Cephalothorax

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Bauplan of a malacostracan; the cephalothorax is the region including cephalon and thorax, marked in yellow. General malacostracan en.svg
Bauplan of a malacostracan; the cephalothorax is the region including cephalon and thorax, marked in yellow.

The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. [1] (The terms prosoma and opisthosoma are equivalent to cephalothorax and abdomen in some groups. The terms prosoma and opisthosoma may be preferred by some researchers in cases such as arachnids, where there is neither fossil nor embryonic evidence animals in this class have ever had separate heads and thoraxes, and where the opisthosoma contains organs atypical of a true abdomen, such as a heart and respiratory organs [2] ). The word cephalothorax is derived from the Greek words for head ( κεφαλή , kephalé) and thorax (θώραξ, thórax ). [3] This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. [1] In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax. [4]

Contents

Arachnid anatomy

Fovea

The fovea is the centre of the cephalothorax and is located behind the head (only in spiders). [5] It is often important in identification. It can be transverse or procurved [6] and can, in some tarantulas (e.g. Ceratogyrus darlingi ) have a "horn". [7]

Clypeus

The clypeus is the space between the anterior of the cephalothorax and the ocularium. It is found in most arachnids. [6] It is connected to the labrum of the invertebrate, between the labrum and the face.

Ocularium

The ocularium is a "turret" for the ocelli found in most arachnids. [8] In harvestmen, it may have the ornament of spines. [9]

Trident

The trident is a small group of (usually three) spines found in harvestmen exclusively. It is located in front of the ocularium. It varies in size amongst species; in some it is completely absent, and in others it is enlarged considerably. [9]

Related Research Articles

<span class="mw-page-title-main">Chelicerata</span> Subphylum of arthropods

The subphylum Chelicerata constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids, as well as a number of extinct lineages, such as the eurypterids and chasmataspidids.

<span class="mw-page-title-main">Arachnid</span> Class of arthropods

Arachnida is a class of joint-legged arthropods, in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons.

<span class="mw-page-title-main">Amblypygi</span> Order of arachnids

Amblypygi is an order of arachnids also known as whip spiders or tailless whip scorpions, not to be confused with whip scorpions or vinegaroons that belong to the related order Thelyphonida. The name "amblypygid" means "blunt tail", a reference to a lack of the flagellum that is otherwise seen in whip scorpions. Amblypygids possess no silk glands or venomous fangs. They rarely bite if threatened, but can grab fingers with their pedipalps, resulting in thorn-like puncture injuries.

<span class="mw-page-title-main">Opiliones</span> Order of arachnids (harvestmen/daddy longlegs)

The Opiliones are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. As of April 2017, over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014.

<span class="mw-page-title-main">Ricinulei</span> Order of spider-like animals

Ricinulei is a small order of arachnids. Like most arachnids, they are predatory, eating small arthropods. They occur today in west-central Africa (Ricinoides) and the Americas as far north as Texas. As of 2021, 91 extant species of ricinuleids have been described worldwide, all in the single family Ricinoididae. In older works they are sometimes referred to as Podogona. Due to their obscurity they do not have a proper common name, though in academic literature they are occasionally referred to as hooded tickspiders.

<span class="mw-page-title-main">Chelicerae</span> Mouthparts of spiders and horseshoe crabs

The chelicerae are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or as a type of pincers. Some chelicerae, such as those found on nearly all spiders, are hollow and contain venom glands, used to inject venom into prey or a perceived threat. Both pseudoscorpions and harvestmen have additional structures on their chelicerae that are used for grooming.

<span class="mw-page-title-main">Solifugae</span> Order of arachnids

Solifugae is an order of animals in the class Arachnida known variously as camel spiders, wind scorpions, sun spiders, or solifuges. The order includes more than 1,000 described species in about 147 genera. Despite the common names, they are neither true scorpions, nor true spiders. Most species of Solifugae live in dry climates and feed opportunistically on ground-dwelling arthropods and other small animals. The largest species grow to a length of 12–15 cm (5–6 in), including legs. A number of urban legends exaggerate the size and speed of the Solifugae, and their potential danger to humans, which is negligible.

<span class="mw-page-title-main">Carapace</span> Part of exoskeleton in some animals

A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron.

<span class="mw-page-title-main">Pygidium</span> Posterior body part or shield of crustaceans and some other arthropods

The pygidium is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.

<span class="mw-page-title-main">Xiphosura</span> Order of marine chelicerates

Xiphosura is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs. They first appeared in the Hirnantian. Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera.

In biology, a tagma is a specialized grouping of multiple segments or metameres into a coherently functional morphological unit. Familiar examples are the head, the thorax, and the abdomen of insects. The segments within a tagma may be either fused or so jointed as to be independently moveable.

<span class="mw-page-title-main">Trigonotarbida</span> Extinct order of arachnids

The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian. These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton (tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tubercles. About seventy species are currently known, with most fossils originating from the Carboniferous coal measures.

<span class="mw-page-title-main">Sternum (arthropod anatomy)</span> Body part in invertebrates

The sternum is the ventral portion of a segment of an arthropod thorax or abdomen.

<span class="mw-page-title-main">Opisthosoma</span> Posterior body part of some arthropods

The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata. Although it is similar in most respects to an abdomen, the opisthosoma is differentiated by its inclusion of the respiratory organs and the heart.

<span class="mw-page-title-main">Spider anatomy</span> Physiology of Spiders (order Araneae)

The anatomy of spiders includes many characteristics shared with other arachnids. These characteristics include bodies divided into two tagmata, eight jointed legs, no wings or antennae, the presence of chelicerae and pedipalps, simple eyes, and an exoskeleton, which is periodically shed.

<span class="mw-page-title-main">Opiliones anatomy</span>

Opiliones are an order of arachnids and share many common characteristics with other arachnids. However, several differences separate harvestmen from other arachnid orders such as spiders. The bodies of opiliones are divided into two tagmata : the abdomen (opisthosoma) and the cephalothorax (prosoma). Unlike spiders, the juncture between the abdomen and cephalothorax is often poorly defined. Harvestmen have chelicerae, pedipalps and four pairs of legs. Most harvestmen have two eyes, although there are eyeless species.

<span class="mw-page-title-main">Tarantula</span> Family of spiders

Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. As of December 2023, 1,100 species have been identified, with 166 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although many other members of the same infraorder (Mygalomorphae) are commonly referred to as "tarantulas" or "false tarantulas". Some of the more common species have become popular in the exotic pet trade. Many New World species kept as pets have setae known as urticating hairs that can cause irritation to the skin, and in extreme cases, cause damage to the eyes.

<span class="mw-page-title-main">Spider</span> Order of arachnids

Spiders are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except for Antarctica, and have become established in nearly every land habitat. As of November 2023, 51,673 spider species in 136 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900.

This glossary describes the terms used in formal descriptions of spiders; where applicable these terms are used in describing other arachnids.

References

  1. 1 2 Eldra Pearl Solomon, Linda R. Berg & Diana W. Martin (2004). "The animal kingdom: an introduction to animal diversity". Biology (7th ed.). Cengage Learning. pp. 534–549. ISBN   978-0-534-49276-2.
  2. Shultz, Stanley; Shultz, Marguerite (2009). The Tarantula Keeper's Guide. Barron's. p. 23. ISBN   978-0-7641-3885-0.{{cite book}}: CS1 maint: multiple names: authors list (link)
  3. Timothy J. Gibb & C. Y. Oseto (2006). "Glossary". Arthropod Collection and Identification: Field and Laboratory Techniques. Academic Press. ISBN   978-0-12-369545-1.
  4. Andrew J. Martinez (2003). "Arthropoda (crabs, shrimps, lobsters)". Marine Life of the North Atlantic: Canada to New England (3rd ed.). Aqua Quest Publications. pp. 144–175. ISBN   978-1-881652-32-8.
  5. Dalton, Steve (2008). Spiders; The Ultimate Predators. A & C Black, London. P.p. 19. ISBN   9781408106976.
  6. 1 2 Smith, A. M. (1990c). Baboon spiders: Tarantulas of Africa and the Middle East. Fitzgerald Publishing, London, pp. 138. Retrieved February 13, 2016.
  7. Gallon, R.C. (2008). "On some poorly known African Harpactirinae, with notes on Avicuscodra arabica Strand, 1908 and Scodra pachypoda Strand, 1908 (Araneae, Theraphosidae)". Bulletin of the British Arachnological Society. 14: 238.
  8. Spiders... Archived 2021-11-28 at the Wayback Machine Yorkshire Naturalists' Union. Retrieved February 13, 2016.
  9. 1 2 Sankey, John & Savory, Theodore. British Harvestmen. Academic Press. P.p. 1-75. ISBN   012619050X.