| Neoephemera antiqua Temporal range: | |
|---|---|
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Ephemeroptera |
| Family: | Neoephemeridae |
| Genus: | Neoephemera |
| Species: | †N. antiqua |
| Binomial name | |
| †Neoephemera antiqua Sinitchenkova, 1999 | |
Neoephemera antiqua is an extinct species of square-gill mayfly in the family Neoephemeridae that is known from early Eocene, Ypresian stage, [1] lake deposits near the small community of Republic in Ferry County, Washington, USA. [2]
Neoephemera antiqua is known only from one fossil, the holotype, number "UWBM76324". It is a single, mostly complete naiad of undetermined sex, preserved as a compression fossil in fine grained shale. [2] The fossil specimen is from the University of Washington site number UWBM A0307B [2] which works sediments from the Tom thumb tuff member [3] of the Klondike Mountain Formation. Outcrops of the formation are found in and around Republic. The type specimen is currently preserved in the paleoentomology collections housed in the Burke Museum of Natural History and Culture, part of the University of Washington in Seattle, Washington, USA. N. antiqua was first studied by Nina D. Sinitchenkova of the Paleontological Institute of the Russian Academy of Sciences, with her 1999 type description being published in the Russian text Palaeontological Journal. [2] The specific epithet antiqua was coined from the Latin "antiquus", meaning old. [2]
When the holotype of Neoephemera antiqua was first described by Sinitchenkova, The Klondike Mountain Formation as assigned a Middle Eocene age. [2] Further refinement of the dating has resulted in the formation being given a slightly older age, placing it in the Ypresian stage of the late Early Eocene. [1] [4] Neoephemera antiqua is the oldest known occurrence of the family Neoephemeridae, being older than the Oligocene species Potamanthellus rubiens , described from Montana in 1977 by Standley Lewis. [2]
The compression fossil of the naiad is preserved dorsal side up and is nearly complete, only missing the head, ends of the fore and middle legs, and the ends of the three caudal filaments. The full naiad is estimated to have been 8.5 millimetres (0.33 in) in length, and is distinguishable from other species of neoepherids by is shorter pronotum and the structure of the posterior margins of the abdominal segments. The side margins of the pronotum are moderately dilated. The gill opercula have a diagonal rib and the anterolateral angles of the mesonotum have distance processes. Both of these features are only found in the modern genus Neoephemera leading to the placement of the fossil species in the genus. [2]
Ulteramus is an extinct genus of parasitic wasp in the sawfly family Pamphiliidae. The genus is solely known from an Eocene fossil found in North America. At the time of its description the new genus was composed of a single species, Ulteramus republicensis.
Abies milleri, an extinct species of fir known from fossil remains found in deposits from the early Eocene Ypresian stage in Washington State, USA, is the oldest confirmed record for the fir genus.
Allorapisma is an extinct genus of lacewing in the moth lacewings family Ithonidae. The genus is solely known from two Eocene fossils found in North America. At the time of description the genus was composed of a single species, Allorapisma chuorum.
Trochodendron nastae is an extinct species of flowering plant in the family Trochodendraceae known from fossil leaves found in the early Eocene Ypresian stage Klondike Mountain Formation deposits of northern Washington state. T. nastae is one of the oldest members of the genus Trochodendron, which includes the living species T. aralioides, native to Japan, southern Korea and Taiwan and the coeval extinct species T. drachukii from the McAbee Fossil Beds near Cache Creek, British Columbia.
Sassafras hesperia is an extinct species of flowering plant in the family Lauraceae.
Corylus johnsonii is an extinct species of hazel known from fossil fruits found in the Klondike Mountain Formation deposits of northern Washington state, dated to the early Eocene Ypresian stage. Based on described features, C. johnsonii is the oldest definite species in the genus Corylus.
Trochodendron drachukii is an extinct species of flowering plants in the family Trochodendraceae known from a fossil fruiting structure found in the early Ypresian age Eocene fossils found in British Columbia, Canada. T. drachukii is one of the oldest members of the genus Trochodendron, which includes the living species T. aralioides, native to Japan, southern Korea and Taiwan and the coeval extinct species T. nastae from Washington state, United States.
Neviusia dunthornei is an extinct species of flowering plants in the family Rosaceae. The species is solely known from the early Eocene, Ypresian stage, Allenby Formation Lacustrine deposits near the town of Princeton, British Columbia.
Acer stonebergae is an extinct maple species in the family Sapindaceae described from two fossil samaras. The species is solely known from the Early Eocene sediments exposed in northeast Washington state, United States, and the adjacent area of south central British Columbia, Canada. It is one of three species belonging to the extinct section Torada.
Fothergilla malloryi is an extinct species of flowering plant in the family Hamamelidaceae known from fossil leaves found in the early Eocene Klondike Mountain Formation deposits of northern Washington state. The F. malloryi leaves are the earliest appearance in the fossil record of a member of the witchalder genus Fothergilla, which includes the living species F. gardenii, and F. major, both of which are native to the southeastern United States. The genus also includes three or four other fossil species with two Asian Miocene species, F. viburnifolia from China, F. ryozenensis from Japan along with one Miocene North American species, F. praeolata of Oregon. Fothergilla durhamensis described from Eocene sediments in King County, Washington is considered dubious in placement, and it was transferred to the genus Platimeliphyllum by Huegele et al. (2021).
Corylopsis reedae is an extinct species of flowering plant in the family Hamamelidaceae known from fossil leaves found in the early Eocene Klondike Mountain Formation deposits of northern Washington state. C. readae is one of the oldest occurrences of the winter-hazel genus Corylopsis, which includes between seven and thirty species, all found in Asia. Fossils from two other occurrences are of similar age to C. readae, with Paleocene specimens from Greenland being placed in the form taxon Corylopsiphyllum and an Eocene Alaskan fossil being included in Corylopsis without species placement.
Rhus malloryi is an extinct species of flowering plant in the sumac family Anacardiaceae. The species is known from fossil leaves found in the early Eocene deposits of northern Washington state, United States. The species was first described from a series of isolated fossil leaves in shale. R. malloryi is one of four sumac species to be described from the Klondike Mountain Formation, and forms a hybrid complex with the other three species.
Neoephemera is a genus of large squaregill mayflies in the family Neoephemeridae first described by McDunnough (1925). and containing approximately six described species in Neoephemera.
Cretomerobius is an extinct genus of lacewings in the neuropteran family Hemerobiidae known from fossils found in Asia. The genus currently contains a single species, the Aptian C. disjunctus.
Paraconcavistylon is an extinct genus of flowering plant in the family Trochodendraceae comprises a single species, Paraconcavistylon wehrii. The genus is known from fossil fruits and leaves found in the early Eocene deposits of northern Washington state, United States, and southern British Columbia, Canada. The species was initially described as a member of the related extinct genus Concavistylon as "Concavistylon" wehrii, but subsequently moved to the new genus Paraconcavistylon in 2020 after additional study.
Pseudolarix wehrii is an extinct species of golden larch in the pine family (Pinaceae). The species is known from early Eocene fossils of northern Washington state, United States, and southern British Columbia, Canada, along with late Eocene mummified fossils found in the Qikiqtaaluk Region, Nunavut, Canada.
Carpinus perryae is an extinct species of hornbeam known from fossil fruits found in the Klondike Mountain Formation deposits of northern Washington state, dated to the early Eocene Ypresian stage. Based on described features, C. perryae is the oldest definite species in the genus Carpinus.
Klondikia is an extinct hymenopteran genus in the ant family Formicidae with a single described species Klondikia whiteae. The species is solely known from the Early Eocene sediments exposed in northeast Washington state, United States. The genus is currently not placed into any ant subfamily, being treated as incertae sedis.
The paleofauna of the Eocene Okanagan Highlands is comprised of Early Eocene arthropods, vertebrates, plus rare nematodes and molluscs found in geological formations of the northwestern North American Eocene Okanagan Highlands. The highlands lake bed series' as a whole are considered one of the great Canadian Lagerstätten. The paleofauna represents that of a late Ypresian upland temperate ecosystem immediately after the Paleocene-Eocene thermal maximum, and before the increased cooling of the middle and late Eocene to Oligocene. The fossiliferous deposits of the region were noted as early as 1873, with small amounts of systematic work happening in the 1880-90s on British Columbian sites, and 1920-30s for Washington sites. Focus and more detailed descriptive work on the Okanagan Highlands site started in the last 1970's. Most of the highlands sites are preserved as compression-impression fossils in "shales", but also includes a rare permineralized biota and an amber biota.