during the Late Cretaceous of Argentina.
Rugops is a monospecific genus of basal abelisaurid theropod dinosaur from Niger that lived during the Late Cretaceous period in what is now the Echkar Formation. The type and only species, Rugops primus, is known only from a partial skull. It was named and described in 2004 by Paul Sereno, Jeffery Wilson and Jack Conrad. Rugops has an estimated length of 4.4–5.3 metres (14–17 ft) and weight of 410 kilograms (900 lb). The top of its skull bears several pits which correlates with overlaying scale and the front of the snout would have had an armour-like dermis.
Aucasaurus is a genus of medium-sized abelisaurid theropod dinosaur from Argentina that lived during the Late Cretaceous of the Anacleto Formation. It was smaller than the related Carnotaurus, although more derived in some ways, such as its extremely reduced arms and almost total lack of fingers. The type skeleton is complete to the thirteenth caudal vertebra, and so is relatively well understood, and is the most complete abelisaurid yet described. However, the skull is damaged, causing some paleontologists to speculate that it was involved in a fight prior to death.
Ilokelesia is an extinct genus of abelisaurid theropod, preserved in the layers of the earliest Late Cretaceous of the Huincul Formation, Neuquén Group, located near Plaza Huincul, Neuquén Province, Argentina. The specimen, consisting of very fragmentary elements of the skull and the axial and appendicular skeleton, was described by Rodolfo Coria and Leonardo Salgado in late 1998.
Quilmesaurus is a genus of carnivorous abelisaurid theropod dinosaur from the Patagonian Upper Cretaceous of Argentina. It was a member of Abelisauridae, closely related to genera such as Carnotaurus. The only known remains of this genus are leg bones which share certain similarities to a variety of abelisaurids. However, these bones lack unique features, which may render Quilmesaurus a nomen vanum.
Majungasaurus is a genus of abelisaurid theropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period, making it one of the last-known non-avian dinosaurs that went extinct during the Cretaceous–Paleogene extinction event. The genus contains a single species, Majungasaurus crenatissimus. This dinosaur is also called Majungatholus, a name which is considered a junior synonym of Majungasaurus.
Ekrixinatosaurus is a genus of abelisaurid theropod which lived approximately 100 to 97 million years ago during the Late Cretaceous period. Its fossils have been found in Argentina. Only one species is currently recognized, Ekrixinatosaurus novasi, from which the specific name honors of Dr. Fernando Novas for his contributions to the study of abelisaurid theropods, while the genus name refers to the dynamiting of the holotype specimen. It was a large abelisaur, measuring between 6.5 and 8 m in length and weighing 800 kg (1,800 lb).
Spinostropheus is a genus of carnivorous neotheropod theropod dinosaur that lived in the Middle Jurassic period and has been found in the Tiouraren Formation, Niger. The type and only species is S. gautieri.
Skorpiovenator is a genus of abelisaurid theropod dinosaur from the Late Cretaceous Huincul Formation of Argentina. It is one of the most complete and informative abelisaurids yet known, described from a nearly complete and articulated skeleton.
Brachyrostra is a clade within the theropod dinosaur family Abelisauridae. It includes the famous genera Carnotaurus, Abelisaurus, Aucasaurus as well as their close relatives from the Cretaceous Period of Argentina and Brazil plus Caletodraco from France. The group was first proposed in an analysis conducted by Juan Canale and colleagues in 2008. They found that all South American abelisaurids described up to that point grouped together as a sub-clade of Abelisauridae, which they named based on the relatively unusual shape of their skulls. They defined the clade Brachyrostra as "all the abelisaurids more closely related to Carnotaurus sastrei than to Majungasaurus crenatissimus."
Rahiolisaurus is a genus of abelisaurid theropod dinosaur which existed in India during the Late Cretaceous period. It was described in 2010, based on fossils recovered from the Lameta Formation in the Indian state of Gujarat. These fossils include elements from at least seven different individuals and are believed to have been from the Maastrichtian stage, sometime between 70 and 66 million years ago, making it one of the last non-avian dinosaurs known in the fossil record. Despite representing a variety of different growth stages, all recovered fossils from the locality indicate a single species, the type species Rahiolisaurus gujaratensis.
Oxalaia is a genus of spinosaurid dinosaur that lived in what is now the Northeast Region of Brazil during the Cenomanian stage of the Late Cretaceous period, sometime between 100.5 and 93.9 million years ago. Its only known fossils were found in 1999 on Cajual Island in the rocks of the Alcântara Formation, which is known for its abundance of fragmentary, isolated fossil specimens. The remains of Oxalaia were described in 2011 by Brazilian palaeontologist Alexander Kellner and colleagues, who assigned the specimens to a new genus containing one species, Oxalaia quilombensis. The species name refers to the Brazilian quilombo settlements. Oxalaia quilombensis is the eighth officially named theropod species from Brazil and the largest carnivorous dinosaur discovered there. It is closely related to the African genus Spinosaurus, and/or may be a junior synonym of this taxon.
Eoabelisaurus is a genus of abelisauroid theropod dinosaur from the Lower Jurassic Cañadón Asfalto Formation of the Cañadón Asfalto Basin in Argentina, South America. The generic name combines a Greek ἠώς, (eos), "dawn", with the name Abelisaurus, in reference to the fact it represents an early relative of the latter. Only one species is currently recognized, E. mefi; the specific name honours the MEF, the Museo Paleontológico "Egidio Feruglio", where discoverer Diego Pol is active. It is characterized by reduced forelimb proportions that show primitive characteristics of the Abelisauridae family.
Arcovenator is an extinct genus of abelisaurid theropod dinosaurs hailing from the Late Cretaceous of France and possibly Spain. The type and only described species is Arcovenator escotae.
Majungasaurinae is a subfamily of large carnivorous theropods from the Upper Cretaceous, found in Madagascar, India, and France. It is a subgroup within the theropod family Abelisauridae, a Gondwanan clade known for their thick and often horned skulls and vestigial arms. The two subfamilies of Abelisauridae are Carnotaurinae, best known from the South American Carnotaurus, and Majungasaurinae, consisting of Madagascar’s Majungasaurus and its closest relatives. Their ancestors emerged in the Middle Jurassic, and the clade lasted until the Upper Cretaceous.
This timeline of ceratosaur research is a chronological listing of events in the history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became the apex predators of the southern hemisphere during the Late Cretaceous. The nature and taxonomic composition of the Ceratosauria has been controversial since the group was first distinguished in the late 19th century. In 1884 Othniel Charles Marsh described the new genus and species Ceratosaurus nasicornis from the Late Jurassic Morrison Formation of the western United States. He felt that it belonged in a new family that he called the Ceratosauridae. He created the new taxon Ceratosauria to include both the Ceratosauridae and the ostrich-like ornithomimids. The idea of the Ceratosauria was soon contested, however. Later that same decade both Lydekker and Marsh's hated rival Edward Drinker Cope argued that the taxon was invalid.
Tralkasaurus is a genus of abelisaurid dinosaur from the Huincul Formation from Río Negro Province in Argentina. The type and only species is Tralkasaurus cuyi, named in 2020 by Mauricio Cerroni and colleagues based on an incomplete skeleton. A medium-sized abelisaurid, Tralkasaurus exhibits a conflicting blend of characteristics found among the early-diverging abelisauroids with others that characterize the highly specialized clade Brachyrostra, and thus its position within the clade is poorly-resolved.
Spectrovenator is a genus of basal abelisaurid theropod dinosaur that lived during the early Cretaceous period in what is now Brazil. It is known by a single species, S. ragei, recovered from the Quiricó Formation.