Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word xylem is derived from the Ancient Greek word ξύλον (xylon), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858.
Vascular plants, also called tracheophytes or collectively Tracheophyta, form a large group of land plants that have lignified tissues for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue to conduct products of photosynthesis. Vascular plants include the clubmosses, horsetails, ferns, gymnosperms, and angiosperms. Scientific names for the group include Tracheophyta, Tracheobionta and Equisetopsida sensu lato. Some early land plants had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones.
In botany, a stoma, also called a stomate, is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bordered by a pair of specialized parenchyma cells known as guard cells that regulate the size of the stomatal opening.
Root pressure is the transverse osmotic pressure within the cells of a root system that causes sap to rise through a plant stem to the leaves.
Guttation is the exudation of drops of xylem sap on the tips or edges of leaves of some vascular plants, such as grasses, and a number of fungi, which are not plants but were previously categorized as such and studied as part of botany.
The apoplast is the extracellular space outside of plant cell membranes, especially the fluid-filled continuum of cell walls of adjacent cells where material can diffuse freely. Fluid and material flows occurring in any extracellular space are called apoplastic flow or apoplastic transport. The apoplastic pathway is one route by which water and solutes are transported and distributed to different places through tissues and organs, contrasting with the symplastic pathway.
Water potential is the potential energy of water per unit volume relative to pure water in reference conditions. Water potential quantifies the tendency of water to move from one area to another due to osmosis, gravity, mechanical pressure and matrix effects such as capillary action. The concept of water potential has proved useful in understanding and computing water movement within plants, animals, and soil. Water potential is typically expressed in potential energy per unit volume and very often is represented by the Greek letter ψ.
Soil moisture is the water content of the soil. It can be expressed in terms of volume or weight. Soil moisture measurement can be based on in situ probes or remote sensing methods.
Moisture stress is a form of abiotic stress that occurs when the moisture of plant tissues is reduced to suboptimal levels. Water stress occurs in response to atmospheric and soil water availability when the transpiration rate exceeds the rate of water uptake by the roots and cells lose turgor pressure. Moisture stress is described by two main metrics, water potential and water content.
Turgor pressure is the force within the cell that pushes the plasma membrane against the cell wall.
The soil-plant-atmosphere continuum (SPAC) is the pathway for water moving from soil through plants to the atmosphere. Continuum in the description highlights the continuous nature of water connection through the pathway. The low water potential of the atmosphere, and relatively higher water potential inside leaves, leads to a diffusion gradient across the stomatal pores of leaves, drawing water out of the leaves as vapour. As water vapour transpires out of the leaf, further water molecules evaporate off the surface of mesophyll cells to replace the lost molecules since water in the air inside leaves is maintained at saturation vapour pressure. Water lost at the surface of cells is replaced by water from the xylem, which due to the cohesion-tension properties of water in the xylem of plants pulls additional water molecules through the xylem from the roots toward the leaf.
The pressure flow hypothesis, also known as the mass flow hypothesis, is the best-supported theory to explain the movement of sap through the phloem. It was proposed by Ernst Münch, a German plant physiologist in 1930. A high concentration of organic substances, particularly sugar, inside cells of the phloem at a source, such as a leaf, creates a diffusion gradient that draws water into the cells from the adjacent xylem. This creates turgor pressure, also known as hydrostatic pressure, in the phloem. Movement of phloem sap occurs by bulk flow from sugar sources to sugar sinks. The movement in phloem is bidirectional, whereas, in xylem cells, it is unidirectional (upward). Because of this multi-directional flow, coupled with the fact that sap cannot move with ease between adjacent sieve-tubes, it is not unusual for sap in adjacent sieve-tubes to be flowing in opposite directions.
Osmoregulation is the active regulation of the osmotic pressure of an organism's body fluids, detected by osmoreceptors, to maintain the homeostasis of the organism's water content; that is, it maintains the fluid balance and the concentration of electrolytes to keep the body fluids from becoming too diluted or concentrated. Osmotic pressure is a measure of the tendency of water to move into one solution from another by osmosis. The higher the osmotic pressure of a solution, the more water tends to move into it. Pressure must be exerted on the hypertonic side of a selectively permeable membrane to prevent diffusion of water by osmosis from the side containing pure water.
A xerophyte is a species of plant that has adaptations to survive in an environment with little liquid water. Examples are typically desert regions like the Sahara, and places in the Alps or the Arctic. Popular examples of xerophytes are cacti, pineapple and some Gymnosperm plants.
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm or culms.
Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. Water is necessary for plants but only a small amount of water taken up by the roots is used for growth and metabolism. The remaining 97–99.5% is lost by transpiration and guttation. Leaf surfaces are dotted with pores called stomata, and in most plants they are more numerous on the undersides of the foliage. The stomata are bordered by guard cells and their stomatal accessory cells that open and close the pore. Transpiration occurs through the stomatal apertures, and can be thought of as a necessary "cost" associated with the opening of the stomata to allow the diffusion of carbon dioxide gas from the air for photosynthesis. Transpiration also cools plants, changes osmotic pressure of cells, and enables mass flow of mineral nutrients and water from roots to shoots. Two major factors influence the rate of water flow from the soil to the roots: the hydraulic conductivity of the soil and the magnitude of the pressure gradient through the soil. Both of these factors influence the rate of bulk flow of water moving from the roots to the stomatal pores in the leaves via the xylem.
A pressure bomb, pressure chamber, or Scholander bomb is an instrument that can measure the approximate water potential of plant tissues. A leaf and petiole or stem segment is placed inside a sealed chamber. Pressurized gas is slowly added to the chamber. As the pressure increases, at some point the liquid contents of the sample will be forced out of the xylem and will be visible at the cut end of the stem or petiole. The pressure that is required to do so is equal and opposite to the water potential of the sample. Pressure bombs are field portable and mechanically simple, which make them the predominant method for water potential measurements in the fields of plant physiology and ecophysiology.
In higher plants water and minerals are absorbed through root hairs which are in contact with soil water and from the root hairs zone a little the root tips.
Stomatal conductance, usually measured in mmol m−2 s−1 by a porometer, estimates the rate of gas exchange and transpiration through the leaf stomata as determined by the degree of stomatal aperture.
Homoiohydry is the capacity of plants to regulate, or achieve homeostasis of, cell and tissue water content. Homoiohydry evolved in land plants to a lesser or greater degree during their transition to land more than 500 million years ago, and is most highly developed in the vascular plants. It is the consequence of a suite of morphological innovations and strategies that enable plant shoots exploring aerial environments to conserve water by internalising the gas exchange surfaces, enclosing them in a waterproof membrane and providing a variable aperture control mechanism, the stomatal guard cells, which regulate the rates of water transpiration and CO2 exchange. In vascular plants, water is acquired from the soil by roots and transported via the xylem to aerial portions of the plant. Water evaporation from the aerial surfaces of the plant is controlled by a waterproof covering of cuticle. Gas exchange with the atmosphere is controlled by stomata, which can open and close to control water loss, and diffusion of carbon dioxide to the chloroplasts takes place in intercellular spaces between chlorenchyma cells in the stem or in the mesophyll tissue of the leaf.
