Pachycaul

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Fouquieria columnaris

Pachycauls are plants with a disproportionately thick trunk, for their height, and relatively few branches. [1] With certain pachycaul species, particularly the more succulent varieties, they are commonly referred to as "caudiciformes", a reference to their trunk development of a moisture-filled caudex for periods of drought.

By comparison, trees with thin twigs, such as oaks ( Quercus ), maples ( Acer ) and Eucalyptus , are called leptocauls; those with moderately thick twigs, such as Plumeria , are called mesocauls. Pachycauls can be the product of exceptional primary growth (as with Arecaceae and Cycadaceae) or disproportionate secondary growth, as with the baobabs ( Adansonia ). The word is derived from the Greek pachy- (meaning "thick" or "stout") and the Latin caulis (meaning "stem"). [2] All of the arboreum (treelike) species of Cactaceae are pachycauls, as are most of the Arecaceae, Cycadaceae and Pandanus . The most extreme pachycauls are the floodplains, or river-bottom, varieties of the African palmyra (Borassus aethiopum), with primary growth up to seven feet (2.1 meters) in thickness, [3] [4] [5] and the Coquito palm (Jubaea chilensis), with its primary growth measuring up to six feet (1.8 meters) thick. [6] [7] [8] The most pachycaulous cycad is Cycas thouarsii at up to five feet (150 centimeters) in diameter. [9] The tallest pachycaul is the Andean wax palm (Ceroxylon quindiuense), at heights of up to 220 feet (67 meters). [10] and around 16 inches (41 cm) in diameter. The most pachycaulous cactus are the barrel cacti (Echinocactus platyacanthus), with primary growth up to 4 ft 4 in (1.32 meters) in diameter. [11] The largest caudiciforme-type pachycaul is the African baobab (Adansonia digitata); another species, called the Glencoe baobab (from Hoedspruit, Limpopo, South Africa) has a basal diameter (not girth) of 52 ft 2 in (15.90 meters). [12] However, this particular tree has suffered a severe trauma and is dying.

Examples occur in the genera [13]

See also

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<i>Adansonia</i> Genus of plants known as baobab

Adansonia is a genus made up of eight species of medium-to-large deciduous trees known as baobabs or adansonias. They are placed in the Malvaceae family, subfamily Bombacoideae. They are native to Madagascar, mainland Africa, and Australia. The trees have also been introduced to other regions such as Asia. A genomic and ecological analysis has suggested that the genus is Madagascan in origin.

<span class="mw-page-title-main">Forb</span> Herbaceous, broad-leaved flowering plant

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<i>Adansonia gregorii</i> Species of tree

Adansonia gregorii, commonly known as the boab and also known by a number of other names, is a tree in the family Malvaceae, endemic to the northern regions of Western Australia and the Northern Territory of Australia.

<i>Phytolacca dioica</i> Species of tree

Phytolacca dioica, commonly known as ombú in Spanish and umbu in Portuguese, is a massive evergreen tree in the Pokeweed Family (Phytolaccaceae) native to the Pampas of South America. As its specific epithet suggests, it is dioecious, with male and female flowers on separate plants. The flowers are pollinated by the butterfly Doxocopa laurentia. It has an umbrella-like canopy that spreads to a diameter of 12 to 15 meters and can attain a height of 12 to 18 meters. This upper growth springs up from a tuberous caudex which, according to Everett "may occupy a circle sixty feet [18 meters] in diameter". One tree of such a size is mentioned by Anglo-Argentine writer William Henry Hudson in his autobiography " Far Away and Long Ago", which was fifty feet girth above the caudex. Because it is derived from herbaceous ancestors, its trunk consists of anomalous secondary thickening rather than true wood. As a result, the ombú grows fast but its wood is soft and spongy enough to be cut with a knife. These properties have led it to be used in the art of bonsai, as it is easily manipulated to create the desired effect. Since the sap is poisonous, the ombú is not grazed by cattle and is immune to locusts and other pests. For similar reasons, the leaves are sometimes used as a laxative or purgative. It is a symbol of Uruguay, Rio Grande do Sul and Argentina, and of gaucho culture, as its canopy is quite distinguishable from afar and provides comfort and shelter from sun and rain.

A system of plant taxonomy, the Goldberg system was published in:

<i>Adansonia digitata</i> Species of plant

Adansonia digitata, the African baobab, is the most widespread tree species of the genus Adansonia, the baobabs, and is native to the African continent and the southern Arabian Peninsula. These are long-lived pachycauls; radiocarbon dating has shown some individuals to be over 2,000 years old. They are typically found in dry, hot savannas of sub-Saharan Africa, where they dominate the landscape and reveal the presence of a watercourse from afar. They have traditionally been valued as sources of food, water, health remedies or places of shelter and are a key food source for many animals. They are steeped in legend and superstition. In recent years, many of the largest, oldest trees have died, for unknown reasons. Common names for the baobab include monkey-bread tree, upside-down tree, and cream of tartar tree.

<i>Adenium obesum</i> Species of plant

Adenium obesum, more commonly known as a desert rose, is a poisonous species of flowering plant belonging to the tribe Nerieae of the subfamily Apocynoideae of the dogbane family, Apocynaceae. It is native to the Sahel regions south of the Sahara, tropical and subtropical eastern and southern Africa, as well as the Arabian Peninsula. Other names for the flower include Sabi star, kudu, mock azalea, and impala lily. Adenium obesum is a popular houseplant and bonsai in temperate regions.

<i>Adansonia madagascariensis</i> Species of flowering plant

Adansonia madagascariensis or Madagascar baobab is a small to large deciduous tree in the family Malvaceae. It is one of six species of baobab endemic to Madagascar, where it occurs in the Madagascar dry deciduous forests.

<i>Borassus aethiopum</i> Species of palm

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The Palmetum of Santa Cruz de Tenerife is a botanical garden of 120,000 m2 specialized in palms (Arecaceae). It is an artificial hill, with views of the ocean, located in Santa Cruz de Tenerife, Canary Islands, Spain. The gardens include a large system of waterfalls, streams and ponds, a museum dedicated to palms, and a display shade house. The project was started in 1995 on a former landfill and only opened to the public in 2014. The valuable palm collection gathers about 600 species of palms and it is focused on the ones native to world islands. Trees and shrubs of other plant families are also displayed, organised in "biogeographical sections". All gardens are maintained with no pesticides and no fertilizers, different species of wild birds are easily seen in the palmetum.

<i>Adansonia grandidieri</i> Species of tree from Madagascar

Adansonia grandidieri is the biggest and most famous of Madagascar's six species of baobabs. It is sometimes known as Grandidier's baobab or the giant baobab. In French it is called Baobab malgache. The local name is renala or reniala. This tree is endemic to the island of Madagascar, where it is an endangered species threatened by the encroachment of agricultural land. This is the tree found at the Avenue of the Baobabs.

<i>Adansonia za</i> Species of flowering plant

Adansonia za is a species of baobab in the genus Adansonia of the family Malvaceae. It was originally named in French as anadzahé. Common names in Malagasy include bojy, boringy, bozy, bozybe, ringy, and za, the last of which gives the plant its specific epithet. Eight Adansonia species are recognized, with six endemic to Madagascar. Adansonia za is the most widespread of the Madagascar endemics.

The Melchior system, "a reference in all taxonomic courses", is a classification system detailing the taxonomic system of the Angiospermae according to A. Engler's Syllabus der Pflanzenfamilien (1964), also known as "modified or updated" Engler system.

<span class="mw-page-title-main">Succulent plant</span> Plants adapted to arid conditions

In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word succulent comes from the Latin word sucus, meaning "juice" or "sap".

<i>Dendrosicyos</i> Species of plant

Dendrosicyos is a monotypic genus in the plant family Cucurbitaceae. The only species is Dendrosicyos socotranus, the cucumber tree. The species is endemic to the island of Socotra in Yemen, and is the only species in the Cucurbitaceae to grow in a tree form. The species name was originally spelled D. socotrana, but this is corrected to masculine grammatical gender according to the International Code of Nomenclature for algae, fungi, and plants.

<span class="mw-page-title-main">Sunland Baobab</span> Baobab tree in South Africa

Sunland Baobab is a well-known enormous baobab in South Africa. The tree is located on Sunland Farm, near Modjadjiskloof, Limpopo Province. In one study the tree was carbon-dated and found to be an estimated 1,060 years old, plus or minus 75 years. Results of other studies have however suggested much higher ages. The tree used to bloom profusely in spring, at some stage providing a refuge to two pairs of owls, and other bird species. Most of the tree died in 2016 and 2017.

References

  1. Gupta, I. C.; S. K. Gupta (1992). Concepts' Dictionary Of Agricultural Sciences. Concept Publishing Company. p. 348. ISBN   978-81-7022-301-6.
  2. Stearn, W.T. (1992). Botanical Latin: History, grammar, syntax, terminology and vocabulary, Fourth edition. David and Charles.
  3. Von Mueller, Ferdinand (1881). Select Extra-tropical Plants. Sydney: Government Printer. p. 50.
  4. Kunkel, Reinhard (1982). Elephants. New York: Harry N. Abrams Inc. pp. Color Plate on pages 100–101. Includes two adult bull Savanna Elephants for size comparison.
  5. Carder, Dr. Al (2005). Giant Trees of Western America and the World. Madeira Park, British Columbia: Harbour Pub. Co. Ltd. p. 130.
  6. anonymous (January 1957). "<not recorded>". Principes. 1 (2): 57.
  7. Caradine, Chris (November 1998). "A Day at Ocoa". The Palm Journal. [no volumes] (143): 20 with photo.
  8. Riverside (California) Sunday Press Enterprise (September 6, 1964) page 39 Excelent photo with three people for size comparison.
  9. Earle, Christopher. "Gymnosperm Database - Cycadales" . Retrieved November 24, 2021.
  10. Corner, Prof. E.J.H. (1966). Natural History of Palms. Berkeley, Calif.: Univ. Calif. Press. p. 289.
  11. Britton, Nathan L.; Rose, Joseph N. (1963). The Cactaceae - Volume 3 (reprint ed.). New York: Dover Pubs. Inc. p. 170.
  12. Esterhuyse, Neels; et al. (2001). Remarkable Trees of South Africa. Pretoria: Briza Pubs. pp. 6 (table) and 156 & 159.
  13. Wickens, G. E.; Pat Lowe (2008). The Baobabs: Pachycauls of Africa, Madagascar and Australia. Springer. pp. 139–140. ISBN   978-1-4020-6430-2.