Koleken Temporal range: Maastrichtian, ~ | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Abelisauridae |
Clade: | † Brachyrostra |
Clade: | † Furileusauria |
Tribe: | † Carnotaurini |
Genus: | † Koleken |
Species: | †K. inakayali |
Binomial name | |
†Koleken inakayali | |
Koleken (meaning "coming from clay and water") is a genus of carnotaurin abelisaurid from the Maastrichtian La Colonia Formation in the Chubut Province of Argentina. The type and only species is K. inakayali, known from one immature specimen about six years old in minimum age. [1]
Koleken is known from only the holotype MPEF-PV 10826 which was initially found and reported in 2015, [2] but was not described as K. inakayali until 2024. The holotype, consists of "closely associated (but disarticulated) remains of the skull and atlas, as well as the articulated postcranial skeleton composed of the posterior most eight dorsal vertebrae, a complete sacrum, eight caudal vertebrae, an almost complete pelvis and hind limbs". It differs from the larger Carnotaurus in anatomical characteristics found in the skull, vertebrae, and leg bones. The specimen likewise is believed to represent a sub-adult based on a lack of fusion in some of the bones. [1]
The generic name, Koleken (IPA: [kɒːlɛkɛn] ), is a name in Teushen spoken by the native population of central Patagonia that means "coming from clay and water", given the specimen was found in a sedimentary section dominated by claystone representing an estuarine environment. The specific name, inakayali (IPA: [ɪnækæjælaɪ] ), honours Inakayal, one of the last chiefs of Tehuelches, native people from central Patagonia. [1]
Koleken is placed in the Carnotaurini tribe of Abelisauridae, in a polytomy with the other carnotaurins in a phylogenetic analysis: [1]
Koleken was found in outcrops of the La Colonia Formation, which dates to the Maastrichtian (Cretaceous)–Paleocene (Paleogene) boundary. [3] This formation is best known for fossils of the abelisaurid theropod Carnotaurus and the saltasauroid titanosaur Titanomachya , [1] [4] as well as fossils of other reptiles including the turtle Patagoniaemys , the snake Alamitophis and the plesiosaurs Kawanectes , Chubutinectes and Sulcusuchus . [2] [5] [6] [7] Mammals discovered from the formation include Reigitherium , Coloniatherium , Argentodites and Ferugliotherium . [8] [9] Remains of an enantiornithine and, possibly, of a neornithine bird have been discovered. [10] [11]
Carnotaurus is a genus of theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 72 and 69 million years ago. The only species is Carnotaurus sastrei. Known from a single well-preserved skeleton, it is one of the best-understood theropods from the Southern Hemisphere. The skeleton, found in 1984, was uncovered in the Chubut Province of Argentina from rocks of the La Colonia Formation. Carnotaurus is a derived member of the Abelisauridae, a group of large theropods that occupied the large predatorial niche in the southern landmasses of Gondwana during the late Cretaceous. Within the Abelisauridae, the genus is often considered a member of the Brachyrostra, a clade of short-snouted forms restricted to South America.
Aucasaurus is a genus of medium-sized abelisaurid theropod dinosaur from Argentina that lived during the Late Cretaceous of the Anacleto Formation. It was smaller than the related Carnotaurus, although more derived in some ways, such as its extremely reduced arms and almost total lack of fingers. The type skeleton is complete to the thirteenth caudal vertebra, and so is relatively well understood, and is the most complete abelisaurid yet described. However, the skull is damaged, causing some paleontologists to speculate that it was involved in a fight prior to death.
Thalassomedon is a genus of plesiosaur, named by Welles in 1943.
Quilmesaurus is a genus of carnivorous abelisaurid theropod dinosaur from the Patagonian Upper Cretaceous of Argentina. It was a member of Abelisauridae, closely related to genera such as Carnotaurus. The only known remains of this genus are leg bones which share certain similarities to a variety of abelisaurids. However, these bones lack unique features, which may render Quilmesaurus a nomen vanum.
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae.
Hydrotherosaurus is an extinct genus of elasmosaurid plesiosaur from the Upper Cretaceous Moreno Formation of Fresno County, California, USA. The only known species, H. alexandrae, was named for Annie Montague Alexander in 1943 by Samuel Paul Welles.
Tuarangisaurus is an extinct genus of elasmosaurid known from New Zealand. The type and only known species is Tuarangisaurus keyesi, named by Wiffen and Moisley in 1986.
Aphrosaurus was an extinct genus of plesiosaur from the Maastrichtian. The type species is Aphrosaurus furlongi, named by Welles in 1943. The holotype specimen was discovered in the Moreno Formation in Fresno County, California in 1939 by rancher Frank C. Piava. A second specimen - LACM 2832 - was also found in the same formation and initially diagnosed as a juvenile of the same species, but has since been removed from the genus.
Aristonectes is an extinct genus of large elasmosaurid plesiosaurs that lived during the Maastrichtian stage of the Late Cretaceous. Two species are known, A. parvidens and A. quiriquinensis, whose fossil remains were discovered in what are now Patagonia and Antarctica. Throughout the 20th century, Aristonectes was a difficult animal for scientists to analyze due to poor fossil preparation, its relationships to other genera were uncertain. After subsequent revisions and discoveries carried out from the beginning of the 21st century, Aristonectes is now recognised as the type genus of the subfamily Aristonectinae, a lineage of elasmosaurids characterized by an enlarged skull and a reduced length of the neck.
The La Colonia Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.
Bonapartenykus is a monospecific genus of alvarezsauroid dinosaur from Argentina that lived during the Late Cretaceous (Campanian-Maastrichtian) in what is now the upper Allen Formation of the Río Negro Province. The type and only species, Bonapartenykus ultimus, is known from a nearly articulated but partial skeleton that was found in close association to two incomplete eggs and several clusters of eggshells belonging to the oogenus Arriagadoolithus. Bonapartenykus was named in 2012 by Federico L. Agnolin, Jaime E. Powell, Fernando E. Novas and Martin Kundrát. Bonapartenykus has an estimated length of 2.5 m (8.2 ft) and weight of 72 kg (159 lb), making it the largest member of the clade Alvarezsauroidea.
Vegasaurus is an extinct genus of elasmosaurid plesiosaur known from the Late Cretaceous Snow Hill Island Formation of Vega Island, Antarctic Peninsula. It contains a single species, Vegasaurus molyi.
Kawanectes is a genus of elasmosaurid plesiosaur, a type of long-necked marine reptile, that lived in the marginal marine environment of Late Cretaceous Patagonia. It contains one species, K. lafquenianum, described in 2016 by O'Gorman.
Tralkasaurus is a genus of abelisaurid dinosaur from the Huincul Formation from Río Negro Province in Argentina. The type and only species is Tralkasaurus cuyi, named in 2020 by Mauricio Cerroni and colleagues based on an incomplete skeleton. A medium-sized abelisaurid, Tralkasaurus exhibits a conflicting blend of characteristics found among the early-diverging abelisauroids with others that characterize the highly specialized clade Brachyrostra, and thus its position within the clade is poorly-resolved.
The Dorotea Formation is a geological formation in the Río de Las Chinas Valley of the Magallanes Basin in Patagonian Chile whose strata date back to the Campanian to Maastrichtian of the Late Cretaceous.
Niebla is a genus of abelisaurid theropod dinosaur from the Late Cretaceous Period (Campanian-Maastrichtian) of Río Negro province, Argentina. The genus contains a single species, Niebla antiqua, and is known from a partial, non-articulated skeleton. The holotype, found in the Allen Formation, represents an adult individual about nine years old in minimum age.
Kookne is a prehistoric bird genus from the Late Cretaceous. Known from a coracoid, the remains of the only known species Kookne yeutensis were found in rocks from the Late Cretaceous (Maastrichtian) Chorrillo Formation of Santa Cruz, Argentina.
Chubutinectes is an extinct genus of elasmosaurid plesiosaur from the Late Cretaceous La Colonia Formation of Argentina. The genus contains a single species, C. carmeloi, known from a partial skeleton and associated gastroliths.
Titanomachya is an extinct genus of titanosaurian sauropod dinosaur from the Late Cretaceous La Colonia Formation of Argentina. The genus contains a single species, T. gimenezi. It is a relatively small titanosaur, weighing around 7.8 tonnes.