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The Cytodeme is the total assembly of organisms that use an identical suite of chromosomes to carry their genes.[ citation needed ] The term was first printed in the 1950s in a book by Heslop-Harrison. [1] Discussing the Deme Terminology [2] - he continued "cytodeme, a population differing in some distinctive cytological feature from others." In most cases the suite is composed of several pairs of homologous chromosomes with or without a pair of sex chromosomes. Since the only acceptable proof of the identity (homology) of chromosomes lies in their ability to pair fully from end to end during meiosis it follows that:
As a general rule for most species all of its members are of the same cytodeme excepting only the infrequent aneuploid aberrants. However some species are known to include several chromosome races which must necessarily belong to different cytodemes. Although it is not intrinsic in the formal definition it is a matter of common observation that members of different cytodemes are essentially cross-incompatible and any hybrids that do arise are usually highly infertile.
The current surgance of interest in cytodemes stems from the realisation that membership of the cytodeme is not necessarily restricted to the members of one species: frequently two or more species are in fact of the same cytodeme. Although fully intercompatible and yielding fertile hybrids when they do cross these species never or rarely cross-breed in the natural environment because they are spatially isolated, geographically and/or ecologically. Typically the species involved are members of the same genus. There are now a substantial number of cases in which two or more species of the same cytodeme are clearly different and even in different genera. Thus all five diploid species recognised in the genus Brassicella and both species recognised in the genus Hutera (7spp. in total) are in the same cytodeme. [3] Perhaps the most extreme case on record concerns the wild grass, teosinte, Euchlaena mexicana and the strikingly different Maize, or Indian Corn, Zea mays , both 2n=20, fully interfertile and yielding fertile hybrids. [4] There is perhaps a remote possibility that two such very different species could have evolved independently from distinct sources and converged in their chromosomal ideotype until they became members of the same cytodeme (including the capacity to cross-breed). It is more likely, though, that the cytodeme arose first complete with its suite of chromosomes and breeding patterns all intact and then, remaining constant in its fundamentals, it diversified into species sometimes so different as to merit generic distinction. Thus, in what may be termed the cytodeme adjunct to Darwinian theory, evolution becomes a two-stage process - first, the establishment of distinct cytodemes reproductively isolated both from one another and from all previously existing cytodemes; second, diversification within cytodeme to yield taxonomically recognisable (but not reproductively isolated) species.
Whereas the second stage is Natural Selection as expounded by Darwin, [5] the first stage is not necessarily Darwinian[ citation needed ]. In the special case of polyploidy it is known that the first stage is not Darwinian. Doubling the chromosome number of the sterile hybrid between two diploid cytodemes yields an allotetraploid which is both fertile within its bounds and essentially incompatible with all previous life forms.
As yet no mechanism is known which could account for the origin of new diploid cytodemes - nor is there any proof that there are any diploid cytodemes of recent origin. Indeed, the evidence is that cytodemes are remarkably long lived. Presumably the Platanus cytodeme existed on Pangaea before that land fragmented. The present day forms of Platanus - P. orientalis in Europe and P. occidentalis in America - are fully intercompatible when brought together artificially in Botanic Gardens yielding fertile hybrids like the London plane, [6] establishing not only that they are in the same cytodeme as one another but also that they are both in the same cytodeme as their common ancestor of approximately 200 million years ago.
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within lineages. Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book On the Origin of Species. He also identified sexual selection as a likely mechanism, but found it problematic.
In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Generally, it means that each cell has genetic material from two different organisms, whereas an individual where some cells are derived from a different organism is called a chimera. Hybrids are not always intermediates between their parents, but can show hybrid vigor, sometimes growing larger or taller than either parent. The concept of a hybrid is interpreted differently in animal and plant breeding, where there is interest in the individual parentage. In genetics, attention is focused on the numbers of chromosomes. In taxonomy, a key question is how closely related the parent species are.
Broccoli is an edible green plant in the cabbage family whose large flowering head, stalk and small associated leaves are eaten as a vegetable. Broccoli is classified in the Italica cultivar group of the species Brassica oleracea. Broccoli has large flower heads, usually dark green, arranged in a tree-like structure branching out from a thick stalk which is usually light green. The mass of flower heads is surrounded by leaves. Broccoli resembles cauliflower, which is a different but closely related cultivar group of the same Brassica species.
The gene pool is the set of all genes, or genetic information, in any population, usually of a particular species.
Broccoflower is either of two edible plants of the species Brassica oleracea with light green heads. The edible portion is the immature flower head (inflorescence) of the plant.
Haldane's rule is an observation about the early stage of speciation, formulated in 1922 by the British evolutionary biologist J.B.S. Haldane, that states that if — in a species hybrid — only one sex is inviable or sterile, that sex is more likely to be the heterogametic sex. The heterogametic sex is the one with two different sex chromosomes; in therian mammals, for example, this is the male.
Self-incompatibility (SI) is a general name for several genetic mechanisms that prevent self-fertilization in sexually reproducing organisms, and thus encourage outcrossing and allogamy. It is contrasted with separation of sexes among individuals (dioecy), and their various modes of spatial (herkogamy) and temporally (dichogamy) separation.
Brassica oleracea is a plant species from family Brassicaceae that includes many common cultivars used as vegetables, such as cabbage, broccoli, cauliflower, kale, Brussels sprouts, collard greens, Savoy cabbage, kohlrabi, and gai lan.
The triangle of U is a theory about the evolution and relationships among the six most commonly known members of the plant genus Brassica. The theory states that the genomes of three ancestral diploid species of Brassica combined to create three common tetraploid vegetables and oilseed crop species. It has since been confirmed by studies of DNA and proteins.
Brassica rapa is a plant species growing in various widely cultivated forms including the turnip ; napa cabbage, bomdong, bok choy, and rapini.
Crocus sativus, commonly known as saffron crocus or autumn crocus, is a species of flowering plant in the iris family Iridaceae. A cormous autumn-flowering cultivated perennial, unknown in the wild, it is best known for the culinary use of its floral stigmas as the spice saffron. Human cultivation of saffron crocus and the trade and use of saffron have endured for more than 3,500 years and span different cultures, continents, and civilizations.
During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids, have developed under a combination of artificial and natural selection. This diversity has led to much confusion in the naming of wheats. This article explains how genetic and morphological characteristics of wheat influence its classification, and gives the most common botanical names of wheat in current use. Information on the cultivation and uses of wheat is at the main wheat page.
In biology, a deme, in the strict sense, is a group of individuals that belong to the same taxonomic group. However, when biologists, and especially ecologists, use the term ‘deme’ they usually refer to it as the definition of a gamodeme: a local group of individuals that interbreed with each other and share a gene pool. The latter definition of a deme is only applicable to sexual reproducing species, while the former is more neutral and also takes asexual reproducing species into account, such as certain plant species. In the following sections the latter definition of a deme will be used.
Georgii Dmitrievich Karpechenko was a Russian and Soviet biologist. His name has sometimes been transliterated as Karpetschenko.
Brassicoraphanus is any intergeneric hybrid between the genera Brassica and Raphanus (radish). The name comes from the combination of the genus names. Both diploid hybrids and allopolyploid hybrids are known and share this name.
The mechanisms of reproductive isolation are a collection of evolutionary mechanisms, behaviors and physiological processes critical for speciation. They prevent members of different species from producing offspring, or ensure that any offspring are sterile. These barriers maintain the integrity of a species by reducing gene flow between related species.
Hybrid speciation is a form of speciation where hybridization between two different species leads to a new species, reproductively isolated from the parent species. Previously, reproductive isolation between two species and their parents was thought to be particularly difficult to achieve, and thus hybrid species were thought to be very rare. With DNA analysis becoming more accessible in the 1990s, hybrid speciation has been shown to be a somewhat common phenomenon, particularly in plants. In botanical nomenclature, a hybrid species is also called a nothospecies. Hybrid species are by their nature polyphyletic.
Somatic fusion, also called protoplast fusion, is a type of genetic modification in plants by which two distinct species of plants are fused together to form a new hybrid plant with the characteristics of both, a somatic hybrid. Hybrids have been produced either between different varieties of the same species or between two different species.
Frank Harlan Lewis was an American botanist, geneticist, taxonomist, systematist, and evolutionist who worked primarily with plants in the genus Clarkia. He is best known for his theories of "catastrophic selection" and "saltational speciation", which are closely aligned with the concepts of quantum evolution and sympatric speciation. The concepts were first articulated in 1958 by Lewis and Peter H. Raven, and later refined in a 1962 paper by Lewis in which he coined the term "catastrophic selection". In 1966, he referred to the same mechanism as "saltational speciation".
This glossary of evolutionary biology is a list of definitions of terms and concepts used in the study of evolutionary biology, population biology, speciation, and phylogenetics, as well as sub-disciplines and related fields. For additional terms from related glossaries, see Glossary of genetics, Glossary of ecology, and Glossary of biology.