Stegotetrabelodon

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Stegotetrabelodon
Temporal range: Late Miocene to Early Pliocene, 8–4.2  Ma
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Stegotetrabelodon11.jpg
Restoration of Stegotetrabelodon syrticus
Stegotetrabelodon molar.jpg
Stegotetrabelodon molar tooth
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Family: Elephantidae
Genus: Stegotetrabelodon
Petrocchi, 1941
Type species
Stegotetrabelodon syrticus
Petrocchi, 1941
Species
  • S. syrticus Petrocchi, 1941
  • S. orbus Maglio, 1970
  • S. lybicus Petrocchi, 1943
  • S. emiratus Sanders, 2022

Stegotetrabelodon is an extinct genus of primitive elephantid from the Late Miocene to Early Pliocene of Africa, the Arabian Peninsula, and Italy.

Species

The earliest species S. emiratus is known from the Late Miocene Baynunah Formation of the United Arab Emirates on the Arabian Peninsula, dating to around 8-6 million years ago. S. orbus is known from the Late Miocene-Early Pliocene in East Africa (Kenya, Uganda), spanning from around 7.5 million years ago to possibly as late as 4.2 million years ago. The species S. syrticus is known from the Sahabi site in Libya, dating to around 6.8–5.3 million years ago, with remains of the species also reported from the Late Miocene of Calabria in southern Italy, which was likely part of Africa during this time. [1]

"Stegotetrabelodon" maluvalensis from the Late Miocene of Pakistan is now placed in Stegolophodon, while "Stegatetrabelodon" gigantorostris from the Late Miocene of China is now placed in Tetralophodon . [2]

Description

An estimate based on the size of limb bones collected from Sahabi suggests that S. syrticus may have reached 4 m (13.1 ft) in shoulder height and 11–12 tonnes (12.1–13.2 short tons) in weight, [3] with a similar estimate of 3.7 m (12.1 ft) and 10–11 tonnes (11.0–12.1 short tons) made for S. emiratus. [1] In comparison to later elephantids, Stegotetrabelodon has several primitive features, including the retention of permanent premolar teeth, an elongated mandibular symphysis and large lower tusks, shared with its gomphothere ancestors. [4] The lower tusks are the longest known among proboscideans, reaching a length of 2.2 metres (7.2 ft) in S.syrticus, though they weighed only around 12.5 kilograms (28 lb), with the upper tusks also being large, reaching a length of 2.8 metres (9.2 ft) and weight exceeding 40 kilograms (88 lb) in S. syrticus. [5]

Ecology

In December 2006, a small contingent of Emirates Natural History Group members found several sets of proboscidean footprints thought to have probably been produced by Stegotetrabelodon preserved in a large calcareous exposure between the sand dunes of "Mleisa 1" [6] in the Western Region [7] of the Emirate of Abu Dhabi, the United Arab Emirates. The longest of these tracks extend for 290 m (950 ft) and 170 m (560 ft). [8] These tracks suggest that like modern elephants, Stegotetrabelodon lived in herds. [9]

Related Research Articles

<span class="mw-page-title-main">Proboscidea</span> Order of elephant-like mammals

Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Proboscideans include some of the largest known land mammals. The largest land mammal of all time may have been a proboscidean; the elephant Palaeoloxodon namadicus has been estimated to be up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, surpassing the paraceratheres, the otherwise largest known land mammals, though this estimate was made based on a single fragmentary femur and is speculative. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.

<span class="mw-page-title-main">Mastodon</span> Extinct genus of proboscideans

A mastodon is a member of the genus Mammut, which strictly defined, was endemic to North America and lived from the late Miocene to the early Holocene. Mastodons belong to the order Proboscidea, the same order as elephants and mammoths. Mammut is the type genus of the extinct family Mammutidae, which diverged from the ancestors of modern elephants at least 27-25 million years ago, during the Oligocene.

<span class="mw-page-title-main">Elephantidae</span> Family of mammals

Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.

<span class="mw-page-title-main">Mammutidae</span> Extinct family of mammals

Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons, which inhabited North America from the Late Miocene until their extinction at beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, μαστός "nipple" and ὀδούς "tooth", referring to their characteristic teeth.

<i>Amebelodon</i> Extinct genus of mammals

Amebelodon is a genus of extinct proboscidean belonging to Amebelodontidae. The most striking attribute of this animal is its lower tusks, which are narrow, elongated, and distinctly flattened with the degree of flattening varying among the different species. One valid species is known for this genus, which was endemic to North America. Other species once assigned to Amebelodon are now assigned to the genus Konobelodon, which was once a subgenus.

<i>Stegodon</i> Genus of extinct proboscidean

Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon remained widespread in South, Southeast and East Asia until the end of the Pleistocene.

<span class="mw-page-title-main">Gomphothere</span> Extinct family of proboscidean mammals

Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to western South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.

<span class="mw-page-title-main">Deinotheriidae</span> Prehistoric family of mammals

Deinotheriidae is a family of prehistoric elephant-like proboscideans that lived during the Cenozoic era, first appearing in Africa, then spreading across southern Asia (Indo-Pakistan) and Europe. During that time, they changed very little, apart from growing much larger in size; by the late Miocene, they had become the largest land animals of their time. Their most distinctive features were their lack of upper tusks and downward-curving tusks on the lower jaw.

<i>Gomphotherium</i> Extinct genus of elephant-like mammals

Gomphotherium is an extinct genus of gomphothere proboscidean from the Neogene of Eurasia, Africa and North America. The genus is probably paraphyletic.

<i>Anancus</i> Genus of proboscideans

Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.

<i>Tetralophodon</i> Extinct genus of mammals

Tetralophodon is an extinct genus of "tetralophodont gomphothere" belonging to the superfamily Elephantoidea, known from the Miocene of Afro-Eurasia.

Primelephas is a genus of Elephantinae that existed during the Miocene and Pliocene epochs. The name of the genus suggests 'first elephant'. These primitive elephantids are thought to be the common ancestor of Mammuthus, the mammoths, and the closely allied genera Elephas and Loxodonta, the Asian and African elephants, diverging some 4-6 million years ago. It had four tusks, which is a trait not shared with its descendants, but common in earlier proboscideans. The type species, Primelephas gomphotheroides, was described by Vincent Maglio in 1970, with the specific epithet indicating the fossil specimens were gomphothere-like. Primelephas korotorensis is the only other species to be assigned to the genus. All fossils found of the Primelephas have been found in Africa, primarily in modern day Chad, Tanzania, Kenya, Ethiopia, and Uganda.

<i>Rhynchotherium</i> Extinct genus of proboscid

Rhynchotherium is an extinct genus of proboscidea endemic to North America and Central America during the Miocene through Pliocene from 13.650 to 3.6 Ma, living for approximately 10 million years.

<i>Archaeobelodon</i>

Archaeobelodon is an extinct genus of proboscidean of the family Amebelodontidae that lived in Europe and North Africa (Egypt) during the Miocene from 16.9 to 16.0 Ma, living for approximately 0.9 million years.

<i>Eubelodon</i> Extinct genus of proboscid

Eubelodon is an extinct genus of gomphothere which lived in North America during the Miocene Epoch. It contains a single species: Eubelodon morrilli.

<span class="mw-page-title-main">Stegodontidae</span> Family of extinct elephant-like mammals

Stegodontidae is an extinct family of proboscideans from Africa and Asia from the Early Miocene to the Late Pleistocene. It contains two genera, the earlier Stegolophodon, known from the Miocene of Asia and the later Stegodon, from the Late Miocene to Late Pleistocene of Africa and Asia which is thought to have evolved from the former. The group is noted for their plate-like lophs on their teeth, which are similar to elephants and different from those of other extinct proboscideans like gomphotheres and mammutids, with both groups having a proal jaw movement utilizing forward strokes of the lower jaw. These similarities with modern elephants were probably convergently evolved. Like elephantids, stegodontids are thought to have evolved from gomphothere ancestors.

The Baynunah Formation is a palaeontological formation located in the western region of the Emirate of Abu Dhabi, the United Arab Emirates. It dates to the Upper Miocene epoch of the Neogene period in the Cenozoic Era.

<span class="mw-page-title-main">Al Dhafra, Abu Dhabi</span> The Gharbiyyah (Western) Region of the Emirate of Abu Dhabi

Al Dhafra Region, known until 2017 as Al Gharbia or the Western Region, is one of three Municipal Regions in the Emirate of Abu Dhabi. Forming the western part of the United Arab Emirates, it is by far the largest region by area, occupying 71 % of the Abu Dhabi emirate's total area, yet the smallest by population and population density, and thus a rather remote region of the Emirate of Abu Dhabi, similar to the Eastern Region. The capital of Al Dhafra Region is Madinat Zayed. The new official name was already the historical name of the westernmost region of former Trucial Oman, which was Dhafrah. It is significant for its natural resources, particularly gas and petroleum. Its reserves of hydrocarbons account for 90% of the Emirate's reserves, which in turn account for 90% of the country's reserves, and are important for the local economy.

<i>Choerolophodon</i> Extinct genus of mammals

Choerolophodon is an extinct genus of proboscidean that lived during the Miocene of Eurasia and Africa. Fossils of Choerolophodon have been found in Africa, Southeast Europe, Turkey, Iraq, Iran, the Indian subcontinent, and China.

<span class="mw-page-title-main">Amebelodontidae</span> Extinct family of mammals

Amebelodontidae is an extinct family of large herbivorous proboscidean mammals related to elephants. They were formerly assigned to Gomphotheriidae, but recent authors consider them a distinct family. They are distinguished from other proboscideans by having flattened lower tusks and very elongate mandibular symphysis. The lower tusks could grow considerable size, with those of Konobelodon reaching 1.61 metres (5.3 ft) in length. Their molar teeth are typically trilophodont, and possessed posttrite conules. In the past, amebelodonts' shovel-like mandibular tusks led to them being portrayed scooping up water plants, however, dental microwear suggests that they were browsers and mixed feeders. The lower tusks have been proposed to have had a variety of functions depending on the species, including stripping bark, cutting through vegetation, as well as possibly digging. They first appeared in Africa during the Early Miocene, and subsequently dispersed into Eurasia and then North America. They became extinct by the beginning of the Pliocene. While some phylogenetic studies have recovered Amebelodontidae as a monophyletic group that forms the sister group to Gomphotheriidae proper, some authors have argued that Amebelodontidae may be polyphyletic, with it being suggested that the shovel-tusked condition arose several times independently within Gomphotheriidae, thus rendering the family invalid.

References

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  2. Wang, Shi-Qi; Saegusa, Haruo; Duangkrayom, Jaroon; He, Wen; Chen, Shan-Qin (December 2017). "A new species of Tetralophodon from the Linxia Basin and the biostratigraphic significance of tetralophodont gomphotheres from the Upper Miocene of northern China". Palaeoworld. 26 (4): 703–717. doi:10.1016/j.palwor.2017.03.005.
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  7. "Des empreintes de 7 millions d'années racontent les éléphants d'autrefois". Futura (in French). Retrieved 2018-10-11.
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