| Holoptychiidae | |
|---|---|
 | |
| Holoptychius sp. | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Sarcopterygii |
| Class: | Porolepimorpha |
| Order: | † Porolepiformes |
| Family: | † Holoptychiidae Owen, 1860 |
| Genera | |
| |
Holoptychiidae is an extinct family of lobe-finned fishes which lived during the Devonian and Carboniferous periods. At least one genus, Laccognathus , is thought to have been amphibious. [1]
The Devonian is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is named after Devon, South West England, where rocks from this period were first studied.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Sarcopterygii — sometimes considered synonymous with Crossopterygii — is a taxon of the bony fish known as the lobe-finned fish or sarcopterygians, characterised by prominent muscular buds (lobes) within the fins, which are supported by appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered bony spines (lepidotrichia) supporting the fins and has no major skeleton within the fins.
Placodermi is a class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scaled or naked, depending on the species. Placoderms were among the first jawed fish; their jaws likely evolved from the first of their gill arches.
The Late Devonian extinction consisted of several extinction events in the Late Devonian Epoch, which collectively represent one of the five largest mass extinction events in the history of life on Earth. The term primarily refers to a major extinction, the Kellwasser event, also known as the Frasnian-Famennian extinction, which occurred around 372 million years ago, at the boundary between the Frasnian stage and the Famennian stage, the last stage in the Devonian Period. Overall, 19% of all families and 50% of all genera became extinct. A second mass extinction called the Hangenberg event, also known as the end-Devonian extinction, occurred 359 million years ago, bringing an end to the Famennian and Devonian, as the world transitioned into the Carboniferous Period.
The Givetian is one of two faunal stages in the Middle Devonian Period. It lasted from 387.7 million years ago to 382.7 million years ago. It was preceded by the Eifelian Stage and followed by the Frasnian Stage. It is named after the town of Givet in France. The oldest forests occurred during the late Givetian. The lower GSSP is located at Jebel Mech Irdane, Tafilalt, Morocco.
Panderichthys is a genus of extinct sarcopterygian from the late Devonian period, about 380 Mya. Panderichthys, which was recovered from Frasnian deposits in Latvia, is represented by two species. P. stolbovi is known only from some snout fragments and an incomplete lower jaw. P. rhombolepis is known from several more complete specimens. Although it probably belongs to a sister group of the earliest tetrapods, Panderichthys exhibits a range of features transitional between tristichopterid lobe-fin fishes and early tetrapods. It is named after the German-Baltic paleontologist Christian Heinrich Pander. Possible tetrapod tracks dating back to before the appearance of Panderichthys in the fossil record were reported in 2010, which suggests that Panderichthys is not a direct ancestor of tetrapods, but nonetheless shows the traits that evolved during the fish-tetrapod evolution
The Frasnian is one of two faunal stages in the Late Devonian Period. It lasted from 382.7 million years ago to 372.2 million years ago. It was preceded by the Givetian Stage and followed by the Famennian Stage.
The Famennian is the later of two faunal stages in the Late Devonian Epoch. The most recent estimate for its duration estimates that it lasted from around 371.1 million years ago to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commission on Stratigraphy, estimated that it lasted from 372.2 million years ago to 358.9 million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage.
The Early Devonian is the first of three epochs comprising the Devonian period, corresponding to the Lower Devonian series. It lasted from 419.2 ± 3.2 to 393.3 ± 1.2 and began with the Lochkovian Stage 419.2 ± 3.2 to 410.8 ± 1.2, which was followed by the Pragian from 410.8 ± 3.2 to 407.6 ± 1.2 and then by the Emsian, which lasted until the Middle Devonian began, 393.3± 1.2 million years ago. During this time, the first ammonoids appeared, descending from bactritoid nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era.
Elpistostegalia or Panderichthyida is an order of prehistoric lobe-finned fishes which lived during the Middle Devonian to Late Devonian period. They represent the advanced tetrapodomorph stock, the fishes more closely related to tetrapods than the osteolepiform fishes. The earliest elpistostegalians, combining fishlike and tetrapod-like characters, are sometimes called fishapods, a phrase coined for the advanced elpistostegalian Tiktaalik. Through a strict cladistic view, the order includes the terrestrial tetrapods.
Laccognathus is an extinct genus of amphibious lobe-finned fish from Europe and North America. They existed from the Middle Devonian to the Late Devonian. The name comes from Greek for 'pitted jaw'.
The geology of the Orkney islands in northern Scotland is dominated by the Devonian Old Red Sandstone (ORS). In the southwestern part of Mainland, this sequence can be seen to rest unconformably on a Moinian type metamorphic basement.
The Rhynchonellata is a class of Lower Cambrian to Recent articulate brachiopods that combines orders from within the Rhynchonelliformea with well developed pedicle attachment. Shell forms vary from those with wide hinge lines to beaked forms with virtually no hinge line and from generally smooth to strongly plicate. Most all are biconvex. Lophophores vary and include both looped and spiraled forms. Although morphologically distinct, included orders follow a consistent phylogenetic sequence.
Laccognathus embryi is an extinct species of porolepiform lobe-finned fish recovered from Ellesmere Island, Canada. It existed during the Frasnian age of the Late Devonian epoch.
Laccognathus panderi is an extinct lobe-finned fish from northern Europe. They existed from the Middle Devonian to the Late Devonian.
Laccognathus grossi is an extinct lobe-finned fish from eastern Europe. They existed during the Middle Devonian. The species is named after Walter Gross, who collected the first Laccognathus fossils belonging to L. panderi.
Ashton F. Embry is a Canadian research scientist at the Geological Survey of Canada.
The Gauja Formation is a Middle Devonian fossil locality in Estonia and Latvia. It is named after the Gauja River, where it is exposed along the banks.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
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