Spectral sensitivity

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Spectral sensitivities (normalized responsivity spectra) of human cone cells, S, M, and L types Cones SMJ2 E.svg
Spectral sensitivities (normalized responsivity spectra) of human cone cells, S, M, and L types
1916 plot of "spectral sensibilities." The author also uses the more modern term "spectral sensitivity" in the same book. Spectral sensibilities.png
1916 plot of "spectral sensibilities." The author also uses the more modern term "spectral sensitivity" in the same book.

Spectral sensitivity is the relative efficiency of detection, of light or other signal, as a function of the frequency or wavelength of the signal.

In visual neuroscience, spectral sensitivity is used to describe the different characteristics of the photopigments in the rod cells and cone cells in the retina of the eye. It is known that the rod cells are more suited to scotopic vision and cone cells to photopic vision, and that they differ in their sensitivity to different wavelengths of light. [2] [3] It has been established that the maximum spectral sensitivity of the human eye under daylight conditions is at a wavelength of 555  nm, while at night the peak shifts to 507 nm. [4]

In photography, film and sensors are often described in terms of their spectral sensitivity, to supplement their characteristic curves that describe their responsivity. [5] A database of camera spectral sensitivity is created and its space analyzed. [6] For X-ray films, the spectral sensitivity is chosen to be appropriate to the phosphors that respond to X-rays, rather than being related to human vision. [7]

In sensor systems, where the output is easily quantified, the responsivity can be extended to be wavelength dependent, incorporating the spectral sensitivity. When the sensor system is linear, its spectral sensitivity and spectral responsivity can both be decomposed with similar basis functions. [8] When a system's responsivity is a fixed monotonic nonlinear function, that nonlinearity can be estimated and corrected for, to determine the spectral sensitivity from spectral input–output data via standard linear methods. [9]

The responses of the rod and cone cells of the retina, however, have a very context-dependent (coupled) nonlinear response, which complicates the analysis of their spectral sensitivities from experimental data. [10] In spite of these complexities, however, the conversion of light energy spectra to the effective stimulus, the excitation of the photopigment, is quite linear, and linear characterizations such as spectral sensitivity are therefore quite useful in describing many properties of color vision. [11]

Spectral sensitivity is sometimes expressed as a quantum efficiency, that is, as probability of getting a quantum reaction, such as a captured electron, to a quantum of light, as a function of wavelength. [12] In other contexts, the spectral sensitivity is expressed as the relative response per light energy, rather than per quantum, normalized to a peak value of 1, and a quantum efficiency is used to calibrate the sensitivity at that peak wavelength. [13] In some linear applications, the spectral sensitivity may be expressed as a spectral responsivity, with units such as amperes per watt. [14] [15] [16]

See also

Related Research Articles

<span class="mw-page-title-main">Color</span> Characteristic of visual perception

Color or colour is the visual perceptual property deriving from the spectrum of light interacting with the photoreceptor cells of the eyes. Color categories and physical specifications of color are associated with objects or materials based on their physical properties such as light absorption, reflection, or emission spectra. By defining a color space, colors can be identified numerically by their coordinates.

Responsivity measures the input–output gain of a detector system. In the specific case of a photodetector, it measures the electrical output per optical input.

<span class="mw-page-title-main">Photodiode</span> Converts light into current

A photodiode is a light-sensitive semiconductor diode. It produces current when it absorbs photons.

<span class="mw-page-title-main">Metamerism (color)</span> Perceived matching of colors in colorimetry

In colorimetry, metamerism is a perceived matching of colors with different (nonmatching) spectral power distributions. Colors that match this way are called metamers.

<span class="mw-page-title-main">Night vision</span> Ability to see in low light conditions

Night vision is the ability to see in low-light conditions, either naturally with scotopic vision or through a night-vision device. Night vision requires both sufficient spectral range and sufficient intensity range. Humans have poor night vision compared to many animals such as cats, in part because the human eye lacks a tapetum lucidum, tissue behind the retina that reflects light back through the retina thus increasing the light available to the photoreceptors.

Luminous efficiency function Description of the average spectral sensitivity of human visual perception of brightness

A luminous efficiency function or luminosity function represents the average spectral sensitivity of human visual perception of light. It is based on subjective judgements of which of a pair of different-colored lights is brighter, to describe relative sensitivity to light of different wavelengths. It is not an absolute reference to any particular individual, but is a standard observer representation of visual sensitivity of theoretical human eye. It is valuable as a baseline for experimental purposes, and in colorimetry. Different luminous efficiency functions apply under different lighting conditions, varying from photopic in brightly lit conditions through mesotopic to scotopic under low lighting conditions. When not specified, the luminous efficiency function generally refers to the photopic luminous efficiency function.

<span class="mw-page-title-main">Color vision</span> Ability to perceive differences in light frequency

Color vision, a feature of visual perception, is an ability to perceive differences between light composed of different wavelengths independently of light intensity. Color perception is a part of the larger visual system and is mediated by a complex process between neurons that begins with differential stimulation of different types of photoreceptors by light entering the eye. Those photoreceptors then emit outputs that are propagated through many layers of neurons and then ultimately to the brain. Color vision is found in many animals and is mediated by similar underlying mechanisms with common types of biological molecules and a complex history of evolution in different animal taxa. In primates, color vision may have evolved under selective pressure for a variety of visual tasks including the foraging for nutritious young leaves, ripe fruit, and flowers, as well as detecting predator camouflage and emotional states in other primates.

Photoreceptor cell Type of neuroepithelial cell

A photoreceptor cell is a specialized type of neuroepithelial cell found in the retina that is capable of visual phototransduction. The great biological importance of photoreceptors is that they convert light into signals that can stimulate biological processes. To be more specific, photoreceptor proteins in the cell absorb photons, triggering a change in the cell's membrane potential.

<span class="mw-page-title-main">Tetrachromacy</span> Type of color vision with four types of cone cells

Tetrachromacy is the condition of possessing four independent channels for conveying color information, or possessing four types of cone cell in the eye. Organisms with tetrachromacy are called tetrachromats.

Trichromacy Possessing of three independent channels for conveying color information

Trichromacy or trichromatism is the possessing of three independent channels for conveying color information, derived from the three different types of cone cells in the eye. Organisms with trichromacy are called trichromats.

<span class="mw-page-title-main">Rod cell</span> Photoreceptor cells that can function in lower light better than cone cells

Rod cells are photoreceptor cells in the retina of the eye that can function in lower light better than the other type of visual photoreceptor, cone cells. Rods are usually found concentrated at the outer edges of the retina and are used in peripheral vision. On average, there are approximately 92 million rod cells in the human retina. Rod cells are more sensitive than cone cells and are almost entirely responsible for night vision. However, rods have little role in color vision, which is the main reason why colors are much less apparent in dim light.

<span class="mw-page-title-main">Cone cell</span> Photoreceptor cells responsible for color vision made to function in bright light

Cone cells, or cones, are photoreceptor cells in the retinas of vertebrate eyes including the human eye. They respond differently to light of different wavelengths, and the combination of their responses is responsible for color vision. Cones function best in relatively bright light, called the photopic region, as opposed to rod cells, which work better in dim light, or the scotopic region. Cone cells are densely packed in the fovea centralis, a 0.3 mm diameter rod-free area with very thin, densely packed cones which quickly reduce in number towards the periphery of the retina. Conversely, they are absent from the optic disc, contributing to the blind spot. There are about six to seven million cones in a human eye, with the highest concentration being towards the macula.

In visual physiology, adaptation is the ability of the retina of the eye to adjust to various levels of light. Natural night vision, or scotopic vision, is the ability to see under low-light conditions. In humans, rod cells are exclusively responsible for night vision as cone cells are only able to function at higher illumination levels. Night vision is of lower quality than day vision because it is limited in resolution and colors cannot be discerned; only shades of gray are seen. In order for humans to transition from day to night vision they must undergo a dark adaptation period of up to two hours in which each eye adjusts from a high to a low luminescence "setting", increasing sensitivity hugely, by many orders of magnitude. This adaptation period is different between rod and cone cells and results from the regeneration of photopigments to increase retinal sensitivity. Light adaptation, in contrast, works very quickly, within seconds.

<span class="mw-page-title-main">Melanopsin</span> Mammalian protein found in Homo sapiens

Melanopsin is a type of photopigment belonging to a larger family of light-sensitive retinal proteins called opsins and encoded by the gene Opn4. In the mammalian retina, there are two additional categories of opsins, both involved in the formation of visual images: rhodopsin and photopsin in the rod and cone photoreceptor cells, respectively.

<span class="mw-page-title-main">Photodetector</span> Sensors of light or other electromagnetic energy

Photodetectors, also called photosensors, are sensors of light or other electromagnetic radiation. There is a wide variety of photodetectors which may be classified by mechanism of detection, such as photoelectric or photochemical effects, or by various performance metrics, such as spectral response. Semiconductor-based photodetectors typically photo detector have a p–n junction that converts light photons into current. The absorbed photons make electron–hole pairs in the depletion region. Photodiodes and photo transistors are a few examples of photo detectors. Solar cells convert some of the light energy absorbed into electrical energy.

In the study of human visual perception, scotopic vision is the vision of the eye under low-light conditions. The term comes from Greek skotos, meaning "darkness", and -opia, meaning "a condition of sight". In the human eye, cone cells are nonfunctional in low visible light. Scotopic vision is produced exclusively through rod cells, which are most sensitive to wavelengths of around 498 nm (blue-green) and are insensitive to wavelengths longer than about 640 nm (red-orange). This condition is called the Purkinje effect.

Abney effect

The Abney effect or the purity-on-hue effect describes the perceived hue shift that occurs when white light is added to a monochromatic light source.

Intrinsically photosensitive retinal ganglion cells (ipRGCs), also called photosensitive retinal ganglion cells (pRGC), or melanopsin-containing retinal ganglion cells (mRGCs), are a type of neuron in the retina of the mammalian eye. The presence of ipRGCs was first suspected in 1927 when rodless, coneless mice still responded to a light stimulus through pupil constriction, This implied that rods and cones are not the only light-sensitive neurons in the retina. Yet research on these cells did not advance until the 1980s. Recent research has shown that these retinal ganglion cells, unlike other retinal ganglion cells, are intrinsically photosensitive due to the presence of melanopsin, a light-sensitive protein. Therefore they constitute a third class of photoreceptors, in addition to rod and cone cells.

The Stiles–Crawford effect is a property of the human eye that refers to the directional sensitivity of the cone photoreceptors.

<span class="mw-page-title-main">Visual perception</span> Ability to interpret the surrounding environment using light in the visible spectrum

Visual perception is the ability to interpret the surrounding environment through photopic vision, color vision, scotopic vision, and mesopic vision, using light in the visible spectrum reflected by objects in the environment. This is different from visual acuity, which refers to how clearly a person sees. A person can have problems with visual perceptual processing even if they have 20/20 vision.

References

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