The Toarcian extinction event, also called the Pliensbachian-Toarcian extinction event, [1] [2] the Early Toarcian mass extinction, [3] the Early Toarcian palaeoenvironmental crisis, [4] or the Jenkyns Event, [5] [6] [7] was an extinction event that occurred during the early part of the Toarcian age, approximately 183 million years ago, during the Early Jurassic. The extinction event had two main pulses, [4] the first being the Pliensbachian-Toarcian boundary event (PTo-E). [8] The second, larger pulse, the Toarcian Oceanic Anoxic Event (TOAE), was a global oceanic anoxic event, [9] representing possibly the most extreme case of widespread ocean deoxygenation in the entire Phanerozoic eon. [10] In addition to the PTo-E and TOAE, there were multiple other, smaller extinction pulses within this span of time. [8]
Occurring during the supergreenhouse climate of the Early Toarcian Thermal Maximum (ETTM), [11] the Early Toarcian extinction was associated with large igneous province volcanism, [12] which elevated global temperatures, [11] acidified the oceans, [13] and prompted the development of anoxia, [14] leading to severe biodiversity loss. [15] The biogeochemical crisis is documented by a high amplitude negative carbon isotope excursions, [16] [17] as well as black shale deposition. [18]
The Early Toarcian extinction event occurred in two distinct pulses, [4] with the first event being classified by some authors as its own event unrelated to the more extreme second event. [19] The first, more recently identified pulse occurred during the mirabile subzone of the tenuicostatum ammonite zone, coinciding with a slight drop in oxygen concentrations and the beginning of warming following a late Pliensbachian cool period. [20] This first pulse, occurring near the Pliensbachian-Toarcian boundary, [21] is referred to as the PTo-E. [8] [9] The TOAE itself occurred near the tenuicostatum–serpentinum ammonite biozonal boundary, [22] specifically in the elegantulum subzone of the serpentinum ammonite zone, during a marked, pronounced warming interval. [20] The TOAE lasted for approximately 500,000 years, [23] [24] [25] though a range of estimates from 200,000 to 1,000,000 years have also been given. [26] The PTo-E primarily affected shallow water biota, while the TOAE was the more severe event for organisms living in deep water. [27]
Geological, isotopic, and palaeobotanical evidence suggests the late Pliensbachian was an icehouse period. [28] [29] [30] These ice sheets are believed to have been thin and stretched into lower latitudes, making them extremely sensitive to temperature changes. [31] A warming trend lasting from the latest Pliensbachian to the earliest Toarcian was interrupted by a "cold snap" in the middle polymorphum zone, equivalent to the tenuicostatum ammonite zone, which was then followed by the abrupt warming interval associated with the TOAE. [32] This global warming, driven by rising atmospheric carbon dioxide, was the mainspring of the early Toarcian environmental crisis. [3] Carbon dioxide levels rose from about 500 ppm to about 1,000 ppm. [33] Seawater warmed by anywhere between 3 °C and 7 °C, depending on latitude. [34] At the height of this supergreenhouse interval, global sea surface temperatures (SSTs) averaged about 21 °C. [3]
The eruption of the Karoo-Ferrar Large Igneous Province is generally attributed to have caused the surge in atmospheric carbon dioxide levels. [12] [6] [35] Argon-argon dating of Karoo-Ferrar rhyolites points to a link between Karoo-Ferrar volcanism and the extinction event, [36] a conclusion reinforced by uranium-lead dating [37] [38] [39] and palaeomagnetism. [40] Occurring during a broader, gradual positive carbon isotope excursion as measured by δ13C values, the TOAE is preceded by a global negative δ13C excursion recognised in fossil wood, organic carbon, and carbonate carbon in the tenuicostatum ammonite zone of northwestern Europe, [41] with this negative δ13C shift being the result of volcanic discharge of light carbon. [16] The global ubiquity of this negative δ13C excursion has been called into question, however, due to its absence in certain deposits from the time, such as the Bächental bituminous marls, [42] though its occurrence in areas like Greece has been cited as evidence of its global nature. [43] The negative δ13C shift is also known from the Arabian Peninsula, [44] the Ordos Basin, [45] and the Neuquén Basin. [46] The negative δ13C excursion has been found to be up to -8% in bulk organic and carbonate carbon, although analysis of compound specific biomarkers suggests a global value of around -3% to -4%. In addition, numerous smaller scale carbon isotope excursions are globally recorded on the falling limb of the larger negative δ13C excursion. [16] Although the PTo-E is not associated with a decrease in δ13C analogous to the TOAE's, volcanism is nonetheless believed to have been responsible for its onset as well, with the carbon injection most likely having an isotopically heavy, mantle-derived origin. [47] The Karoo-Ferrar magmatism released so much carbon dioxide that it disrupted the imprint of the 9 Myr long-term carbon cycle that was otherwise steady and stable during the Jurassic and Early Cretaceous. [48] The values of 187Os/188Os rose from ~0.40 to ~0.53 during the PTo-E and from ~0.42 to ~0.68 during the TOAE, and many scholars conclude this change in osmium isotope ratios evidences the responsibility of this large igneous province for the biotic crises. [49] Mercury anomalies from the approximate time intervals corresponding to the PTo-E and TOAE have likewise been invoked as tell-tale evidence of the ecological calamity's cause being a large igneous province, [50] [51] [52] although some researchers attribute these elevated mercury levels to increased terrigenous flux. [53] There is evidence that the motion of the African Plate suddenly changed in velocity, shifting from mostly northward movement to southward movement. Such shifts in plate motion are associated with similar large igneous provinces emplaced in other time intervals. [54] A 2019 geochronological study found that the emplacement of the Karoo-Ferrar large igneous province and the TOAE were not causally linked, and simply happened to occur rather close in time, contradicting mainstream interpretations of the TOAE. The authors of the study conclude that the timeline of the TOAE does not match up with the course of activity of the Karoo-Ferrar magmatic event. [55]
The large igneous province also intruded into coal seams, releasing even more carbon dioxide and methane than it otherwise would have. [56] [57] [16] Magmatic sills are also known to have intruded into shales rich in organic carbon, causing additional venting of carbon dioxide into the atmosphere. [58] Carbon release via metamorphic heating of coal has been criticised as a major driver of the environmental perturbation, however, on the basis that coal transects themselves do not show the δ13C excursions that would be expected if significant quantities of thermogenic methane were released, suggesting that much of the degassed emissions were either condensed as pyrolytic carbon or trapped as coalbed methane. [59]
In addition, possible associated release of deep sea methane clathrates has been potentially implicated as yet another cause of global warming. [60] [61] [62] Episodic melting of methane clathrates dictated by Milankovitch cycles has been put forward as an explanation fitting the observed shifts in the carbon isotope record. [63] [26] [64] Other studies contradict and reject the methane hydrate hypothesis, however, concluding that the isotopic record is too incomplete to conclusively attribute the isotopic excursion to methane hydrate dissociation, [65] that carbon isotope ratios in belemnites and bulk carbonates are incongruent with the isotopic signature expected from a massive release of methane clathrates, [66] that much of the methane released from ocean sediments was rapidly sequestered, buffering its ability to act as a major positive feedback, [67] and that methane clathrate dissociation occurred too late to have had an appreciable causal impact on the extinction event. [68] Hypothetical release of methane clathrates extremely depleted in heavy carbon isotopes has furthermore been considered unnecessary as an explanation for the carbon cycle disruption. [69]
It has also been hypothesised that the release of cryospheric methane trapped in permafrost amplified the warming and its detrimental effects on marine life. [70] [71] Obliquity-paced carbon isotope excursions have been interpreted as some researchers as reflective of permafrost decline and consequent greenhouse gas release. [72] [73]
The TOAE is believed to be the second largest anoxic event of the last 300 Ma, [74] and possibly the largest of the Phanerozoic. [10] A positive δ13C excursion, likely resulting from the mass burial of organic carbon during the anoxic event, is known from the falciferum ammonite zone, chemostratigraphically identifying the TOAE. [75] Large igneous province resulted in increased silicate weathering and an acceleration of the hydrological cycle, [76] [77] as shown by a increased amount of terrestrially derived organic matter found in sedimentary rocks of marine origin during the TOAE. [78] [79] Concentrations of phosphorus, magnesium, and manganese rose in the oceans. [80] A -0.5% excursion in δ44/40Ca provides further evidence of increased continental weathering. [81] Osmium isotope ratios confirm further still a major increase in weathering. [82] The enhanced continental weathering in turn led to increased eutrophication that helped drive the anoxic event in the oceans. [14] [83] [84] Continual transport of continentally weathered nutrients into the ocean enabled high levels of primary productivity to be maintained over the course of the TOAE. [24] Rising sea levels contributed to ocean deoxygenation; [85] as rising sea levels inundated low-lying lands, organic plant matter was transported outwards into the ocean. [86] An alternate model for the development of anoxia is that epicontinental seaways became salinity stratified with strong haloclines, chemoclines, and thermoclines. This caused mineralised carbon on the seafloor to be recycled back into the photic zone, driving widespread primary productivity and in turn anoxia. [87] The freshening of the Arctic Ocean by way of melting of Northern Hemisphere ice caps was a likely trigger of such stratification and a slowdown of global thermohaline circulation. [88] Stratification also occurred due to the freshening of surface water caused by an enhanced water cycle. [89] [90] Rising seawater temperatures amidst a transition from icehouse to greenhouse conditions further retarded ocean circulation, aiding the establishment of anoxic conditions. [91] Geochemical evidence from what was then the northwestern European epicontinental sea suggests that a shift from cooler, more saline water conditions to warmer, fresher conditions prompted the development of significant density stratification of the water column and induced anoxia. [24] Extensive organic carbon burial induced by anoxia was a negative feedback loop retarding the otherwise pronounced warming and may have caused global cooling in the aftermath of the TOAE. [92] In anoxic and euxinic marine basins in Europe, organic carbon burial rates increased by ~500%. [5] Furthermore, anoxia was not limited to oceans; large lakes also experienced oxygen depletion and black shale deposition. [93] [94]
Euxinia occurred in the northwestern Tethys Ocean during the TOAE, as shown by a positive δ34S excursion in carbonate-associated sulphate occurs synchronously with the positive δ13C excursion in carbonate carbon during the falciferum ammonite zone. This positive δ34S excursion has been attributed to the depletion of isotopically light sulphur in the marine sulphate reservoir that resulted from microbial sulphur reduction in anoxic waters. [95] Similar positive δ34S excursions corresponding to the onset of TOAE are known from pyrites in the Sakahogi and Sakuraguchi-dani localities in Japan, with the Sakahogi site displaying a less extreme but still significant pyritic positive δ34S excursion during the PTo-E. [96] Euxinia is further evidenced by enhanced pyrite burial in Zázrivá, Slovakia, [97] enhanced molybdenum burial totalling about 41 Gt of molybdenum, [98] and δ98/95Mo excursions observed in sites in the Cleveland, West Netherlands, and South German Basins. [99] Valdorbia, a site in the Umbria-Marche Apennines, also exhibited euxinia during the anoxic event. [26] There is less evidence of euxinia outside the northwestern Tethys, and it likely only occurred transiently in basins in Panthalassa and the southwestern Tethys. [100] Due to the clockwise circulation of the oceanic gyre in the western Tethys and the rough, uneven bathymetry in the northward limb of this gyre, oxic bottom waters had relatively few impediments to diffuse into the southwestern Tethys, which spared it from the far greater prevalence of anoxia and euxinia that characterised the northern Tethys. [101] The Panthalassan deep water site of Sakahogi was mainly anoxic-ferruginous across the interval spanning the late Pliensbachian to the TOAE, but transient sulphidic conditions did occur during the PTo-E and TOAE. [102] In northeastern Panthalassa, in what is now British Columbia, euxinia dominated anoxic bottom waters. [103]
The early stages of the TOAE were accompanied by a decrease in the acidity of seawater following a substantial decrease prior to the TOAE. Seawater pH then dropped close to the middle of the event, strongly acidifying the oceans. [13] The sudden decline of carbonate production during the TOAE is widely believed to be the result of this abrupt episode of ocean acidification. [104] [105] [106] Additionally, the enhanced recycling of phosphorus back into seawater as a result of high temperatures and low seawater pH inhibited its mineralisation into apatite, helping contribute to oceanic anoxia. The abundance of phosphorus in marine environments created a positive feedback loop whose consequence was the further exacerbation of eutrophication and anoxia. [107]
The extreme and rapid global warming at the start of the Toarcian promoted intensification of tropical storms across the globe. [108] [109]
The extinction event associated with the TOAE primarily affected marine life as a result the collapse of the carbonate factory. [110] Brachiopods were particularly severely hit, [111] [2] [112] with the TOAE representing one of the most dire crises in their evolutionary history. [113] Brachiopod taxa of large size declined significantly in abundance. [114] Uniquely, the brachiopod genus Soaresirhynchia thrived during the later stages of the TOAE due to its low metabolic rate and slow rate of growth, making it a disaster taxon. [115] The species S. bouchardi is known to have been a pioneer species that colonised areas denuded of brachiopods in the northwestern Tethyan region. [116] Ostracods also suffered a major diversity loss, [117] [118] with almost all ostracod clades’ distributions during the time interval corresponding to the serpentinum zone shifting towards higher latitudes to escape intolerably hot conditions near the Equator. [20] Bivalves likewise experienced a significant turnover. [118] The decline of bivalves exhibiting high endemism with narrow geographic ranges was particularly severe. [1] At Ya Ha Tinda, a replacement of the pre-TOAE bivalve assemblage by a smaller, post-TOAE assemblage occurred, [119] while in the Cleveland Basin, the inoceramid Pseudomytiloides dubius experienced the Lilliput effect. [120] Ammonoids, having already experienced a major morphological bottleneck thanks to the Gibbosus Event, [121] about a million years before the Toarcian extinction, suffered further losses in the Early Toarcian diversity collapse. [122] Belemnite richness in the northwestern Tethys dropped during the PTo-E but slightly increased across the TOAE. [123] Belemnites underwent a major change in habitat preference from cold, deep waters to warm, shallow waters. [10] Their average rostrum size also increased, though this trend heavily varied depending on the lineage of belemnites. [123] The Toarcian extinction was unbelievably catastrophic for corals; 90.9% of all Tethyan coral species and 49% of all genera were wiped out. [124] Calcareous nannoplankton that lived in the deep photic zone suffered, with the decrease in abundance of the taxon Mitrolithus jansae used as an indicator of shoaling of the oxygen minimum zone in the Tethys and the Hispanic Corridor. [125] Other affected invertebrate groups included echinoderms, [126] radiolarians, [127] dinoflagellates, [126] and foraminifera. [128] [129] [117] Trace fossils, an indicator of bioturbation and ecological diversity, became highly undiverse following the TOAE. [130]
Carbonate platforms collapsed during both the PTo-E and the TOAE. Enhanced continental weathering and nutrient runoff was the dominant driver of carbonate platform decline in the PTo-E, while the biggest culprits during the TOAE were heightened storm activity and a decrease in the pH of seawater. [27]
The recovery from the mass extinction among benthos commenced with the recolonisation of barren locales by opportunistic pioneer taxa. Benthic recovery was slow and sluggish, being regularly set back thanks to recurrent episodes of oxygen depletion, which continued for hundreds of thousands of years after the main extinction interval. [131] Evidence from the Cleveland Basin suggests it took ~7 Myr for the marine benthos to recover, on par with the Permian-Triassic extinction event. [132] [133] Many marine invertebrate taxa found in South America migrated through the Hispanic Corridor into European seas after the extinction event, aided in their dispersal by higher sea levels. [134]
The TOAE had minor effects on marine reptiles, in stark contrast to the major impact it had on many clades of marine invertebrates. In fact, in the Southwest German Basin, ichthyosaur diversity was higher after the extinction interval, although this may be in part a sampling artefact resulting from a sparse Pliensbachian marine vertebrate fossil record. [135]
The TOAE is suggested to have caused the extinction of various clades of dinosaurs, including coelophysids, dilophosaurids, and many basal sauropodomorph clades, as a consequence of the remodelling of terrestrial ecosystems caused by global climate change. [15] Some heterodontosaurids and thyreophorans also perished in the extinction event. [136] In the wake of the extinction event, many derived clades of ornithischians, sauropods, and theropods emerged, with most of these post-extinction clades greatly increasing in size relative to dinosaurs before the TOAE. [15] Eusauropods were propelled to ecological dominance after their survival of the Toarcian cataclysm. [137] Megalosaurids experienced a diversification event in the latter part of the Toarcian that was possibly a post-extinction radiation that filled niches vacated by the mass death of the Early Toarcian extinction. [138] Insects may have experienced blooms as fish moved en masse to surface waters to escape anoxia and then died in droves due to limited resources. [139]
The volcanogenic extinction event initially impacted terrestrial ecosystems more severely than marine ones. A shift towards a low diversity assemblage of cheirolepid conifers, cycads, and Cerebropollenites-producers adapted for high aridity from a higher diversity ecological assemblage of lycophytes, conifers, seed ferns, and wet-adapted ferns is observed in the palaeobotanical and palynological record over the course of the TOAE. [83] The coincidence of the zenith of Classopolis and the decline of seed ferns and spore producing plants with increased mercury loading implicates heavy metal poisoning as a key contributor to the floristic crisis during the Toarcian mass extinction. [140] Poisoning by mercury, along with chromium, copper, cadmium, arsenic, and lead is speculated to be responsible for heightened rates of spore malformation and dwarfism concomitant with enrichments in all these toxic metals. [141]
The TOAE was associated with widespread phosphatisation of marine fossils believed to result from the warming-induced increase in weathering that increased phosphate flux into the ocean. This produced exquisitely preserved lagerstätten across the world, such as Ya Ha Tinda, Strawberry Bank, and the Posidonia Shale. [142]
As is common during anoxic events, black shale deposition was widespread during the deoxygenation events of the Toarcian. [143] [18] [144] Toarcian anoxia was responsible for the deposition of commercially extracted oil shales, [145] particularly in China. [146] [147]
Enhanced hydrological cycling caused clastic sedimentation to accelerate during the TOAE; the increase in clastic sedimentation was synchronous with excursions in 187Os/188Os, 87Sr/86Sr, and δ44/40Ca. [148]
Additionally, the Toarcian was punctuated by intervals of extensive kaolinite enrichment. [149] These kaolinites correspond to negative oxygen isotope excursions and high Mg/Ca ratios and are thus reflective of climatic warming events that characterised much of the Toarcian. [150] Likewise, illitic/smectitic clays were also common during this hyperthermal perturbation. [151]
The Intertropical Convergence Zone (ITCZ) migrated southwards across southern Gondwana, turning much of the region more arid. This aridification was interrupted, however, in the spinatus ammonite biozone and across the Pliensbachian-Toarcian boundary itself. [152]
The large rise in sea levels resulting from the intense global warming led to the formation of the Laurasian Seaway, which enabled the flow of cool water low in salt content to flow into the Tethys Ocean from the Arctic Ocean. The opening of this seaway may have potentially acted as a mitigating factor that ameliorated to a degree the oppressively anoxic conditions that were widespread across much of the Tethys. [153]
The enhanced hydrological cycle during early Toarcian warming caused lakes to grow in size. [45] During the anoxic event, the Sichuan Basin was transformed into a giant lake, [154] [155] which was believed to be approximately thrice as large as modern-day Lake Superior. [156] Lacustrine sediments deposited as a result of this lake's existence are represented by the Da’anzhai Member of the Ziliujing Formation. [157] Roughly ~460 gigatons (Gt) of organic carbon and ~1,200 Gt of inorganic carbon were likely sequestered by this lake over the course of the TOAE. [156]
The TOAE and the Palaeocene-Eocene Thermal Maximum have been proposed as analogues to modern anthropogenic global warming based on the comparable quantity of greenhouse gases released into the atmosphere in all three events. [61] Some researchers argue that evidence for a major increase in Tethyan tropical cyclone intensity during the TOAE suggests that a similar increase in magnitude of tropical storms is bound to occur as a consequence of present climate change. [109]
An extinction event is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity.
The Jurassic is a geologic period and stratigraphic system that spanned from the end of the Triassic Period 201.4 million years ago (Mya) to the beginning of the Cretaceous Period, approximately 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era as well as the eighth period of the Phanerozoic Eon and is named after the Jura Mountains, where limestone strata from the period were first identified.
Approximately 251.9 million years ago, the Permian–Triassicextinction event forms the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras. It is Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It is also the greatest known mass extinction of insects. It is the greatest of the "Big Five" mass extinctions of the Phanerozoic. There is evidence for one to three distinct pulses, or phases, of extinction.
The Triassic–Jurassic (Tr-J) extinction event (TJME), often called the end-Triassic extinction, marks the boundary between the Triassic and Jurassic periods, 201.4 million years ago. It is one of five major extinction events, profoundly affecting life on land and in the oceans. In the seas, about 23–34% of marine genera disappeared. On land, all archosauromorph reptiles other than crocodylomorphs, dinosaurs, and pterosaurs became extinct; some of the groups which died out were previously abundant, such as aetosaurs, phytosaurs, and rauisuchids. Plants, crocodylomorphs, dinosaurs, pterosaurs and mammals were left largely untouched, allowing the dinosaurs, pterosaurs, and crocodylomorphs to become the dominant land animals for the next 135 million years.
The Paleocene–Eocene thermal maximum (PETM), alternatively ”Eocene thermal maximum 1 (ETM1)“ and formerly known as the "Initial Eocene" or “Late Paleocene thermal maximum", was a geologically brief time interval characterized by a 5–8 °C global average temperature rise and massive input of carbon into the ocean and atmosphere. The event began, now formally codified, at the precise time boundary between the Paleocene and Eocene geological epochs. The exact age and duration of the PETM remain uncertain, but it occurred around 55.8 million years ago (Ma) and lasted about 200 thousand years (Ka).
The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician-Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 445 million years ago (Ma). It is often considered to be the second-largest known extinction event just behind the end-Permian mass extinction, in terms of the percentage of genera that became extinct. Extinction was global during this interval, eliminating 49–60% of marine genera and nearly 85% of marine species. Under most tabulations, only the Permian-Triassic mass extinction exceeds the Late Ordovician mass extinction in biodiversity loss. The extinction event abruptly affected all major taxonomic groups and caused the disappearance of one third of all brachiopod and bryozoan families, as well as numerous groups of conodonts, trilobites, echinoderms, corals, bivalves, and graptolites. Despite its taxonomic severity, the Late Ordovician mass extinction did not produce major changes to ecosystem structures compared to other mass extinctions, nor did it lead to any particular morphological innovations. Diversity gradually recovered to pre-extinction levels over the first 5 million years of the Silurian period.
The Late Devonian extinction consisted of several extinction events in the Late Devonian Epoch, which collectively represent one of the five largest mass extinction events in the history of life on Earth. The term primarily refers to a major extinction, the Kellwasser event, also known as the Frasnian-Famennian extinction, which occurred around 372 million years ago, at the boundary between the Frasnian age and the Famennian age, the last age in the Devonian Period. Overall, 19% of all families and 50% of all genera became extinct. A second mass extinction called the Hangenberg event, also known as the end-Devonian extinction, occurred 359 million years ago, bringing an end to the Famennian and Devonian, as the world transitioned into the Carboniferous Period.
An anoxic event describes a period wherein large expanses of Earth's oceans were depleted of dissolved oxygen (O2), creating toxic, euxinic (anoxic and sulfidic) waters. Although anoxic events have not happened for millions of years, the geologic record shows that they happened many times in the past. Anoxic events coincided with several mass extinctions and may have contributed to them. These mass extinctions include some that geobiologists use as time markers in biostratigraphic dating. On the other hand, there are widespread, various black-shale beds from the mid-Cretaceous which indicate anoxic events but are not associated with mass extinctions. Many geologists believe oceanic anoxic events are strongly linked to the slowing of ocean circulation, climatic warming, and elevated levels of greenhouse gases. Researchers have proposed enhanced volcanism (the release of CO2) as the "central external trigger for euxinia."
The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 184.2 Ma and 174.7 ±0.8 Ma. It follows the Pliensbachian and is followed by the Aalenian.
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma, approximately. The Aptian succeeds the Barremian and precedes the Albian, all part of the Lower/Early Cretaceous.
The Ziliujing Formation is a geological formation in China, It is Early Jurassic in age. It is part of the stratigraphy of the Sichuan Basin. The dinosaur Gongxianosaurus and indeterminate theropod material are known from the Dongyuemiao Member of the formation, as well as dinosaur footprints, Zizhongosaurus and indeterminate prosauropods from the Da'anzhai Member. The basal sauropod Sanpasaurus is known from the Maanshan Member. An possible unnamed stegosaur and the pliosauroid plesiosaur Sinopliosaurus are also known from this formation but they were found an indeterminate member. An unnamed teleosaurid known from a complete skull has also been found in the formation, pending a formal description. The deposition environment during the Da'anzhai Member in the lower Toarcian is thought to have been that of a giant freshwater lake encompassing the whole of the Sichuan basin, around 3 times larger than Lake Superior, coeval with the Toarcian Oceanic Anoxic Event around 183 Ma. The Sinemurian-Pliensbachian boundary event has been recorded on the top of the Dongyuemiao Member, while lower parts of this member are of Earliest Sinemurian age.
The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced with a Paleozoic fauna rich in suspension feeder and pelagic animals.
The Posidonia Shale geologically known as the Sachrang Formation, is an Early Jurassic geological formation of southwestern and northeast Germany, northern Switzerland, northwestern Austria, southern Luxembourg and the Netherlands, including exceptionally well-preserved complete skeletons of fossil marine fish and reptiles.
The Cenomanian-Turonian boundary event, also known as the Cenomanian-Turonian extinction, Cenomanian-Turonian Oceanic Anoxic Event, and referred to also as the Bonarelli Event or Level, was an anoxic extinction event in the Cretaceous period. The Cenomanian-Turonian oceanic anoxic event is considered to be the most recent truly global oceanic anoxic event in Earth's geologic history. There was a large carbon cycle disturbance during this time period, signified by a large positive carbon isotope excursion. However, apart from the carbon cycle disturbance, there were also large disturbances in the ocean's nitrogen, oxygen, phosphorus, sulphur, and iron cycles.
Euxinia or euxinic conditions occur when water is both anoxic and sulfidic. This means that there is no oxygen (O2) and a raised level of free hydrogen sulfide (H2S). Euxinic bodies of water are frequently strongly stratified; have an oxic, highly productive, thin surface layer; and have anoxic, sulfidic bottom water. The word "euxinia" is derived from the Greek name for the Black Sea (Εὔξεινος Πόντος (Euxeinos Pontos)) which translates to "hospitable sea". Euxinic deep water is a key component of the Canfield ocean, a model of oceans during part of the Proterozoic eon (a part specifically known as the Boring Billion) proposed by Donald Canfield, an American geologist, in 1998. There is still debate within the scientific community on both the duration and frequency of euxinic conditions in the ancient oceans. Euxinia is relatively rare in modern bodies of water, but does still happen in places like the Black Sea and certain fjords.
The Calcare di Sogno is a geological formation in Italy, dated to roughly between 182-169 million years ago and covering the Lower Toarcian-Late Bajocian stagess of the Jurassic Period in the Mesozoic Era. Thallatosuchian remains are known from the formation, as well fishes and other taxa.
The Sorthat Formation is a geologic formation on the island of Bornholm, Denmark and in the Rønne Graben in the Baltic Sea. It is of Latest Pliensbachian to Late Toarcian age. Plant fossils have been recovered from the formation, along with several traces of invertebrate animals. The Sorthat Formation is overlain by fluvial to lacustrine gravels, along with sands, clay and in some places coal beds that are part of the Aalenian-Bathonian Bagå Formation. Until 2003, the Sorthat Formation was included as the lowermost part of the Bagå Formation, recovering the latest Pliensbachian to lower Aalenian boundary. The Sorthat strata reflect a mostly marginally deltaic to marine unit. Large streams fluctuated to the east, where a large river system was established at the start of the Toarcian. In the northwest, local volcanism that started in the lower Pliensbachian extended along the North Sea, mostly from southern Sweden. At this time, the Central Skåne Volcanic Province and the Egersund Basin expelled most of their material, with influences on the local tectonics. The Egersund Basin has abundant fresh porphyritic nephelinite lavas and dykes of lower Jurassic age, with a composition nearly identical to those found in the clay pits. That indicates the transport of strata from the continental margin by large fluvial channels of the Sorthat and the connected Röddinge Formation that ended in the sea deposits of the Ciechocinek Formation green series.
The Selli Event, also known as OAE1a, was an oceanic anoxic event (OAE) of global scale that occurred during the Aptian stage of the Early Cretaceous, about 120.5 million years ago (Ma). The OAE is associated with large igneous province volcanism and an extinction event of marine organisms driven by global warming, ocean acidification, and anoxia.
The Breistroffer Event (OAE1d) was an oceanic anoxic event (OAE) that occurred during the middle Cretaceous period, specifically in the latest Albian, around 101 million years ago (Ma).
The Paquier Event (OAE1b) was an oceanic anoxic event (OAE) that occurred around 111 million years ago (Ma), in the Albian geologic stage, during a climatic interval of Earth's history known as the Middle Cretaceous Hothouse (MKH).
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