Moros Temporal range: Cenomanian, | |
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Diagram showing known remains | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | † Pantyrannosauria |
Genus: | † Moros Zanno et al., 2019 [1] |
Species: | †M. intrepidus |
Binomial name | |
†Moros intrepidus Zanno et al., 2019 [1] | |
Moros is a genus of small tyrannosauroid theropod dinosaur that lived during the Late Cretaceous period in what is now Utah. It contains a single species, M. intrepidus. [1] Moros represents one of the earliest known diagnostic tyrannosauroid material from North America. [1]
Moros was first discovered at the Stormy Theropod site located in Emery County in the U.S. state of Utah. Palaeontologists had been researching the area for ten years when, in 2013, limb bones were seen jutting out of a hillside, prompting the excavation. [2] The bones were described as of a new species in February, 2019. [3] The type species, Moros intrepidus, was named and described by Lindsay E. Zanno, Ryan T. Tucker, Aurore Canoville, Haviv M. Avrahami, Terry A. Gates, and Peter J. Makovicky. The generic name is derived from the Greek term Moros (an embodiment of impending doom), in reference to the establishment of the tyrannosauroid lineage in North America that would soon dominate the continent by the end of the Cretaceous. The specific name is the Latin word intrepidus ("intrepid"), referring to the hypothesized dispersal of tyrannosauroids from Asia throughout North America following the arrival of Moros. [1]
The holotype specimen, NCSM 33392, was found in the lower Mussentuchit Member of the Cedar Mountain Formation dating from the Cenomanian age. The layer has a maximimum age of 96.4 million years. The holotype consists of a right leg, specifically the thighbone, shinbone, second and fourth metatarsal, and the third and fourth phalanx of the fourth toe. Lines of arrested growth, or LAGs, indicate that it represents a subadult individual of six or seven years old, nearing its maximum size. Additionally, two premaxillary teeth were referred to the species, specimens NCSM 33393 and NCSM 33276. [1]
Moros was a small-bodied, cursorial tyrannosauroid with an estimated leg length of 1.2 m (3.9 ft) and a body mass of 78 kg (172 lb). [1] The foot bones of Moros were extremely slender, with metatarsal proportions found to be more similar to ornithomimids than to other Late Cretaceous tyrannosauroids. [1]
In their 2019 phylogenetic analyses, Zanno and colleagues recovered Moros as a basal pantyrannosaurian alongside Asian taxa from the middle of the Cretaceous, such as Xiongguanlong and Timurlengia . [1] This phylogenetic affinity with Asian basal tyrannosauroids suggests that Moros was part of a transcontinental exchange between the biotas of Asia and North America during the mid-Cretaceous that is well-documented in other taxa. [1]
Moros intepidus was featured in the 2022 American science fiction action film Jurassic World Dominion. It was the first feathered dinosaur confirmed for the film, but was incorrectly depicted as being duck-sized, as opposed to being a deer-sized animal. [4]
Tyrannosaurus is a genus of large theropod dinosaur. The type species Tyrannosaurus rex, often shortened to T. rex or colloquially T-Rex, is one of the best represented theropods. It lived throughout what is now western North America, on what was then an island continent known as Laramidia. Tyrannosaurus had a much wider range than other tyrannosaurids. Fossils are found in a variety of rock formations dating to the latest Campanian-Maastrichtian ages of the late Cretaceous period, 72.7 to 66 million years ago. It was the last known member of the tyrannosaurids and among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event.
Alectrosaurus is a genus of tyrannosauroid theropod dinosaur that lived in Asia during the Late Cretaceous period, about some 96 million years ago in what is now the Iren Dabasu Formation.
Alioramus is a genus of tyrannosaurid theropod dinosaurs from the Late Cretaceous period of Asia. It currently contains two species. The type species, A. remotus is known from a partial skull and three foot bones recovered from the Mongolian Nemegt Formation, which was deposited in a humid floodplain about 70 million years ago. These remains were named and described by Soviet paleontologist Sergei Kurzanov in 1976. A second species, A. altai, known from a much more complete skeleton also from the Nemegt Formation, was named and described by Stephen L. Brusatte and colleagues in 2009. Its relationships to other tyrannosaurid genera were at first unclear, with some evidence supporting a hypothesis that Alioramus was closely related to the contemporary species Tarbosaurus bataar. However, the discovery of Qianzhousaurus indicates that it belongs to a distinct branch of tyrannosaurs, namely the tribe Alioramini.
Tyrannosauroidea is a superfamily of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic Tyrannosaurus. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia.
Dryptosaurus is a genus of basal eutyrannosaurian theropod dinosaur that lived on the island continent of Appalachia approximately 67.6 million years ago during the end of the Maastrichtian age of the Late Cretaceous period. Dryptosaurus was a large, bipedal, ground-dwelling carnivore that could grow up to 7.5 metres (25 ft) long and weigh up to 756–1,500 kilograms (1,667–3,307 lb). Although it is now largely unknown outside of academic circles, the famous 1897 painting of the genus by Charles R. Knight made Dryptosaurus one of the more widely known dinosaurs of its time, in spite of its poor fossil record. First described by Edward Drinker Cope in 1866 and later renamed by Othniel Charles Marsh in 1877, Dryptosaurus is among the first theropod dinosaurs ever known to science.
Tyrannosaurinae is one of the two extinct subfamilies of Tyrannosauridae, a family of coelurosaurian theropods that consists of at least three tribes and several genera. All fossils of these genera have been found in the Late Cretaceous deposits of western North America and east Asia. Compared to the related subfamily Albertosaurinae, tyrannosaurines overall are more robust and larger though the alioramins were gracile by comparison. This subfamily also includes the oldest known tyrannosaurid genus Lythronax as well as the youngest and most famous member of the group, Tyrannosaurus rex. There were at least 30 different species of tyrannosaurines.
Peloroplites is a monospecific genus of nodosaurid dinosaur from Utah that lived during the Late Cretaceous in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Peloroplites cedrimontanus, is known from a partial skull and postcranial skeleton. It was named in 2008 by Kenneth Carpenter and colleagues. Peloroplites was 6 metres long and weighed 2 tonnes, making it one of the largest known nodosaurids, and came from a time when ankylosaurids and nodosaurids were attaining large sizes.
Xiongguanlong is an extinct genus of tyrannosauroid theropod from the Early Cretaceous period of what is now China. The type and only species is X. baimoensis. The generic name comes from Jiayuguan City and the Mandarin word "long" which means dragon. The specific epithet, "baimoensis" is a latinization of the Mandarin word for "white ghost" in reference to one of the geological features of the type locality.
Sinotyrannus was an early, large-bodied genus within the superfamily Tyrannosauroidea. This dinosaur had a single type species, S. kazuoensis, with the only known specimen containing a partial skull, some vertebrae, and a hip, all of which were found in the Early Cretaceous aged Jiufotang Formation of Liaoning, China. While it exhibited greater body size that would put it on par with the later tyrannosaurids such as Tyrannosaurus, Sinotyrannus was probably a member of the basal tyrannosaur family known as the Proceratosauridae. This family originated in the Jurassic, whose members are known from Europe and Asia. Sinotyrannus, alongside another early tyrannosaur, Yutyrannus, appears to have been oddly large when compared to most tyrannosaurs of the Early Cretaceous, such as Dilong. Most of the world during the Early Cretaceous was dominated by more basal tetanurans, such as the megalosaurs and allosaurs, with tyrannosaurids themselves only taking over after both groups started to decline. However, Sinotyrannus and Yutyrannus appear to be exceptions to this. Sinotyrannus was the largest theropod in the Jiufotang Formation, reaching up to 10 meters in overall length and having a general mass similar to that of a large rhino.
Qiupalong is an extinct genus of ornithomimosaurian theropod that was discovered in the Late Cretaceous Qiupa Formation of Henan, China. The genus contains a single species, Q. henanensis, the specific epithet for which was named for the province of Henan. Uniquely, Qiupalong is one of the few Late Cretaceous non-avian dinosaurs known from both Asia and Laramidia. Specimens from Russia and Alberta have been referred to the genus without being assigned to the type species.
Juratyrant is a tyrannosauroid dinosaur genus from the late Jurassic period of England. The genus contains a single species, Juratyrant langhami, which was once classed as a species of Stokesosaurus.
Lythronax is a genus of tyrannosaurid dinosaur that lived in North America around 81.9-81.5 million years ago during the Late Cretaceous period. The only known specimen was discovered in Utah in the Wahweap Formation of the Grand Staircase–Escalante National Monument in 2009, and it consists of a partial skull and skeleton. In 2013, it became the basis of the new genus and species Lythronax argestes; the generic name Lythronax means "gore king", and the specific name argestes originates from the Greek poet Homer's name for the wind from the southwest, in reference to the specimen's geographic provenance in North America.
Siats (/see-ats/) is an extinct genus of large theropod dinosaur known from the Late Cretaceous Cedar Mountain Formation of Utah, United States. It contains a single species, Siats meekerorum. It was initially classified as a megaraptoran, a clade of large theropods with very controversial relationships. Siats may be a neovenatorid allosauroid, a coelurosaur of uncertain phylogenetic position, or a tyrannosauroid.
Suskityrannus is a genus of small tyrannosauroid theropod from the Late Cretaceous in southern Laramidia. It contains a single species, Suskityrannus hazelae, and the type specimen was found in the Turonian-age Moreno Hill Formation of the Zuni Basin in western New Mexico.
Lindsay E. Zanno is an American vertebrate paleontologist and a leading expert on theropod dinosaurs and Cretaceous paleoecosystems. She is the Head of Paleontology at the North Carolina Museum of Natural Sciences and an Associate Research Professor in the Department of Biological Sciences at North Carolina State University.
This timeline of tyrannosaur research is a chronological listing of events in the history of paleontology focused on the tyrannosaurs, a group of predatory theropod dinosaurs that began as small, long-armed bird-like creatures with elaborate cranial ornamentation but achieved apex predator status during the Late Cretaceous as their arms shrank and body size expanded. Although formally trained scientists did not begin to study tyrannosaur fossils until the mid-19th century, these remains may have been discovered by Native Americans and interpreted through a mythological lens. The Montana Crow tradition about thunder birds with two claws on their feet may have been inspired by isolated tyrannosaurid forelimbs found locally. Other legends possibly inspired by tyrannosaur remains include Cheyenne stories about a mythical creature called the Ahke, and Delaware stories about smoking the bones of ancient monsters to have wishes granted.
Timurlengia is an extinct genus of tyrannosauroid theropod dinosaur found in Uzbekistan, in the Bissekty Formation in the Kyzylkum Desert, hailing from the Turonian age of the early Late Cretaceous. The type species is Timurlengia euotica.
Phuwiangvenator is an extinct genus of megaraptoran theropod that lived during the Early Cretaceous period in what is now Thailand. It contains only the type species, P. yaemniyomi. The generic name of Phuwiangvenator comes from the Phu Wiang mountains, where the holotype was discovered, and the Latin word "venator" meaning hunter. The specific name, "yaemniyomi", is in honor of Sudham Yaemniyom, who was a historical paleontologist from Thailand and the first person to discover fossils there.
Iani is an extinct genus of rhabdodontomorph iguanodontian dinosaur from the Late Cretaceous Cedar Mountain Formation of Utah, United States. The genus contains a single species, I. smithi, known from a partial skeleton including the skull. Its discovery serves as a link between the genus Tenontosaurus and the Rhabdodontidae, with both along with Iani being members of the clade Rhabdodontomorpha.
Fona is an extinct genus of thescelosaurine thescelosaurid ornithischian dinosaurs from the 'mid'-Cretaceous Cedar Mountain Formation of Utah, United States. The genus contains a single species, F. herzogae, known from several partial skeletons and skulls. Based on anatomical similarities to the closely related and similarly aged Oryctodromeus, Fona was likely a semi-fossorial animal, likely spending a significant amount of time in underground burrows. It also represents the oldest known definitive thescelosaurine.