Veterupristisaurus | |
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Skeletal diagram showing known remains | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | † Carcharodontosauria |
Family: | † Carcharodontosauridae |
Genus: | † Veterupristisaurus Rauhut, 2011 |
Species: | †V. milneri |
Binomial name | |
†Veterupristisaurus milneri Rauhut, 2011 | |
Synonyms | |
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Veterupristisaurus is a potentially dubious extinct genus of carcharodontosaurid theropod dinosaur known from the Jurassic of Tendaguru, Lindi Region of southeastern Tanzania. [1]
Veterupristisaurus is known from the holotype specimen MB R 1938, an isolated middle caudal vertebra. Two partially fused posterior middle caudal vertebrae, MB R 2166, from the same locality as the holotype, are referred to this genus and most probably came from the same individual. The anterior caudal vertebra, MB R 1940, may also represent this genus. The holotype was collected in the St (EH) locality of the Tendaguru in German East Africa, from the Middle Dinosaur Member of the Tendaguru Formation, dating to the late Kimmeridgian to earliest Tithonian faunal stage of the Late Jurassic, about 154-150 million years ago. The holotype was originally referred to Ceratosaurus? roechlingi by Werner Janensch in 1925. [2]
Veterupristisaurus was named by Oliver W. M. Rauhut in 2011 and the type species is Veterupristisaurus milneri. The generic name translates as "old shark lizard". It refers to the fact that Veterupristisaurus is currently the oldest known representative of the "shark-toothed lizards", the carcharodontosaurids. The specific name honours the paleontologist Angela C. Milner. [1]
Veterupristisaurus was a large bipedal animal. The length of the holotype vertebra is about 123 mm (4.8 in). Veterupristisaurus has been estimated to have been about 8 m (26 ft) in length and to have weighed 1.65 t (1.82 short tons; 1.65 t), [3] based on the more complete and closely related Acrocanthosaurus . There are teeth from Tendaguru Formation that probably come from it. [4] Whether the individual represented by the holotype represents an adult individual cannot be determined based on the available material. It is diagnosed by a spinoprezygapophyseal lamina in the middle caudal vertebrae extending anteriorly to the midwidth of the base of the prezygapophysis and being flanked laterally by a short, parallel lamina extending from the lateral margin of the prezygapophysis posteriorly. Thus, Rauhut considered a sister-group relationship between Veterupristisaurus and Acrocanthosaurus within the Carcharodontosauridae. [1] In 2025, the study that described Tameryraptor, which had Rauhut as a co-author, noted its only diagnostic character was shared with Lusovenator , making the validity of Veterupristisaurus questionable. [5]
Veterupristisaurus is generally recovered as a member of the Carcharodontosauridae [1] [6] although some studies find it to be a Carcharodontosaurian outside of Carcharodontosauridae instead. [7]
Cau (2024) found Veterupristisaurus to be a carcharodontosaurid forming a clade with Sauroniops , Lusovenator, Eocarcharia , and Concavenator .
Carcharodontosauridae |
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Acrocanthosaurus is a genus of carcharodontosaurid dinosaur that existed in what is now North America during the Aptian and early Albian stages of the Early Cretaceous, from 113 to 110 million years ago. Like most dinosaur genera, Acrocanthosaurus contains only a single species, A. atokensis. It had a continent-wide range, with fossil remains known from the U.S. states of Oklahoma, Texas, and Wyoming in the west, and Maryland in the east.
Carcharodontosaurus is a genus of large carnivorous theropod dinosaur that lived in Northwest Africa from about 100 to 94 million years ago during the Cenomanian age of the Cretaceous. Two teeth of the genus, now lost, were first described from Algeria by French paleontologists Charles Depéret and Justin Savornin as Megalosaurus saharicus. A partial skeleton initially referred to this genus was collected by crews of German paleontologist Ernst Stromer during a 1914 expedition to Egypt. Stromer did not report the Egyptian find until 1931, in which he dubbed the novel genus Carcharodontosaurus, making the type species C. saharicus. Although this skeleton was destroyed during the Second World War, it was subsequently redescribed as the holotype of a distinct carcharodontosaurid genus, Tameryraptor. In 1995, a nearly complete skull of C. saharicus, the first well-preserved specimen to be found in almost a century, was discovered in the Kem Kem Beds of Morocco, which was officially designated as the neotype in 2007. In the same year, fossils unearthed from the Echkar Formation of northern Niger were described and named as another species, C. iguidensis, though this species might belong to a different genus.
Torvosaurus is a genus of large megalosaurine theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle and Late Jurassic period in what is now Colorado, Portugal, Germany, and possibly England, Spain, Tanzania, and Uruguay. It contains two currently recognized species, Torvosaurus tanneri and Torvosaurus gurneyi, plus a third unnamed species from Germany.
Carcharodontosauridae is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae as a family, which, in modern paleontology, indicates a clade within Carnosauria. Carcharodontosaurids include some of the largest land predators ever known: Giganotosaurus, Mapusaurus, Carcharodontosaurus, and Tyrannotitan all rivaled Tyrannosaurus in size. Estimates give a maximum weight of 8–10 metric tons for the largest carcharodontosaurids, while the smallest carcharodontosaurids were estimated to have weighed at least 500 kilograms (1,100 lb).
Elaphrosaurus is a genus of ceratosaurian theropod dinosaur that lived approximately 154 to 150 million years ago during the Late Jurassic Period in what is now Tanzania in Africa. Elaphrosaurus was a medium-sized but lightly built member of the group that could grow up to 6.2 m (20 ft) long. Morphologically, this dinosaur is significant in two ways. Firstly, it has a relatively long body but is very shallow-chested for a theropod of its size. Secondly, it has very short hindlimbs in comparison with its body. Phylogenetic analyses indicate that this genus is likely a ceratosaur. Earlier suggestions that it is a late surviving coelophysoid have been examined but generally dismissed. Elaphrosaurus is currently believed to be a very close relative of Limusaurus, an unusual beaked ceratosaurian which may have been either herbivorous or omnivorous.
Tendaguria is a genus of herbivorous sauropod dinosaur from the Late Jurassic of Lindi Region, Tanzania.
Metriacanthosauridae is an extinct family of allosauroid theropod dinosaurs that lived from the Middle Jurassic to the Early Cretaceous. The family is split into two subgroups: Metriacanthosaurinae, which includes dinosaurs closely related to Metriacanthosaurus, and another group composed of the close relatives of Yangchuanosaurus. Metriacanthosaurids are considered carnosaurs, belonging to the Allosauroidea superfamily. The group includes species of large range in body size. Of their physical traits, most notable are their neural spines. The records of the group are mostly confined to Asia, though Metriacanthosaurus is known from Europe. Metriacanthosauridae is used as a senior synonym of Sinraptoridae.
Kelmayisaurus is an extinct genus of allosauroid theropod dinosaur from the Early Cretaceous. It was roughly 10–12 meters long and its name refers to the petroleum-producing city of Karamay in the Xinjiang province of western China near where it was found.
Spinostropheus is a genus of carnivorous neotheropod theropod dinosaur that lived in the Middle Jurassic period and has been found in the Tiouraren Formation, Niger. The type and only species is S. gautieri.
Poekilopleuron is a genus of carnosaurian theropod dinosaur, which lived during the middle Bathonian of the Jurassic, about 168 to 166 million years ago. The genus has been used under many different spelling variants, although only one, Poekilopleuron, is valid. The type species is P. bucklandii, named after William Buckland, and many junior synonyms of it have also been erected. Little material is currently known, as the holotype was destroyed in World War II, although many casts of the material still exist.
Concavenator is a genus of carcharodontosaurian dinosaur that lived in Spain during the Early Cretaceous epoch, about 125 million years ago. The genus contains a single species, Concavenator corcovatus named and described in 2010 from a nearly complete skeleton collected from Las Hoyas fossil site of La Huérguina Formation.
Ostafrikasaurus is a genus of theropod dinosaur from the Late Jurassic period of what is now Lindi Region, Tanzania. It is known only from fossil teeth discovered sometime between 1909 and 1912, during an expedition to the Tendaguru Formation by the Natural History Museum of Berlin. Eight teeth were originally attributed to the dubious dinosaur genus Labrosaurus, and later to Ceratosaurus, both known from the North American Morrison Formation. Subsequent studies attributed two of these teeth to a spinosaurid dinosaur, and in 2012, Ostafrikasaurus crassiserratus was named by French palaeontologist Eric Buffetaut, with one tooth as the holotype, and the other referred to the same species. The generic name comes from the German word for German East Africa, the former name of the colony in which the fossils were found, while the specific name comes from the Latin words for "thick" and "serrated", in reference to the form of the animal's teeth.
Sauroniops is a controversial, potentially dubious, genus of carnivorous basal carcharodontosaurid theropod dinosaur known from the Late Cretaceous of Morocco. It is mainly known from the Gara Sbaa Formation of the Kem Kem beds and contains a single species, S. platytholus.
Siats (/see-ats/) is an extinct genus of large theropod dinosaurs known from the Late Cretaceous Cedar Mountain Formation of Utah, United States. The genus contains a single species, Siats meekerorum. It was initially classified as a megaraptoran, a clade of large theropods with controversial relationships. Alternative positions within the Neovenatoridae, Allosauroidea, and Tyrannosauroidea have also been proposed.
Taurovenator is a large carcharodontosaurid theropod from the late Cretaceous Huincul Formation of Argentina that lived during the Cenomanian age of the Late Cretaceous. It is monotypic, containing only one species, T. violantei.
Wamweracaudia is a large herbivorous sauropod dinosaur from the Late Jurassic Tendaguru Formation of Tanzania, Africa, 155-145 million years ago.
Oceanotitan is a genus of sauropod dinosaur known from the Upper Jurassic Lourinha Formation of Portugal. It is represented by a single specimen consisting of several tail vertebrae and appendicular bones. It contains one species, Oceanotitan dantasi. Oceanotitan is classified as possibly one of the earliest members of the Somphospondyli, a group of sauropods that includes the titanosaurs.
Siamraptor is an extinct genus of carcharodontosaurian dinosaur, containing the single species S. suwati, known from the Khok Kruat Formation of Thailand. It is possibly the first definitive named carcharodontosaurian species known from Southeast Asia. However a later 2024 study found it to be an early tetanuran outside of the group Orionides, although the describers of Alpkarakush still found it within the Carcharodontosauria the same year.
Asfaltovenator is a genus of possibly allosauroid dinosaur from the Lower Jurassic Cañadón Asfalto Formation of Chubut Province, Argentina. The type and only species is Asfaltovenator vialidadi.
Lusovenator is a genus of carcharodontosaurian theropod dinosaur, from the Late Jurassic (Kimmeridgian) Praia de Amoreira Porto-Novo Member and the Late Jurassic-Early Cretaceous (Tithonian-Berriasian) Assenta Member of the Lourinhã Formation in present-day Portugal. It includes one species, Lusovenator santosi.
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