Venom glands are a wide variety of salivary glands in insects and reptiles modified to produce venom and store it without harm to the organism itself. [1] [2]
In reptiles the glands are characterized by their ability to synthesize venom, store venom for long periods of time, stabilize the venom while in the gland itself, and rapidly activate the toxin upon deployment from the gland. [1] In insects however, specifically some parasitoid insects in the genus Hymenoptera, the venom gland has been shown to have additional functions such as the regulation of the immune response, paralysis, castration, and developmental alteration of the parasite's host. [2]
Lizard is the common name used for all squamate reptiles other than snakes, encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The grouping is paraphyletic as some lizards are more closely related to snakes than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon.
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved venom apparatus, such as fangs or a stinger, in a process called envenomation. Venom is often distinguished from poison, which is a toxin that is passively delivered by being ingested, inhaled, or absorbed through the skin, and toxungen, which is actively transferred to the external surface of another animal via a physical delivery mechanism.
Vipers are snakes in the family Viperidae, found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, New Zealand, Ireland, and various other isolated islands. They are venomous and have long, hinged fangs that permit deep penetration and injection of their venom. Three subfamilies are currently recognized. They are also known as viperids. The name "viper" is derived from the Latin word vipera, -ae, also meaning viper, possibly from vivus ("living") and parere, referring to the trait viviparity common in vipers like most of the species of Boidae.
The Atractaspididae (atractaspidids) are a family of venomous snakes found in Africa and the Middle East, commonly called mole vipers, stiletto snakes, or burrowing asps. Currently, 12 genera are recognized.
Snake venom is a highly toxic saliva containing zootoxins that facilitates in the immobilization and digestion of prey. This also provides defense against threats. Snake venom is usually injected by unique fangs during a bite, though some species are also able to spit venom.
Envenomation is the process by which venom is injected by the bite or sting of a venomous animal.
The Mexican beaded lizard is a species of lizard in the family Helodermatidae, one of the two species of venomous beaded lizards found principally in Mexico and southern Guatemala. It and the other members of the same genus, including the Gila monster, are the only lizards known to have evolved an overt venom delivery system. The Mexican beaded lizard is larger than the Gila monster, with duller coloration, black with yellowish bands. As it is a specialized predator that feeds primarily upon eggs, the primary use of its venom is still a source of debate among scientists. This venom has been found to contain several enzymes useful for manufacturing drugs in the treatment of diabetes, and research on the pharmacological use of its venom is ongoing.
Venomous mammals are tetrapods of the class Mammalia that produce venom, which they use to kill or disable prey, to defend themselves from predators or conspecifics or in agonistic encounters. Mammalian venoms form a heterogeneous group with different compositions and modes of action, from four orders of mammals: Eulipotyphla, Monotremata, Primates, and Chiroptera. To explain the rarity of venom delivery in Mammalia, Mark Dufton of the University of Strathclyde has suggested that modern mammalian predators do not need venom because they are able to kill quickly with their teeth or claws, whereas venom, no matter how sophisticated, requires time to disable prey.
Metapleural glands are secretory glands that were considered unique to ants and basal in the evolutionary history of ants. They are responsible for the production of an antibiotic fluid that then collects in a reservoir on the posterior of the ant's alitrunk. These reservoirs are also referred to as the bulla and vary in size between ant species and also between castes of the same species. As of 2023, research has found a metapleural gland convergently evolved in the wasp species Pelecinus polyturator.
Toxicofera is a proposed clade of scaled reptiles (squamates) that includes the Serpentes (snakes), Anguimorpha and Iguania. Toxicofera contains about 4,600 species, of extant Squamata. It encompasses all venomous reptile species, as well as numerous related non-venomous species. There is little morphological evidence to support this grouping; however, it has been recovered by all molecular analyses as of 2012.
Venomous snakes are species of the suborder Serpentes that are capable of producing venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The venom is typically delivered by injection using hollow or grooved fangs, although some venomous snakes lack well-developed fangs. Common venomous snakes include the families Elapidae, Viperidae, Atractaspididae, and some of the Colubridae. The toxicity of venom is mainly indicated by murine LD50, while multiple factors are considered to judge the potential danger to humans. Other important factors for risk assessment include the likelihood that a snake will bite, the quantity of venom delivered with the bite, the efficiency of the delivery mechanism, and the location of a bite on the body of the victim. Snake venom may have both neurotoxic and hemotoxic properties. There are about 600 venomous snake species in the world.
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders.
Euchambersia is an extinct genus of therocephalian therapsids that lived during the Late Permian in what is now South Africa and China. The genus contains two species. The type species E. mirabilis was named by paleontologist Robert Broom in 1931 from a skull missing the lower jaw. A second skull, belonging to a probably immature individual, was later described. In 2022, a second species, E. liuyudongi, was named by Jun Liu and Fernando Abdala from a well-preserved skull. It is a member of the family Akidnognathidae, which historically has also been referred by as the synonymous Euchambersiidae.
The Gila monster is a species of venomous lizard native to the Southwestern United States and the northwestern Mexican state of Sonora. It is a heavy, slow-moving reptile, up to 56 centimetres (22 in) long, and it is the only venomous lizard native to the United States. Its venomous close relatives, the four beaded lizards inhabit Mexico and Guatemala. The Gila monster is sluggish in nature, so it is not generally dangerous and very rarely poses a real threat to humans. However, it has a fearsome reputation and is sometimes killed despite the species being protected by state law in Arizona.
Venomoid surgery is a surgical procedure performed on venomous snakes to prevent them from producing venom. The procedure has been used for snakes kept as pets or used in public demonstrations in order to remove the risk of injury or death when handled, but is considered illegal and unethical. Most venomoid procedures consist of either removing the venom gland itself, or severing the duct between the gland and the fang. Removal of fangs is uncommon, as snakes frequently regenerate teeth, and the more invasive procedure of removing the underlying maxillary bone would be fatal. However, the duct and gland have been known to regenerate, and supposedly "safe" snakes have killed mice and successfully envenomated humans.
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.
Uatchitodon is an extinct genus of Late Triassic reptile known only from isolated teeth. Based on the structure of the teeth, Uatchitodon was probably a carnivorous archosauromorph. Folded grooves on the teeth indicate that the animal was likely venomous, with the grooves being channels for salivary venom. The teeth are similar to those of living venomous squamates such as Heloderma and venomous snakes. Uatchitodon is the earliest known venomous reptile.
Chemical defense is a strategy employed by many organisms to avoid consumption by producing toxic or repellent metabolites or chemical warnings which incite defensive behavioral changes. The production of defensive chemicals occurs in plants, fungi, and bacteria, as well as invertebrate and vertebrate animals. The class of chemicals produced by organisms that are considered defensive may be considered in a strict sense to only apply to those aiding an organism in escaping herbivory or predation. However, the distinction between types of chemical interaction is subjective and defensive chemicals may also be considered to protect against reduced fitness by pests, parasites, and competitors. Repellent rather than toxic metabolites are allomones, a sub category signaling metabolites known as semiochemicals. Many chemicals used for defensive purposes are secondary metabolites derived from primary metabolites which serve a physiological purpose in the organism. Secondary metabolites produced by plants are consumed and sequestered by a variety of arthropods and, in turn, toxins found in some amphibians, snakes, and even birds can be traced back to arthropod prey. There are a variety of special cases for considering mammalian antipredatory adaptations as chemical defenses as well.
Asobara is a genus of parasitoid wasps in the family Braconidae. It contains around forty species. The genus is best known for the Drosophila parasitoid Asobara tabida, which is notable as both a model for parasitoid wasp infection in insects, and also as a representative of the hologenome theory of evolution. Asobara tabida is commensally infected with Wolbachia, and cannot reproduce in the absence of Wolbachia infection. As such, the genome of Asobara is directly tied to the genome of its commensal Wolbachia symbiont, and the two are considered to have a hologenome.
Toxungen comprises a secretion or other body fluid of one or more biological toxins that is transferred by one animal to the external surface of another animal via a physical delivery mechanism. Toxungens can be delivered through spitting, spraying, or smearing. As one of three categories of biological toxins, toxungens can be distinguished from poisons, which are passively transferred via ingestion, inhalation, or absorption across the skin, and venoms, which are delivered through a wound generated by a bite, sting, or other such action. Toxungen use offers the evolutionary advantage of delivering toxins into the target's tissues without the need for physical contact.