The omasum, also known as the bible, [1] the fardel, [1] the manyplies [1] and the psalterium, [1] is the third compartment of the stomach in ruminants. The omasum comes after the rumen and reticulum and before the abomasum. Different ruminants have different omasum structures and function based on the food that they eat and how they developed through evolution. [2]
The omasum can be found on the right side of the cranial portion of the rumen. [3] The omasum receives food from the reticulum through the reticulo-omasal orifice [3] and provides food to the abomasum through the omaso-abomasal orifice. [4] The omasum is spherical [5] to crescent shape [6] and has multiple leaflets similar to that of a book [7] called omasal laminae. [4] The omasal laminae are made of thin muscular layers covered with a nonglandular mucous membrane. [4] The omasal laminae come from the sides of the large curvature and project towards the inside of the omasum, extending from the reticulo-omasal orifice to the omaso-abomasal orifice. [8] [4] The laminae greatly increase the surface area of the omasum. [3] [9] The laminae are covered in omasal papillae that are claw-like in some ruminants or blunt cones in others. [4] [2] These papillae further increase the surface area but they also provide increased friction against the food particles. [3]
The function of the omasum is not completely understood. [5] During the second contraction phase of the reticulum, the reticule-omasal sphincter opens for a few seconds allowing a small volume of finely dispersed and well-fermented ingesta to enter the omasum. [3]
The omasum has two physiological compartments: omasal canal that transfers food from the reticulum to the omasum, and the inter-laminate recesses between the mucosal laminae which provide the area for absorption. [2] The omasum is where food particles that are small enough get transferred into the abomasum for enzymatic digestion. [5] [2] In ruminants with a more sophisticated omasum, the large surface area [9] allows it to play a key role in the absorption of water, electrolytes, [2] [4] volatile fatty acids, minerals, and the fermentation of food. [5]
Young ruminants that are still drinking milk have an esophageal groove that allows milk to bypass the rumen and go straight from the esophagus to the omasum. [10]
An early version of the omasum is seen in early ruminants like duikers and muntjacs, where it is a little more than a strainer sieve which prevents un-chewed foods from entering the abomasum. [2]
The smallest omasum belongs to ruminants that consume high quality diets like the moose and roe deer, while the largest belongs to those who are un-selective grass and roughage eaters like cattle and sheep. [2]
The omasum is not only bigger in grass and roughage eaters but there is greater differentiation in the book-like structure; seen as an increase in the number of laminae. [2]
Artiodactyls are mammals belonging to the order Artiodactyla. Typically, they are ungulates which bear weight equally on two of their five toes: the third and fourth, often in the form of a hoof. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two is that many artiodactyls digest plant cellulose in one or more stomach chambers rather than in their intestine as perissodactyls do. The advent of molecular biology, along with new fossil discoveries, found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.
Digestion is the breakdown of large insoluble food compounds into small water-soluble components so that they can be absorbed into the blood plasma. In certain organisms, these smaller substances are absorbed through the small intestine into the blood stream. Digestion is a form of catabolism that is often divided into two processes based on how food is broken down: mechanical and chemical digestion. The term mechanical digestion refers to the physical breakdown of large pieces of food into smaller pieces which can subsequently be accessed by digestive enzymes. Mechanical digestion takes place in the mouth through mastication and in the small intestine through segmentation contractions. In chemical digestion, enzymes break down food into the small compounds that the body can use.
Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The process, which takes place in the front part of the digestive system and therefore is called foregut fermentation, typically requires the fermented ingesta to be regurgitated and chewed again. The process of rechewing the cud to further break down plant matter and stimulate digestion is called rumination. The word "ruminant" comes from the Latin ruminare, which means "to chew over again".
Paratuberculosis is a contagious, chronic and sometimes fatal infection that primarily affects the small intestine of ruminants. It is caused by the bacterium Mycobacterium avium subspecies paratuberculosis. Infections normally affect ruminants, but have also been seen in a variety of nonruminant species, including rabbits, foxes, and birds. Horses, dogs, and nonhuman primates have been infected experimentally. Paratuberculosis is found worldwide, with some states in Australia being the only areas proven to be free of the disease. At least in Canada, the signs of BJD usually start when cattle are four to seven years of age, and then usually only are diagnosed in one animal at a time. Cattle "with signs of Johne’s disease shed billions of bacteria through their manure and serve as a major source of infection for future calves."
Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the physical digestive process of rumination.
A monogastric organism has a simple single-chambered stomach. Examples of monogastric herbivores are horses and rabbits. Examples of monogastric omnivores include humans, pigs, hamsters and rats. Furthermore, there are monogastric carnivores such as cats. A monogastric organism is comparable to ruminant organisms, such as cattle, goats, or sheep. Herbivores with monogastric digestion can digest cellulose in their diets by way of symbiotic gut bacteria. However, their ability to extract energy from cellulose digestion is less efficient than in ruminants.
The rumen, also known as a paunch, is the largest stomach compartment in ruminants and the larger part of the reticulorumen, which is the first chamber in the alimentary canal of ruminant animals. The rumen's microbial favoring environment allows it to serve as the primary site for microbial fermentation of ingested feed. The smaller part of the reticulorumen is the reticulum, which is fully continuous with the rumen, but differs from it with regard to the texture of its lining.
The abomasum, also known as the maw, rennet-bag, or reed tripe, is the fourth and final stomach compartment in ruminants. It secretes rennet, which is used in cheese creation.
The reticulorumen represents the first chamber in the alimentary canal of ruminant animals. It is composed of the rumen and reticulum. The reticulum differs from the rumen with regard to the texture of its lining. The rumen wall is covered in small, finger-like projections called papillae, whereas the reticulum is lined with ridges that form a hexagonal honeycomb pattern. The ridges are approximately 0.1–0.2 mm wide and are raised 0.5 cm above the reticulum wall. The hexagons in the reticulum are approximately 2–5 cm wide in cattle. Despite the differences in the texture of the lining of the two parts of the reticulorumen, it represents one functional space.
The reticulum is the second chamber in the four-chamber alimentary canal of a ruminant animal. Anatomically it is the smaller portion of the reticulorumen along with the rumen. Together these two compartments make up 84% of the volume of the total stomach.
Abomasitis is a relatively rare ruminant disease characterized by inflammation of abomasum usually caused by larval development in young calves, lambs, and goat kids. It occurs with gastroenteritis, but can also be a side effect of other diseases. However, it is seldom diagnosed as a separate condition.
Grass tetany is a metabolic disease involving magnesium deficiency, which can occur in such ruminant livestock as beef cattle, dairy cattle and sheep, usually after grazing on pastures of rapidly growing grass, especially in early spring.
A cannulated cow or fistulated cow refers to a cow that has been surgically fitted with a cannula. A cannula acts as a porthole-like device that allows access to the rumen of a cow, to perform research and analysis of the digestive system and to allow veterinarians to transplant rumen contents from one cow to another.
Cattle are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus Bos. Mature female cattle are called cows and mature male cattle are bulls. Young female cattle (heifers), young male cattle, and castrated male cattle (steers) are all colloquially called "cows".
Paramphistomum cervi, the type species of Paramphistomum, is a parasitic flat worm belonging to the class Trematoda. It is a tiny fluke mostly parasitising livestock ruminants, as well as some wild mammals. Uniquely, unlike most parasites, the adult worms are relatively harmless, but it is the developing juveniles that cause serious disease called paramphistomiasis, especially in cattle and sheep. Its symptoms include profuse diarrhoea, anaemia, lethargy, and often result in death if untreated.
Paramphistomum is a genus of parasitic flatworms belonging to the digenetic trematodes. It includes flukes which are mostly parasitising livestock ruminants, as well as some wild mammals. They are responsible for the serious disease called paramphistomiasis, also known as amphistomosis, especially in cattle and sheep. Its symptoms include profuse diarrhoea, anaemia, lethargy, and often result in death if untreated. They are found throughout the world, and most abundantly in livestock farming regions such as Australia, Asia, Africa, Eastern Europe, and Russia.
Ruminal tympany, also known as ruminal bloat, is a disease of ruminant animals, characterized by an excessive volume of gas in the rumen. Ruminal tympany may be primary, known as frothy bloat, or secondary, known as free-gas bloat.
Ostertagia ostertagi, commonly known as the medium stomach worm or brown stomach worm, is a parasitic nematode of cattle. O. ostertagi can also be found to a lesser extent in sheep, goats, wild ruminants, and horses. It causes ostertagiosis, which is potentially fatal in cattle. It is found worldwide and is economically important to cattle industries, particularly those found in temperate climates.
Methanogens are a group of microorganisms that produce methane as a byproduct of their metabolism. They play an important role in the digestive system of ruminants. The digestive tract of ruminants contains four major parts: rumen, reticulum, omasum and abomasum. The food with saliva first passes to the rumen for breaking into smaller particles and then moves to the reticulum, where the food is broken into further smaller particles. Any indigestible particles are sent back to the rumen for rechewing. The majority of anaerobic microbes assisting the cellulose breakdown occupy the rumen and initiate the fermentation process. The animal absorbs the fatty acids, vitamins and nutrient content on passing the partially digested food from the rumen to the omasum. This decreases the pH level and initiates the release of enzymes for further breakdown of the food which later passes to the abomasum to absorb remaining nutrients before excretion. This process takes about 9–12 hours.
Displaced abomasum in cattle occurs when the abomasum, also known as the true stomach, which typically resides on the floor of the abdomen, fills with gas and rises to the top of the abdomen, where it is said to be ‘displaced’. When the abomasum moves from its normal position it prevents the natural passage of gas and feed through the digestive system, creating a restriction. As cattle are ruminants, which have a 4 chambered stomach composed of a rumen, reticulum, omasum and abomasum. Ruminants require this specialized digestive system in order to properly process and break down their high fiber and cellulose rich diets. As this type of digestive system is quite complex it is at a greater risk for incidence. Due to the natural anatomy of cattle it is more common to have the abomasum displace to the left, known as a left-displaced abomasum, than to the right, right-displaced abomasum. When the abomasum becomes displaced there also becomes a chance of an abomasal volvulus, twist, developing. An abomasal volvulus occurs when the abomasum, which is already out of place, will rotate and cut off blood and nutrient supply to the abomasum. Cattle which develop an abomasal twist require immediate vet attention to regain blood supply and food passage through the digestive system or the abomasum will begin to shut down due to lack of blood supply and toxicity development.
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