Beishanlong

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Beishanlong
Temporal range: Late Aptian, 123–113  Ma
Skeleton of Beishanlong grandis.JPG
Restored skeleton of Beishanlong and Xiongguanlong
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Ornithomimosauria
Family: Deinocheiridae
Genus: Beishanlong
Makovicky et al., 2010
Species:
B. grandis
Binomial name
Beishanlong grandis
Makovicky et al., 2010

Beishanlong is a genus of giant ornithomimosaurian theropod dinosaur from the Early Cretaceous of China. It is the second-largest ornithomimosaur discovered, only surpassed by Deinocheirus.

Contents

Discovery and naming

Restoration Beishanlong grandis.jpg
Restoration

Three fossils of Beishanlong were in the early twenty-first century found in northwestern China at the White Ghost Castle site, in the province of Gansu. The type species is Beishanlong grandis, described and named online in 2009 by a team of Chinese and American paleontologists, and formally published in January 2010 by the same Peter Makovicky, Li Daiqing, Gao Keqin, Matthew Lewin, Gregory Erickson and Mark Norrell. The generic name combines a references to the Bei Shan, the "North Mountains", with a Chinese long, "dragon". The specific name means "large" in Latin, in reference to the body size. [1]

Beishanlong lived in the late Aptian [2] stage, with its fossils being uncovered in layers of the Xinminpu Group, in the Xiagou Formation. The holotype is FRDC-GS GJ (06) 01-18, found in 2006, consisting of a partial skeleton lacking the skull. [1] The paratypes consist of two specimens found in 2007: one consisting of remains of hindlimbs, the other, FRDC-GS JB(07)01-01, being a pair of pubes. A fourth fossil found in 1999, IVPP V12756 consisting of foot bones, was tentatively referred to the species. [1]

Description

Size comparison Beishanlong Scale.svg
Size comparison

Beishanlong is of a considerable size, approximating the largest-known individuals of Gallimimus , which have been estimated to reach eight metres. According to the description, Beishanlong "is one of the largest definitive ornithomimosaurs yet described, though histological analysis shows that the holotype individual was still growing at its death." A histological study of the bone structure of the fibula found thirteen or fourteen growth lines, indicating the individual was subadult, though growth had already slowed. [1] The size of this subadult individual is estimated at 5.9–7 m (19–23 ft) in length and 375–626 kilograms (827–1,380 lb) in body mass. [3] [1] [4] [5]

The build of Beishanlong was rather robust. The arms and legs were long, though lacking the extremely elongated hands, feet and claws of later forms. [1]

Classification

Beishanlong was by the describers assigned to the Ornithomimosauria, in a more basal position. Beishanlong was closely related to fellow ornithomimosaurian Harpymimus . Together they formed a polytomy with the main ornithomimosaurian branch just below Garudimimus . [1] In 2014 Yuong-Nam Lee et al. recovered Beishanlong as a member of Deinocheiridae basal to a clade containing Garudimimus and Deinocheirus . [6]

See also

Related Research Articles

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Gallimimus is a genus of theropod dinosaur that lived in what is now Mongolia during the Late Cretaceous period, about seventy million years ago (mya). Several fossils in various stages of growth were discovered by Polish-Mongolian expeditions in the Gobi Desert of Mongolia during the 1960s; a large skeleton discovered in this region was made the holotype specimen of the new genus and species Gallimimus bullatus in 1972. The generic name means "chicken mimic", referring to the similarities between its neck vertebrae and those of the Galliformes. The specific name is derived from bulla, a golden capsule worn by Roman youth, in reference to a bulbous structure at the base of the skull of Gallimimus. At the time it was named, the fossils of Gallimimus represented the most complete and best preserved ornithomimid material yet discovered, and the genus remains one of the best known members of the group.

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<span class="mw-page-title-main">Ornithomimosauria</span> Extinct clade of theropod dinosaurs

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<i>Dromiceiomimus</i> Extinct genus of reptiles

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<i>Deinocheirus</i> Genus of theropod dinosaurs

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<i>Arkansaurus</i> Extinct genus of dinosaurs

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<i>Garudimimus</i> Ornithomimosaur genus from the Late Cretaceous

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<i>Archaeornithomimus</i> Extinct genus of dinosaurs

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<i>Harpymimus</i> Extinct genus of dinosaurs

Harpymimus is a basal ornithomimosaurian theropod dinosaur from the Early Cretaceous Period of what is now Mongolia. Unlike later, more derived ornithomimosaurs, Harpymimus still possessed teeth, although they appear to have been restricted to the dentary of the lower jaw.

<span class="mw-page-title-main">Bayan Shireh Formation</span> Geological formation in Mongolia

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<i>Nqwebasaurus</i> Extinct genus of dinosaur

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<span class="mw-page-title-main">Deinocheiridae</span> Extinct family of dinosaurs

Deinocheiridae is a family of ornithomimosaurian dinosaurs, living in Asia and the Americas from the Albian until the Maastrichtian. The family was originally named by Halszka Osmólska and Roniewicz in 1970, including only the type genus Deinocheirus. In a 2014 study by Yuong-Nam Lee and colleagues and published in the journal Nature, it was found that Deinocheiridae was a valid family. Lee et al. found that based on a new phylogenetic analysis including the recently discovered complete skeletons of Deinocheirus, the type genus, as well as Garudimimus and Beishanlong, could be placed as a successive group, with Beishanlong as the most primitive and Deinocheirus as most derived. The family Garudimimidae, named in 1981 by Rinchen Barsbold, is now a junior synonym of Deinocheiridae as the latter family includes the type genus of the former. The group existed from 115 to 69 million years ago, with Beishanlong living from 115 to 100 mya, Garudimimus living from 98 to 83 mya, and Deinocheirus living from 71 to 69 mya. Other genera included are Paraxenisaurus, and possibly Harpymimus and Hexing.

<i>Suzhousaurus</i> Extinct genus of dinosaurs

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<i>Xiongguanlong</i> Extinct genus of dinosaurs

Xiongguanlong is an extinct genus of tyrannosauroid theropod from the Early Cretaceous period of what is now China. The type and only species is X. baimoensis. The generic name comes from Jiayuguan City and the Mandarin word "long" which means dragon. The specific epithet, "baimoensis" is a latinization of the Mandarin word for "white ghost" in reference to one of the geological features of the type locality.

Hexing is an extinct genus of basal ornithomimosaur dinosaur known from the Early Cretaceous of northeastern China. It contains a single species, Hexing qingyi.

<span class="mw-page-title-main">Timeline of ornithomimosaur research</span>

This timeline of ornithomimosaur research is a chronological listing of events in the history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Although fragmentary, probable, ornithomimosaur fossils had been described as far back as the 1860s, the first ornithomimosaur to be recognized as belonging to a new family distinct from other theropods was Ornithomimus velox, described by Othniel Charles Marsh in 1890. Thus the ornithomimid ornithomimosaurs were one of the first major Mesozoic theropod groups to be recognized in the fossil record. The description of a second ornithomimosaur genus did not happen until nearly 30 years later, when Henry Fairfield Osborn described Struthiomimus in 1917. Later in the 20th century, significant ornithomimosaur discoveries began occurring in Asia. The first was a bonebed of "Ornithomimus" asiaticus found at Iren Debasu. More Asian discoveries took place even later in the 20th century, including the disembodied arms of Deinocheirus mirificus and the new genus Gallimimus bullatus. The formal naming of the Ornithomimosauria itself was performed by Rinchen Barsbold in 1976.

<i>Afromimus</i> Extinct genus of dinosaurs

Afromimus is a genus of theropod dinosaur from the Early Cretaceous Elrhaz Formation of Niger. It contains a single species, A. tenerensis, named in 2017 by Paul Sereno from parts of the right leg, vertebrae, and ribs found in the Ténéré Desert. It was originally classified as an ornithomimosaurian, but subsequently it was argued to be an abelisauroid.

<i>Paraxenisaurus</i> Genus of deinocheirid dinosaur (fossil)

Paraxenisaurus is an extinct genus of ornithomimosaurian theropod from the Late Cretaceous Cerro del Pueblo Formation of Coahuila in Mexico. The genus contains a single species, P. normalensis, which is known from a few bones of tail, hips, hands, and feet. The specific epithet was given in honor of the Benemérita Normal School of Coahuila, a teacher training institution, where the fossils were reposited. It is a member of the family Deinocheiridae and is the only member of that clade known from Laramidia.

<i>Oksoko avarsan</i> Extinct species of dinosaur

Oksoko is a genus of oviraptorid dinosaur from the Late Cretaceous of Asia, that lived in what is now the Nemegt Formation in Mongolia. It includes the type species Oksoko avarsan.

References

  1. 1 2 3 4 5 6 7 Makovicky, Peter J.; Li, Daqing; Gao, Ke-Qin; Lewin, Matthew; Erickson, Gregory M.; Norell, Mark A. (2010). "A giant ornithomimosaur from the Early Cretaceous of China". Proceedings of the Royal Society B: Biological Sciences. 277 (1679): 191–198. doi:10.1098/rspb.2009.0236. PMC   2842665 . PMID   19386658.
  2. Suarez, Marina B.; Milder, Timothy; Peng, Nan; Suarez, Celina A.; You, Hailu; Li, Daqing; Dodson, Peter (2018-12-13). "Chemostratigraphy of the Lower Cretaceous dinosaur-bearing Xiagou and Zhonggou formations, Yujingzi Basin, northwest China". Journal of Vertebrate Paleontology. 38 (sup1): 12–21. doi:10.1080/02724634.2018.1510412. ISSN   0272-4634. S2CID   202865132.
  3. Paul, G.S., 2010, The Princeton Field Guide to Dinosaurs, Princeton University Press p. 112
  4. Claudia Inés Serrano-Brañas; Belinda Espinosa-Chávez; S. Augusta Maccracken; Cirene Gutiérrez-Blando; Claudio de León-Dávila; José Flores Ventura (2020). "Paraxenisaurus normalensis, a large deinocheirid ornithomimosaur from the Cerro del Pueblo Formation (Upper Cretaceous), Coahuila, Mexico". Journal of South American Earth Sciences. 101: Article 102610. doi:10.1016/j.jsames.2020.102610.
  5. Chinzorig, Tsogtbaatar; Cullen, Thomas; Phillips, George; Rolke, Richard; Zanno, Lindsay E. (2022-10-19). "Large-bodied ornithomimosaurs inhabited Appalachia during the Late Cretaceous of North America". PLOS One. 17 (10). e0266648. doi: 10.1371/journal.pone.0266648 . PMC   9581415 .
  6. Yuong-Nam Lee, Rinchen Barsbold, Philip J. Currie, Yoshitsugu Kobayashi, Hang-Jae Lee, Pascal Godefroit, François Escuillié & Tsogtbaatar Chinzorig (2014) "Resolving the long-standing enigmas of a giant ornithomimosaur Deinocheirus mirificus". Nature (advance online publication) doi : 10.1038/nature13874.