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Salt pruning is the process by which saline mists generated by seawater are driven ashore by winds and thus over time alter the shape of trees or shrubs. The process degrades foliage and branches on the windward side of the plant that faces the body of saline water, more than it does the foliage on the landward side. The resultant growth form is asymmetrical, appearing "swept back" away from the ocean.
There are numerous examples worldwide of this phenomenon, though it usually appears on coasts near saline water. In the eastern United States on Long Island, occurrences of salt-pruned Quercus stellata are observable in Flax Marsh. [1] In San Diego County, California, a colony of Pinus torreyana has been salt-pruned by spray from the Pacific Ocean. [2]
The logo of the Torrey Pines Golf Course in La Jolla, California, features a salt-pruned Torrey pine.
In the case of Fire Island's Sunken Forest, the saline mists prevent new oak shoots from growing above the impacted area. [3] These oaks grow "pruned" by the wind with bundles of bushes and stems at the top to keep it top-heavy. They also grow much shorter than regular oaks due to the salt keeping them below or around the height of the hill.
The first known record of salt pruning was in 1805, though it was called a "storm of salt" by British botanist Richard Anthony Salisbury. [4] Much later, Bertram Whittier Wells discovered a similar effect at Cape Fear, though he described the oak tree's bent form to be the "Salt Spray Climax." [5]
Later research done in Cape Fear by Stephen G. Boyce between the years 1951-52 found the oaks were not experiencing necrosis due to the abrasion from the salt content and pressure, but instead had grown accustomed to it. The plant life in the Cape Cod area showed similar salt pruning. While the leaves did not experience necrosis, they did appear to curl in on themselves. [6]
A shrub or bush is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple stems and shorter height, less than 6–10 m (20–33 ft) tall. Small shrubs, less than 2 m (6.6 ft) tall are sometimes termed as subshrubs. Many botanical groups have species that are shrubs, and others that are trees and herbaceous plants instead.
Pinus nigra, the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as well as Crimea and in the high mountains of Northwest Africa.
The Torrey pine is a rare pine species in California, United States. It is a critically endangered species growing only in coastal San Diego County, and on Santa Rosa Island, offshore from Santa Barbara in Santa Barbara County. The Torrey pine is endemic to the California coastal sage and chaparral ecoregion.
Pruning is a horticultural, arboricultural, and silvicultural practice involving the selective removal of certain parts of a plant, such as branches, buds, or roots.
Pinus rigida, the pitch pine, is a small-to-medium-sized pine. It is native to eastern North America, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuitable for growth, such as acidic, sandy, and low-nutrient soils.
Pinus elliottii, commonly known as slash pine, is a conifer tree native to the Southeastern United States. Slash pine is named after the "slashes" – swampy ground overgrown with trees and bushes – that constitute its habitat. Other common names include swamp pine, yellow slash pine, and southern Florida pine. Slash pine has two different varieties: P. e. var. elliottii and P. e. var. densa. Historically, slash pine has been an important economic timber for naval stores, turpentine, and resin. The wood of slash pine is known for its unusually high strength, especially for a pine. It exceeds many hardwoods and is even comparable to very dense woods such as ironwood.
Table Mountain pine, Pinus pungens, also called hickory pine, prickly pine, or mountain pine, is a small pine native to the Appalachian Mountains in the United States.
Jack pine, also known as grey pine or scrub pine, is a North American pine.
The stone pine, botanical name Pinus pinea, also known as the Italian stone pine, Mediterranean stone pine, umbrella pine and parasol pine, is a tree from the pine family (Pinaceae). The tree is native to the Mediterranean region, occurring in Southern Europe and the Levant. The species was introduced into North Africa millennia ago, and is also naturalized in the Canary Islands, South Africa and New South Wales.
Torrey Pines State Natural Reserve is 2,000 acres (810 ha) of coastal state park located in San Diego, California, off North Torrey Pines Road. Although it is located within San Diego city limits, it remains one of the wildest stretches of land (8 km²) on the Southern California coast. It is bordered immediately on the south by Torrey Pines Municipal Golf Course and on the north by the city of Del Mar. The reserve was designated a National Natural Landmark in 1977.
Torrey Pines Golf Course is a 36-hole municipal golf facility on the west coast of the United States, owned by the city of San Diego, California. It sits on the coastal cliffs overlooking the Pacific Ocean in the community of La Jolla, just south of Torrey Pines State Reserve. Opened in 1957, it was built on the site of Camp Callan, a U.S. Army installation during World War II.
Pinus halepensis, commonly known as the Aleppo pine, also known as the Jerusalem pine, is a pine native to the Mediterranean region.
The Atlantic coastal pine barrens is a now rare temperate coniferous forest ecoregion of the Northeast United States distinguished by unique species and topographical features, generally nutrient-poor, often acidic soils and a pine tree distribution once controlled by frequent fires.
Cronartium ribicola is a species of rust fungus in the family Cronartiaceae that causes the disease white pine blister rust. Other names include: Rouille vésiculeuse du pin blanc (French), white pine Blasenrost (German), moho ampolla del pino blanco (Spanish).
The plant pathogenic fungus Leucostoma kunzei is the causal agent of Leucostoma canker, a disease of spruce trees found in the Northern Hemisphere, predominantly on Norway spruce and Colorado blue spruce. This disease is one of the most common and detrimental stem diseases of Picea species in the northeastern United States, yet it also affects other coniferous species. Rarely does it kill its host tree; however, the disease does disfigure by killing host branches and causing resin exudation from perennial lesions on branches or trunks.
The California coastal sage and chaparral is a Mediterranean forests, woodlands, and scrub ecoregion located in southwestern California and northwestern Baja California (Mexico). It is part of the larger California chaparral and woodlands ecoregion.
The Sierra Madre del Sur pine–oak forests is a subtropical coniferous forest ecoregion in the Sierra Madre del Sur mountain range of southern Mexico.
The Southeastern conifer forests are a temperate coniferous forest ecoregion of the southeastern United States. It is the largest conifer forest ecoregion east of the Mississippi River. It is also the southernmost instance of temperate coniferous forest within the Nearctic realm.
Pine-pine gall rust, also known as western gall rust, is a fungal disease of pine trees. It is caused by Endocronartium harknessii, an autoecious, endocyclic, rust fungus that grows in the vascular cambium of the host. The disease is found on pine trees with two or three needles, such as ponderosa pine, jack pine and scots pine. It is very similar to pine-oak gall rust, but its second host is another Pinus species. The fungal infection results in gall formation on branches or trunks of infected hosts. Gall formation is typically not detrimental to old trees, but has been known to kill younger, less stable saplings. Galls can vary from small growths on branch extremities to grapefruit-sized galls on trunks.
The wildlife of the Channel Islands of California is wide and diverse, including many endemic species. While the land wildlife is slightly limited, there being only one large, naturally predatory, and native mammal, the small island fox, marine life can include anything from kelp forests to great white sharks.
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