Betula nigra

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Betula nigra
RiverBirchBark01.JPG
The bark of a young river birch
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Fagales
Family: Betulaceae
Genus: Betula
Subgenus: Betula subg. Neurobetula
Species:
B. nigra
Binomial name
Betula nigra
L.
Betula nigra range map 1.png
Natural range of Betula nigra

Betula nigra, the black birch, river birch or water birch, is a species of birch native to the Eastern United States from New Hampshire west to southern Minnesota, and south to northern Florida and west to Texas. It is one of the few heat-tolerant birches in a family of mostly cold-weather trees which do not thrive in USDA Zone 6 and up. B. nigra commonly occurs in floodplains and swamps. [2]

Contents

Description

Betula nigra is a deciduous tree growing to 25–30 meters (80–100 ft) with a trunk 50 to 150 centimeters (20 to 60 in) in diameter. The base of the tree is often divided into multiple slender trunks. [2] [3]

Bark

Bark characteristics of the river birch differ during its youth stage, maturation, and old growth. The bark of a young river birch can vary from having a salmon-pink to brown-gray tint and can be described as having loose layers of curling, paper thin scales. As the tree matures, the salmon-pink color is exchanged for a reddish-brown with a dark grey base color. The scales on a mature tree lack the loose curling and are closely pressed into thick, irregular plates. These scales are slightly separated from the trunk and can shift outward to the side. Once the river birch ages past maturity, the scales become thicker towards the base of the trunk and are divided in deep furrows. [3]

Leaves and fruit

The twigs are glabrous or thinly hairy. There is an absence of terminal buds, and lateral buds often have a hook at the tip of the bud, which differs from other species in the family Betulaceae. [4] The leaves are alternate, ovate, 4–8 centimeters (1+123+14 in) long and 3–6 centimeters (1+142+14 in) broad, with a serrated margin and five to twelve pairs of veins. The upper surface of the leaf is dark green in color, while the underside can be described as having a light yellow-green color. [5] The leaves turn yellow in Autumn. The flowers are wind-pollinated catkins 3–6 centimeters (1+142+14 in) long, the male catkins pendulous, the female catkins erect. The fruit is unusual among birches in maturing in late spring; it is composed of numerous tiny winged seeds packed between the catkin bracts. [2] [6]

Taxonomy

Betula nigra is a tree that falls into the family of Betulaceae, which is commonly known as the Birch or Alder Family. This family comprises six genera (Alnus, Betula, Carpinus, Corylus, Ostrya, and Ostryopsis) and includes alders, birches, hornbeams, and hazelnuts. Species within this family, along with Betula nigra, are shrubs or trees that grow along stream sides or in poorly drained soils throughout the Northern Hemisphere. Betulaceae is included within the order Fagales, which branches from the Rosid clade. [7] [8]

Habitat and range

The river birch is often found in low-elevation regions from as north as Massachusetts to as south as northern Florida. It can be found extending west to Kansas and east to the coast where proper habitat conditions occur. As its name depicts, this birch is found along stream-sides. It can also be a prominent species found in forested wetland communities and in areas containing moist soil, such as floodplains. [4] States include: Alabama, Arkansas, Connecticut, Delaware, Florida, Georgia, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maryland, Massachusetts, Mississippi, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Oklahoma, Pennsylvania, South Carolina, Tennessee, Texas, Virginia, West Virginia, and Wisconsin. [9]

River birch is best placed in USDA hardiness zones 4–9.[ citation needed ]

Conservation status in the United States

It is listed as threatened in New Hampshire. [10]

Ecology

In states in which mining is prevalent, the river birch is often used for reclamation and erosion control, as it is well suited for soils that are too acidic for other species of hardwoods. In West Virginia, they have been found to establish within mine refuse sites after being blown from neighboring areas. [9]

As the species occurs predominately in flood plains and along stream banks, it is described as being moderately tolerant of flooding. Saplings were observed to survive up to 30 days of continuous flooding in some regions. While the species is tolerant of excessive water, it is intolerant of shade. Seeds will not germinate without a large amount of direct sunlight. [9]

This species is utilized by many local bird species, such as waterfowl, ruffed grouse, and wild turkey. Many waterfowl use the cover for nesting sites, while the ruffed grouse and wild turkey use the seeds as a food source. [9] Deer have been known to graze on saplings or reachable branches. [4] It is a larval host for over fifteen moth species, including Acronicta betulae , Acrobasis betulivorella , Bucculatrix coronatella , Nemoria bistriaria , Nites betulella , Orgyia leucostigma , and Pseudotelphusa betulella . [11]

Germination

Seeds are typically produced annually. Seasonal development begins in the fall as male catkins begin to form and mature. The emergence of female catkins corresponds with the return of leaves around early spring. Male and female fruit matures during the spring season or in the early summer months. [9]

Once mature, the seeds are predominantly spread by wind or water from neighboring stream channels. Seeds spread by water are generally more successful as the moist banks of stream channels, where the seeds are deposited, are favorable for germination and sturdy establishment. Successful germination often occurs in large numbers along sandbars, where alluvial soil is present. [9]

Cultivation and uses

While its native habitat is wet ground, it will grow on higher land, and its bark is quite distinctive, making it a favored ornamental tree for landscape use. A number of cultivars with much whiter bark than the normal wild type have been selected for garden planting, including 'Heritage' and 'Dura Heat'; these are notable as the only white-barked birches resistant to the bronze birch borer ( Agrilus anxius ) in warm areas of the southeastern United States of America. [12]

Native Americans used the boiled sap as a sweetener similar to maple syrup, and the inner bark as a survival food. [12] The river birch is not typically used in the commercial lumber industry, due to knotting, but its strong, closely grained wood is sometimes used for local furniture, woodenware, and fuel. [5] [9]

Trunk of a river birch River Birch Trunk.JPG
Trunk of a river birch
Middle of the tree River Birch Middle.JPG
Middle of the tree
River birch leaves & seeds River Birch Seeds.JPG
River birch leaves & seeds

Essential oils

The essential oils derived from leaves, inner bark, and buds of B. nigra are mostly composed of eugenol, linalool, palmitic acid, and heptacosane with many more compounds in smaller concentrations. [13] The combined essential oils are phytotoxic to lettuce (Lactuca sativa) and perennial ryegrass (Lolium perenne) seedlings. They have also demonstrated insecticidal, nematicidal, and antibacterial properties. [13]

Related Research Articles

<span class="mw-page-title-main">Alder</span> Genus of flowering plants in the birch family Betulaceae

Alders are trees comprising the genus Alnus in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few species extending into Central America, as well as the northern and southern Andes.

<span class="mw-page-title-main">Birch</span> Genus of flowering plants in the family Betulaceae

A birch is a thin-leaved deciduous hardwood tree of the genus Betula, in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus Betula contains 30 to 60 known taxa of which 11 are on the IUCN 2011 Red List of Threatened Species. They are a typically rather short-lived pioneer species widespread in the Northern Hemisphere, particularly in northern areas of temperate climates and in boreal climates. Birch wood, the wood of the birch, is used for a wide range of purposes.

<span class="mw-page-title-main">Hornbeam</span> Genus of flowering plants

Hornbeams are hardwood trees in the plant genus Carpinus in the family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere.

<i>Alnus glutinosa</i> Species of flowering plant in the birch family Betulaceae

Alnus glutinosa, the common alder, black alder, European alder, European black alder, or just alder, is a species of tree in the family Betulaceae, native to most of Europe, southwest Asia and northern Africa. It thrives in wet locations where its association with the bacterium Frankia alni enables it to grow in poor quality soils. It is a medium-sized, short-lived tree growing to a height of up to 30 metres (98 feet). It has short-stalked rounded leaves and separate male and female flowers in the form of catkins. The small, rounded fruits are cone-like and the seeds are dispersed by wind and water.

<i>Betula pendula</i> Species of birch

Betula pendula, commonly known as silver birch, warty birch, European white birch, or East Asian white birch, is a species of tree in the family Betulaceae, native to Europe and parts of Asia, though in southern Europe, it is only found at higher altitudes. Its range extends into Siberia, China, and southwest Asia in the mountains of northern Turkey, the Caucasus, and northern Iran. It has been introduced into North America, where it is known as the European white birch or weeping birch and is considered invasive in some states in the United States and parts of Canada. The tree can also be found in more temperate regions of Australia.

<i>Juglans cinerea</i> Species of tree

Juglans cinerea, commonly known as butternut or white walnut, is a species of walnut native to the eastern United States and southeast Canada.

<i>Betula papyrifera</i> Species of tree

Betula papyrifera is a short-lived species of birch native to northern North America. Paper birch is named after the tree's thin white bark, which often peels in paper-like layers from the trunk. Paper birch is often one of the first species to colonize a burned area within the northern latitudes, and is an important species for moose browsing. Primary commercial uses for paper birch wood are as boltwood and sawlogs, while secondary products include firewood and pulpwood. It is the provincial tree of Saskatchewan and the state tree of New Hampshire.

<i>Betula alleghaniensis</i> Species of flowering plant in the birch family Betulaceae

Betula alleghaniensis, the yellow birch, golden birch, or swamp birch, is a large tree and an important lumber species of birch native to northeastern North America. Its vernacular names refer to the golden color of the tree's bark. In the past its scientific name was Betula lutea.

<i>Betula lenta</i> Species of plant

Betula lenta is a species of birch native to eastern North America, from southern Maine west to southernmost Ontario, and south in the Appalachian Mountains to northern Georgia.

<i>Carpinus caroliniana</i> Species of tree

Carpinus caroliniana, the American hornbeam, is a small hardwood understory tree in the genus Carpinus. American hornbeam is also known as blue-beech, ironwood, musclewood and muscle beech. It is native to eastern North America, from Minnesota and southern Ontario east to Maine, and south to eastern Texas and northern Florida. It also grows in Canada. It occurs naturally in shaded areas with moist soil, particularly near the banks of streams or rivers, and is often a natural constituent understory species of the riverine and maritime forests of eastern temperate North America.

<i>Betula populifolia</i> Species of birch

Betula populifolia, known as the gray birch, is a deciduous tree in the family Betulaceae. It is native to eastern North America and is most commonly found in the northeast United States as well as southern Quebec, New Brunswick, and Nova Scotia. The tree is a pioneer species that is commonly found in sites following disturbance, such as fire or logging. Gray birches don't have as much economic value as other birch species but are still commonly used as ornamental trees.

<i>Salix nigra</i> Species of tree

Salix nigra, the black willow, is a species of willow native to eastern North America, from New Brunswick and southern Ontario west to Minnesota, and south to northern Florida and Texas.

<i>Populus deltoides</i> Species of tree

Populus deltoides, the eastern cottonwood or necklace poplar, is a species of cottonwood poplar native to North America, growing throughout the eastern, central, and southwestern United States as well as the southern Canadian prairies, the southernmost part of eastern Canada, and northeastern Mexico.

<i>Carya glabra</i> Species of tree

Carya glabra, the pignut hickory, is a common, but not abundant species of hickory in the oak-hickory forest association in the Eastern United States and Canada. Other common names are pignut, sweet pignut, coast pignut hickory, smoothbark hickory, swamp hickory, and broom hickory. The pear-shaped nut ripens in September and October, has a sweet maple like smell, and is an important part of the diet of many wild animals. The wood is used for a variety of products, including fuel for home heating. Its leaves turn yellow in the Fall.

<i>Betula occidentalis</i> Species of birch

Betula occidentalis, the water birch or red birch, is a species of birch native to western North America, in Canada from Yukon east to Northwestern Ontario and southwards, and in the United States from eastern Washington east to western North Dakota, and south to eastern California, northern Arizona and northern New Mexico, and southwestern Alaska. It typically occurs along streams in mountainous regions, sometimes at elevations of 2,100 metres and in drier areas than paper birch.

<i>Betula cordifolia</i> Species of birch

Betula cordifolia, the mountain paper birch or heartleaf birch is a birch species native to Eastern Canada and the Northeastern United States. Until recently it was considered a variety of Betula papyrifera, with which it shares many characteristics, and it was classified as B. papyrifera var. cordifolia (Regel) Fern.

<i>Alnus serrulata</i> Species of tree

Alnus serrulata, the hazel alder or smooth alder, is a thicket-forming shrub in the family Betulaceae. It is native to eastern North America and can be found from western Nova Scotia and southern New Brunswick south to Florida and Texas.

<i>Salix arbusculoides</i> Species of flowering plant

Salix arbusculoides is a species of flowering plant in the willow family known by the common name little tree willow. It is native to northern North America, where its distribution extends across Alaska and most of Canada.

<i>Betula chichibuensis</i> Species of birch

Betula chichibuensis, commonly known as Chichibu birch, is a species of birch native exclusively to limestone outcrops in the Okuchichibu and Kitakami Mountains of central and northeast Honshu, Japan. The tree is rated as Critically Endangered on the IUCN Red List due to its extreme rarity and limited range.

References

  1. Stritch, L. (2014). "Betula nigra". IUCN Red List of Threatened Species . 2014: e.T194551A2348770. doi: 10.2305/IUCN.UK.2014-3.RLTS.T194551A2348770.en . Retrieved 19 November 2021.
  2. 1 2 3 Furlow, John J. (1997). "Betula nigra". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 3. New York and Oxford: Oxford University Press via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
  3. 1 2 Wojtech, Michael (2011). Bark:A Field Guide to Trees of the Northeast. Hanover and London: University Press of New England. pp. 110–111. ISBN   9781584658528.
  4. 1 2 3 Leopold, Donald Joseph; McComb, William C.; Muller, Robert N. (1998). Trees of the Central Hardwood Forests of North America. Portland: Timber Press. pp. 258–259. ISBN   0-88192-406-7.
  5. 1 2 McClung, Lester (1968). Forest Trees of West Virginia. Charleston: West Virginia Department of Natural Resources. p. 35.
  6. Grelen, H. E. (1990). "Betula nigra". In Burns, Russell M.; Honkala, Barbara H. (eds.). Hardwoods. Silvics of North America. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA). Vol. 2 via Southern Research Station.
  7. Murrell, Zack (2010). Vascular Plant Taxonomy, 6th edition. Dubuque: Kendall Hunt. pp. 258–259. ISBN   978-0-7575-7615-7.
  8. Glimn-Lacy, Kaufman (2006). Botany Illustrated. Springer. pp.  84. ISBN   0-387-28870-8.
  9. 1 2 3 4 5 6 7 Sullivan, Janet (1993). "Betula nigra". Fire Effects Information System (FEIS). US Department of Agriculture (USDA), Forest Service (USFS), Rocky Mountain Research Station, Fire Sciences Laboratory. Retrieved 2018-04-01.
  10. USDA, NRCS (n.d.). "Betula nigra". The PLANTS Database (plants.usda.gov). Greensboro, North Carolina: National Plant Data Team. Retrieved 26 January 2018.
  11. Robinson, Gaden S.; Ackery, Phillip R.; Kitching, Ian; Beccaloni, George W.; Hernández, Luis M. (2023). "HOSTS - The Hostplants and Caterpillars Database at the Natural History Museum". www.nhm.ac.uk. Natural History Museum. doi:10.5519/havt50xw . Retrieved 27 September 2022.
  12. 1 2 Harlow, W. M., & Harrar, E. S. (1969). Textbook Of Dendrology 5th ed., LOC# 68-17188
  13. 1 2 Woods, Katherine E; Chhetri, Bhuwan; Jones, Chelsea (October 2013). "Bioactivities and Compositions of Betula nigra Essential Oils". Journal of Medicinally Active Plants. 2 (1). doi:10.7275/R5639MP8. S2CID   80716012 . Retrieved 30 July 2016.