Root cap

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Root tip magnified 100x. 1. Meristem 2. Columellae (statocytes with statolithes) 3. Lateral part of the tip 4. Dead cells 5. Elongation zone Root-tip-tag.png
Root tip magnified 100×. 1. Meristem 2. Columellae (statocytes with statolithes) 3. Lateral part of the tip 4. Dead cells 5. Elongation zone

The root cap is a type of tissue at the tip of a plant root. [1] It is also called calyptra. Root caps contain statocytes which are involved in gravity perception in plants. [1] If the cap is carefully removed the root will grow randomly. The root cap protects the growing tip in plants. [1] It secretes mucilage to ease the movement of the root through soil, [1] and may also be involved in communication with the soil microbiota. [1]

The purpose of the root cap is to enable downward growth of the root, with the root cap covering the sensitive tissue in the root. [2] Thanks to the presence of statocytes, the root cap enables geoperception or gravitropism. This allows the plant to grow downwards (with gravity) or upwards (against gravity). [3]

The root cap is absent in some parasitic plants [4] :138 and some aquatic plants, in which a sac-like structure called the root pocket may form instead. [5] :2–76

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<span class="mw-page-title-main">Lateral root</span> Plant root

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<span class="mw-page-title-main">Statocyte</span>

Statocytes are gravity-sensing (gravitropic) cells in higher plants. They contain amyloplasts-statoliths – starch-filled amyloplastic organelles – which sediment at the lowest part of the cells. In the roots, sedimentation of the statoliths towards the lower part of the statocytes constitutes a signal for the production and redistribution of auxin. When stems or roots are not exactly aligned with the gravity vector, statoliths move and adjust to gravity. This is followed by a triggering of the asymmetrical distribution of auxin that causes the curvature and growth of stems against the gravity vector, as well as growth of roots along the gravity vector. Statocytes are present in the elongating region of coleoptiles, shoots and inflorescence stems. In roots, the root cap is the only place where sedimentation is observed, and only the central columella cells of the root cap serve as gravity-sensing statocytes. They can initiate differential growth patterns, bending the root towards the vertical axis.

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The exodermis is a physiological barrier that has a role in root function and protection. The exodermis is a membrane of variable permeability responsible for the radial flow of water, ions, and nutrients. It is the outer layer of a plant's cortex. The exodermis serves a double function as it can protect the root from invasion by foreign pathogens and ensures that the plant does not lose too much water through diffusion through the root system and can properly replenish its stores at an appropriate rate.

References

  1. 1 2 3 4 5 Raven, J.A.; Edwards, D. (2001). "Roots: evolutionary origins and biogeochemical significance". Journal of Experimental Botany. 52 (90001): 381–401. doi: 10.1093/jexbot/52.suppl_1.381 . PMID   11326045.
  2. Burgess, Jeremy (1985-05-16). Introduction to Plant Cell Development. CUP Archive. ISBN   9780521316118.
  3. Kuya, Noriyuki; Sato, Seiichi (2011). "The relationship between profiles of plagiogravitropism and morphometry of columella cells during the development of lateral roots of Vigna angularis". Advances in Space Research. 47 (3): 553–562. Bibcode:2011AdSpR..47..553K. doi:10.1016/j.asr.2010.09.009.
  4. Jeffrey, Edward Charles (2007). The Anatomy of Woody Plants. Pomeroy, Ohio: Carpenter Press. ISBN   978-1-4067-1634-4.
  5. Gupta, P.K. (2007). Genetics: Classical to Modern. Rastogi Publications. ISBN   978-8-1713-3896-2.