Heishanosaurus

Last updated

Heishanosaurus
Temporal range: Aptian–Albian
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Choristodera
Genus: Heishanosaurus
Dong et al., 2020
Type species
Heishanosaurus pygmaeus
Dong et al., 2020

Heishanosaurus is an extinct genus of choristodere reptile from the Early Cretaceous of China. The type and currently only known species is Heishanosaurus pygmaeus. It is unusual as it is much more primitive than other known choristoderes from the Early Cretaceous of Asia, and retains many plesiomorphic characters.

Contents

Discovery

The type specimens are represented by the blocks IVPP V25322, 25323 and 25324, were discovered in September 2003 at the Badaohao locality, in Heishan County, Liaoning Province, China, in sediments belonging to the Shahai Formation. The Shahai Formation is generally thought to overlie the Jiufotang Formation and has an uncertain Aptian-Albian age, based on the radiometric dating of the Jiufotang Formation. The remains are disarticulated inside the blocks, and were scanned using microcomputed tomography. [1]

Description

Remains include disarticulated skull bones, along with limited postcranial remains. Based on the size of the skull, the animal likely around 16 cm long, similar in size to Cteniogenys . The skull retains an open lower temporal fenestra, as in Cteniogenys, Coeruleodraco and neochoristoderes, but which is closed in the majority of Early Cretaceous Asian "non-neochoristoderans". [1]

Phylogeny

In a strict consensus phylogenetic analysis, it was recovered as a basal choristodere, more derived than Cteniogenys, but more primitive than neochoristoderes or the weakly supported clade containing all other Asian Early Cretaceous "non-neochoristoderans" with closed lower temporal fenestrae, which was informally named the "allochoristoderes" in the study. Heishanosaurus was recovered in a polytomy with the Late Jurassic Coeruleodraco, despite its much younger age. [1]

Choristodera

Cteniogenys sp.

Heishanosaurus pygmaeus

Coeruleodraco jurassicus

Neochoristodera

Ikechosaurus pijiagouensis

Ikechosaurus sunailinae

Tchoiria namsari

Tchoiria klauseni

Champsosaurus

C. gigas

C. albertensis

Simoedosaurus

S. lemoinei

S. dakotensis

"Allochoristodera"

Monjurosuchus splendens

Philydrosaurus proseilus

Lazarussuchus

L. inexpectatus

Lazarussuchus sp.

L. dvoraki

Khurendukhosaurus orlovi

Hyphalosaurus sp.

Hyphalosaurus lingyuanensis

Shokawa ikoi

Related Research Articles

<i>Champsosaurus</i> Extinct genus of reptiles

Champsosaurus is an extinct genus of crocodile-like choristodere reptile, known from the Late Cretaceous and early Paleogene periods of North America and Europe (Campanian–Paleocene). The name Champsosaurus is thought to come from champsai, (χαμψαι) said in an Ancient Greek source to be an Egyptian word for "crocodiles", and sauros, (σαύρος) Greek for "lizard". The morphology of Champsosaurus resembles that of gharials, with a long, elongated snout. It was native to freshwater environments where it likely preyed on fish, similar to living gharials.

<span class="mw-page-title-main">Choristodera</span> Extinct order of reptiles

Choristodera is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the Miocene. Choristoderes are morphologically diverse, with the best known members being the crocodile-like neochoristoderes such as Champsosaurus. Other choristoderans had lizard-like or long necked morphologies. Choristoderes appear to have been confined to the Northern Hemisphere, having been found in North America, Asia, and Europe, and possibly also North Africa. Choristoderes are generally thought to be derived neodiapsids that are close relatives or members of Sauria.

<i>Psittacosaurus</i> Early Cretaceous dinosaur genus

Psittacosaurus is a genus of extinct ceratopsian dinosaur from the Early Cretaceous of what is now Asia, existing between 126 and 101 million years ago. It is notable for being the most species-rich non-avian dinosaur genus. Up to 12 species are known, from across China, Mongolia, Russia, and Thailand. The species of Psittacosaurus were obligate bipeds at adulthood, with a high skull and a robust beak. One individual was found preserved with long filaments on the tail, similar to those of Tianyulong. Psittacosaurus probably had complex behaviours, based on the proportions and relative size of the brain. It may have been active for short periods of time during the day and night, and had well-developed senses of smell and vision.

<i>Hyphalosaurus</i> Genus of extinct freshwater aquatic reptiles

Hyphalosaurus is a genus of freshwater aquatic reptiles, belonging to the extinct order Choristodera. They lived during the early Cretaceous period, about 123-120 million years ago. The genus contains two species, H. lingyuanensis from the Yixian Formation and H. baitaigouensis from both the Yixian and Jiufotang Formation of Liaoning Province, China. They are among the best-known animals from the Jehol Biota, with thousands of fossil specimens representing all growth stages in scientific and private collections.

<i>Liaoningosaurus</i> Extinct genus of dinosaurs

Liaoningosaurus is an unusual genus of basal ankylosaurid dinosaur from the Liaoning Province, China that lived during the Early Cretaceous in what is now the Yixian and Jiufotang Formation. The type and only species, Liaoningosaurus paradoxus, is known from more than 20 specimens, with some representing juveniles. It was named in 2001 by Xu, Wang and You.

<i>Shokawa</i> Extinct genus of reptiles

Shokawa is an extinct genus of choristoderan diapsid reptile, known from the Lower Cretaceous of Japan. It is only known from one species, Shokawa ikoi. The only known remains are a postcranial specimen lacking the skull, discovered at the KO2 locality in sediments belonging to the Okurodani Formation near the village of Shokawa in Gifu Prefecture. Shokawa possessed a long neck with at least 16 cervical vertebrae, and closely resembles and is closely related to the smaller choristoderan, Hyphalosaurus. The generic name refers to the village near where it was found, while the specific name honors the collector of the first specimen, one Mr. Ikoi Shibata.

<i>Lazarussuchus</i> Extinct genus of reptiles

Lazarussuchus is an extinct genus of amphibious reptile, known from the Cenozoic of Europe. It is the youngest known member of Choristodera, an extinct order of aquatic reptiles that first appeared in the Middle Jurassic. Fossils have been found in Late Paleocene, Late Oligocene, Early Miocene and possibly Late Miocene deposits in France, Germany, and the Czech Republic. Two species have been named: the type species L. inexpectatus ("unexpected") from the late Oligocene of France. and L. dvoraki from the early Miocene of the Czech Republic. It was not a large animal; with the total preserved body and tail length of L. inexpectatus being just over 30 centimetres. A complete specimen of Lazarussuchus with preserved soft tissue was found from the Late Paleocene of France, but has not been assigned to a species.

Hongshanopterus is a genus of pterodactyloid pterosaur from the Lower Cretaceous Jiufotang Formation of Liaoning, China.

<i>Gobiconodon</i> Extinct genus of mammals

Gobiconodon is an extinct genus of carnivorous mammals belonging to the family Gobiconodontidae. Undisputed records of Gobiconodon are restricted to the Early Cretaceous of Asia and North America, but isolated teeth attributed to the genus have also been described from formations in England and Morocco dating as far back as the Middle Jurassic. Species of Gobiconodon varied considerably in size, with G. ostromi, one of the larger species, being around the size of a modern Virginia opossum. Like other gobiconodontids, it possessed several speciations towards carnivory, such as shearing molariform teeth, large canine-like incisors and powerful jaw and forelimb musculature, indicating that it probably fed on vertebrate prey. Unusually among predatory mammals and other eutriconodonts, the lower canines were vestigial, with the first lower incisor pair having become massive and canine-like. Like the larger Repenomamus there might be some evidence of scavenging.

<i>Similicaudipteryx</i> Extinct genus of dinosaurs

Similicaudipteryx, meaning "similar to Caudipteryx", is a genus of theropod dinosaur of the family Caudipteridae.

<i>Cteniogenys</i> Genus of reptiles

Cteniogenys is a genus of choristodere, a morphologically diverse group of aquatic reptiles. It is part of the monotypic family Cteniogenidae. The type and only named species, C. antiquus, was named in 1928 by Charles W. Gilmore. The holotype, VP.001088, was collected in the Morrison Formation, Wyoming in 1881 by William H. Reed. More specimens have been discovered since then, including specimens from the Late Jurassic of Portugal and Middle Jurassic of Britain, which have not been assigned to species.

<i>Monjurosuchus</i> Extinct genus of reptiles

Monjurosuchus is a genus of choristoderan reptile that lived in what is now China and Japan during the Early Cretaceous. It has large eyes, a rounded skull, robust legs with short claws, and a long, thin tail. Fossils have been found that preserve soft tissue, showing that it had soft skin and webbed feet.

<i>Tchoiria</i> Extinct genus of diapsid reptiles

Tchoiria () is a genus of neochoristoderan reptile from the Early Cretaceous of Mongolia. The name Tchoiria comes from the city of Choir which is nearby to where the holotype was found. Tchoiria is thought to have a similar diet to another neochoristoderan reptile, Champsosaurus, due to morphology of the skull. It would hunt in freshwater environments, like the living gharials, where it would prey on many different types of fish and turtles.

<span class="mw-page-title-main">Neochoristodera</span> Extinct suborder of reptiles

Neochoristodera is a lineage of specialised crocodile-like fully aquatic choristodere reptiles. Noted for their long jaws and large size, these animals were predominant across the Northern Hemisphere, occurring in freshwater and coastal environments across the Cretaceous and early Cenozoic.

<i>Philydrosaurus</i> Extinct genus of reptiles

Philydrosaurus is an extinct genus of choristoderan which existed in China during the Early Cretaceous. The type species P. proseilus was named in 2005. Philydrosaurus was found from the Jiufotang Formation and is slightly younger than Monjurosuchus, which was found from the Yixian Formation.

Kielanobaatar is an extinct genus of albionbaatarid multituberculate which existed in Shahai and Fuxin formations, northeastern China, during the early Cretaceous. It was first named by Nao Kusuhashi, Yaoming Hu, Yuanqing Wang, Takeshi Setoguchi and Hiroshige Marsuoka in 2010 and the type species is Kielanobaatar badaohaoensis, named after Zofia Kielan-Jaworowska, a leading specialist on Mesozoic mammals. It is known from a small fragment of the left lower jaw, including the third and fourth premolars, as well as two upper premolars. Kielanobaatar is the first record of an Asian albionbaatarid multituberculate. Since the albionbaataridae is known primarily from Europe, this discovery supports the idea of faunal exchange of terrestrial vertebrates between Europe and Asia in the Early Cretaceous.

<i>Chuanqilong</i> Extinct genus of dinosaurs

Chuanqilong is a monospecific genus of basal ankylosaurid dinosaur from the Liaoning Province, China that lived during the Early Cretaceous in what is now the Jiufotang Formation. The type and only species, Chuanqilong chaoyangensis, is known from a nearly complete skeleton with a skull of a juvenile individual. It was described in 2014 by Fenglu Han, Wenjie Zheng, Dongyu Hu, Xing Xu, and Paul M. Barrett. Chuanqilong shows many similarities with Liaoningosaurus and may represent a later ontogenetic stage of the taxon.

<i>Ikrandraco</i> Genus of lonchodraconid pterosaur from the Early Cretaceous

Ikrandraco is a genus of lonchodraconid pterodactyloid pterosaur known from the Lower Cretaceous Jiufotang Formation of northeastern China, and the Cambridge Greensand of England. It is notable for its unusual skull, which features a crest on the lower jaw.

<span class="mw-page-title-main">Sinemydidae</span> Extinct family of turtles

Sinemydidae is an extinct family of turtles from Cretaceous to Paleocene deposits in Asia and North America. Their exact position is engimatic, they have alternatively been considered stem-group cryptodires, but also "crownward stem-turtles" alongside Macrobaenidae, Paracryptodira, Xinjiangchelyidae, Thalassochelydia and Sandownidae outside of crown Testudines.

<i>Coeruleodraco</i> Extinct genus of choristoderan reptiles

Coeruleodraco is an extinct genus of choristoderan known from the Late Jurassic (Oxfordian) Tiaojishan Formation in China. Coeruleodraco is significant as the most complete Jurassic choristodere taxon, as the only other named Jurassic choristodere Cteniogenys is based on fragmentary remains. Although similar to Philydrosaurus in its proportions and postcranial characters, it is distinct in retaining several apparently plesiomorphic characters, including a short snout, paired external nares and an open lower temporal fenestra.

References

  1. 1 2 3 Dong, Liping; Matsumoto, Ryoko; Kusuhashi, Nao; Wang, Yuanqing; Wang, Yuan; Evans, Susan E. (2020-08-02). "A new choristodere (Reptilia: Choristodera) from an Aptian–Albian coal deposit in China". Journal of Systematic Palaeontology. 18 (15): 1223–1242. doi:10.1080/14772019.2020.1749147. ISSN   1477-2019. S2CID   219047160.