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The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Ma) to the start of the Silurian Period 443.8 Mya.
Approximately 251.9 million years ago, the Permian–Triassicextinction event forms the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras. It is the Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It is also the greatest known mass extinction of insects. It is the greatest of the "Big Five" mass extinctions of the Phanerozoic. There is evidence for one to three distinct pulses, or phases, of extinction.
The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician-Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 445 million years ago (Ma). It is often considered to be the second-largest known extinction event, in terms of the percentage of genera that became extinct. Extinction was global during this interval, eliminating 49–60% of marine genera and nearly 85% of marine species. Under most tabulations, only the Permian-Triassic mass extinction exceeds the Late Ordovician mass extinction in biodiversity loss. The extinction event abruptly affected all major taxonomic groups and caused the disappearance of one third of all brachiopod and bryozoan families, as well as numerous groups of conodonts, trilobites, echinoderms, corals, bivalves, and graptolites. Despite its taxonomic severity, the Late Ordovician mass extinction did not produce major changes to ecosystem structures compared to other mass extinctions, nor did it lead to any particular morphological innovations. Diversity gradually recovered to pre-extinction levels over the first 5 million years of the Silurian period.
The Late Devonian extinction consisted of several extinction events in the Late Devonian Epoch, which collectively represent one of the five largest mass extinction events in the history of life on Earth. The term primarily refers to a major extinction, the Kellwasser event, also known as the Frasnian-Famennian extinction, which occurred around 372 million years ago, at the boundary between the Frasnian stage and the Famennian stage, the last stage in the Devonian Period. Overall, 19% of all families and 50% of all genera became extinct. A second mass extinction called the Hangenberg event, also known as the end-Devonian extinction, occurred 359 million years ago, bringing an end to the Famennian and Devonian, as the world transitioned into the Carboniferous Period.
The Tremadocian is the lowest stage of Ordovician. Together with the later Floian Stage it forms the Lower Ordovician Epoch. The Tremadocian lasted from 485.4 to 477.7 million years ago. The base of the Tremadocian is defined as the first appearance of the conodont species Iapetognathus fluctivagus at the Global Boundary Stratotype Section and Point (GSSP) section on Newfoundland.
The Cambrian–Ordovician extinction event, also known as the Cambrian-Ordovician boundary event, was an extinction event that occurred approximately 485 million years ago (mya) in the Paleozoic era of the early Phanerozoic eon. It was preceded by the less-documented End-Botomian mass extinction around 517 million years ago, and the Dresbachian extinction event about 502 million years ago.
The Andean-Saharan glaciation, also known as the Early Paleozoic Ice Age (EPIA), the Early Paleozoic Icehouse, the Late Ordovician glaciation, the end-Ordovician glaciation, or the Hirnantian glaciation, occurred during the Paleozoic from approximately 460 Ma to around 420 Ma, during the Late Ordovician and the Silurian period. The major glaciation during this period was formerly thought only to consist of the Hirnantian glaciation itself but has now been recognized as a longer, more gradual event, which began as early as the Darriwilian, and possibly even the Floian. Evidence of this glaciation can be seen in places such as Arabia, North Africa, South Africa, Brazil, Peru, Bolivia, Chile, Argentina, and Wyoming. More evidence derived from isotopic data is that during the Late Ordovician, tropical ocean temperatures were about 5 °C cooler than present day; this would have been a major factor that aided in the glaciation process.
In the geologic timescale, the Gorstian is an age of the Ludlow Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 427.4 ± 0.5 Ma and 425.6 ± 0.9 Ma, approximately. The Gorstian Age succeeds the Homerian Age and precedes the Ludfordian Age. The age is named after Gorsty village southwest of Ludlow. The base of the age is marked by Graptolites tumescens and Graptolites incipiens. The type section is located in a quarry in the Elton Formation at Pitch Coppice, Shropshire, United Kingdom.
In the geological timescale, the Llandovery Epoch occurred at the beginning of the Silurian Period. The Llandoverian Epoch follows the massive Ordovician-Silurian extinction events, which led to a large decrease in biodiversity and an opening up of ecosystems.
The Daptocephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the upper Teekloof Formation west of 24°E, the majority of the Balfour Formation east of 24°E, and the Normandien Formation in the north. It has numerous localities which are spread out from Colesberg in the Northern Cape, Graaff-Reniet to Mthatha in the Eastern Cape, and from Bloemfontein to Harrismith in the Free State. The Daptocephalus Assemblage Zone is one of eight biozones found in the Beaufort Group and is considered Late Permian (Lopingian) in age. Its contact with the overlying Lystrosaurus Assemblage Zone marks the Permian-Triassic boundary.
The Lau event was the last of three relatively minor mass extinctions during the Silurian period. It had a major effect on the conodont fauna, but barely scathed the graptolites, though they suffered an extinction very shortly thereafter termed the Kozlowskii event that some authors have suggested was coeval with the Lau event and only appears asynchronous due to taphonomic reasons. It coincided with a global low point in sea level caused by glacioeustasy and is closely followed by an excursion in geochemical isotopes in the ensuing late Ludfordian faunal stage and a change in depositional regime.
The Ireviken event was the first of three relatively minor extinction events during the Silurian period. It occurred at the Llandovery/Wenlock boundary. The event is best recorded at Ireviken, Gotland, where over 50% of trilobite species became extinct; 80% of the global conodont species also became extinct in this interval.
The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced with a Paleozoic fauna rich in suspension feeder and pelagic animals.
The Steptoean Positive Carbon Isotope Excursion (SPICE) was a geological event which occurred about 495 million years ago near the beginning of the International Furongian Epoch of the Cambrian Period. The SPICE represents an increase in the atmospheric 13C to 12C ratio, known as the δ13C, and lasted for around 2 to 4 million years.
Athyridida is an order of Paleozoic brachiopods included in the Rhynchonellata, which makes up part of the articulate brachiopods.
The Lilliput effect is a decrease in body size in animal species that have survived a major extinction. There are several hypotheses as to why these patterns appear in the fossil record, some of which are: the survival of small taxa, dwarfing of larger lineages, and the evolutionary miniaturization from larger ancestral stocks. The term was coined in 1993 by Adam Urbanek in his paper concerning the extinction of graptoloids and is derived from the island of Lilliput inhabited by a miniature race of people in Gulliver’s Travels. This size decrease may just be a temporary phenomenon restricted to the survival period of the extinction event. In 2019 Atkinson et al. coined the term the Brobdingnag effect to describe a related phenomenon operating in the opposite direction, whereby new species evolving after the Triassic-Jurassic mass extinction originated at small body sizes before undergoing a size increase. The term is also from Gulliver's Travels where Brobnignag is a land inhabited by a race of giants.
The Capitanian mass extinction event, also known as the end-Guadalupian extinction event, the Guadalupian-Lopingian boundary mass extinction, the pre-Lopingian crisis, or the Middle Permian extinction, was an extinction event that predated the end-Permian extinction event. The mass extinction occurred during a period of decreased species richness and increased extinction rates near the end of the Middle Permian, also known as the Guadalupian epoch. It is often called the end-Guadalupian extinction event because of its initial recognition between the Guadalupian and Lopingian series; however, more refined stratigraphic study suggests that extinction peaks in many taxonomic groups occurred within the Guadalupian, in the latter half of the Capitanian age. The extinction event has been argued to have begun around 262 million years ago with the Late Guadalupian crisis, though its most intense pulse occurred 259 million years ago in what is known as the Guadalupian-Lopingian boundary event.
The Toarcian extinction event, also called the Pliensbachian-Toarcian extinction event, the Early Toarcian mass extinction, the Early Toarcian palaeoenvironmental crisis, or the Jenkyns Event, was an extinction event that occurred during the early part of the Toarcian age, approximately 183 million years ago, during the Early Jurassic. The extinction event had two main pulses, the first being the Pliensbachian-Toarcian boundary event (PTo-E). The second, larger pulse, the Toarcian Oceanic Anoxic Event (TOAE), was a global oceanic anoxic event, representing possibly the most extreme case of widespread ocean deoxygenation in the entire Phanerozoic eon. In addition to the PTo-E and TOAE, there were multiple other, smaller extinction pulses within this span of time.
The Dasberg Event was a minor extinction event that occurred during the Famennian, the final stage of the Devonian period. It is often considered to be one of the events contributing to the Late Devonian extinction, which is believed by many palaeontologists to have been a protracted event that took place over millions of years.
The Šilalė Event was an extinction event affecting conodonts during the Přídolí, the final stage of the Silurian period.
References
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- ↑ Whittingham, Misha; Spiridonov, Andrej; Radzevičius, Sigitas (3 December 2021). "Dynamic ecophenotypy in the Silurian Monograptidae (Graptolithina)". Earth and Environmental Science Transactions of the Royal Society of Edinburgh . 113 (1): 29–38. doi: 10.1017/S1755691021000402 . S2CID 244877517 . Retrieved 31 March 2023.