Brachauchenius Temporal range: Late Cretaceous, | |
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Holotype specimen of B. lucasi (USNM 4989) | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Superorder: | † Sauropterygia |
Order: | † Plesiosauria |
Family: | † Pliosauridae |
Subfamily: | † Brachaucheninae |
Genus: | † Brachauchenius Williston, 1903 [1] |
Type species | |
†Brachauchenius lucasi Williston, 1903 [1] |
Brachauchenius (meaning 'short neck') is an extinct genus of pliosaurid that lived in North America (United States) during the Late Cretaceous.
The type species, Brachauchenius lucasi, lived in the Western Inland Sea of North America around 100.5-89.3 million years ago, from the Cenomonian to the Turonian of the Cretaceous. An older specimen from the Barremian of Colombia was considered as part of this genus, representing the first re-occurrence of the non-rhomaleosaurid pliosaurs after a Berriasian-Hauterivian hiatus. [2] However, subsequent analysis of this Colombian specimen shows that it was enough distinctive to warrant a new genus and species, named as Stenorhynchosaurus munozi . [3]
The first known (type) specimen (USNM 4989) was collected by Charles Hazelius Sternberg from Ottawa County, Kansas in 1884. [4] [5] It had a skull length of up to 1 m (3.3 ft). [6] The species was named by Samuel W. Williston. [1] [7] The individual to which this specimen belongs would have measured 4–5 m (13–16 ft) long. [6] Brachauchenius represents the last known occurrence of a pliosaur in North America.
A larger specimen (FHSM VP-321 - skull length 170 cm) was collected by George Fryer Sternberg in 1952 from the Fairport Chalk of Russell County, Kansas, and later described by Carpenter. [8] Schumacher and Everhart (2005) reported on the age and locality of both Kansas specimens. [9] This specimen is now reassigned to its own genus and species, Megacephalosaurus eulerti . [6]
In 2013, Benson et al. referred a partial cranium from the Chalk Group of England (previously referred to Polyptychodon ) to Brachauchenius indet. [10]
The cladogram below follows a 2011 analysis by paleontologists Hilary F. Ketchum and Roger B. J. Benson, and reduced to genera only. [11]
Pliosauroidea |
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Plesiosauroidea is an extinct clade of carnivorous marine reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroids are known from the Jurassic and Cretaceous periods. After their discovery, some plesiosauroids were said to have resembled "a snake threaded through the shell of a turtle", although they had no shell.
Elasmosaurus (;) is a genus of plesiosaur that lived in North America during the Campanian stage of the Late Cretaceous period, about 80.5 million years ago. The first specimen was discovered in 1867 near Fort Wallace, Kansas, US, and was sent to the American paleontologist Edward Drinker Cope, who named it E. platyurus in 1868. The generic name means "thin-plate reptile", and the specific name means "flat-tailed". Cope originally reconstructed the skeleton of Elasmosaurus with the skull at the end of the tail, an error which was made light of by the paleontologist Othniel Charles Marsh, and became part of their "Bone Wars" rivalry. Only one incomplete Elasmosaurus skeleton is definitely known, consisting of a fragmentary skull, the spine, and the pectoral and pelvic girdles, and a single species is recognized today; other species are now considered invalid or have been moved to other genera.
Kronosaurus is a potentially dubious genus of extinct short-necked pliosaur. With an estimated length of 9 to 10.9 meters, it was among the largest pliosaurs, and is named after the leader of the Greek Titans, Kronos. It lived in the Early Cretaceous period. Fossil material has been recovered from the Toolebuc Formation and Wallumbilla Formations (Aptian) of Queensland and New South Wales in Australia.
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled pleisiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.
Liopleurodon is an extinct genus of large, carnivorous marine reptile belonging to the Thalassophonea, a clade of short-necked pliosaurid plesiosaurs. The two species of Liopleurodon lived from the Callovian Stage of the Middle Jurassic to the Kimmeridgian stage of the Late Jurassic Period. It was the apex predator of the Middle to Late Jurassic seas that covered Europe. The largest species, L. ferox, is estimated to have grown up to 6.6 metres (22 ft) in length, but could have been larger.
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae. Their diet mainly consisted of crustaceans and molluscs.
Plesiopleurodon is an extinct genus of Mesozoic marine reptiles, belonging to Sauropterygia, known from the Late Cretaceous of North America. It was named by Kenneth Carpenter based on a complete skull with a mandible, cervical vertebra, and a coracoid. In naming the specimen, Carpenter noted "Of all known pliosaurs, Plesiopleurodon wellesi most closely resembles Liopleurodon ferox from the Oxfordian of Europe, hence the generic reference." It was initially described as a pliosaur due to it short neck, a common trait of the family, although it is in the order Plesiosauria. However, later exploration into the relationships of both orders indicate that not all pliosaurs have short necks and not all plesiosaurs have long necks.
Trinacromerum is an extinct genus of sauropterygian reptile, a member of the polycotylid plesiosaurs. It contains two species, T. bentonianum and T. kirki. Specimens have been discovered in the Late Cretaceous fossil deposits of what is now modern Kansas and Manitoba.
Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874.
Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. Several species survived into the final stage of the Cretaceous, the Maastrichtian.
Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing three species, D. osborni, D. bonneri and D. tropicensis, as well as a questionably referred fourth species, D. herschelensis. Dolichorhynchops was an oceangoing prehistoric reptile. Its Greek generic name means "long-nosed face".
Leptocleidus is an extinct genus of plesiosaur, belonging to the family Leptocleididae.
Simolestes is an extinct pliosaurid genus that lived in the Middle to Late Jurassic. The type specimen, BMNH R. 3319 is an almost complete but crushed skeleton diagnostic to Simolestes vorax, dating back to the Callovian of the Oxford Clay formation, England. The genus is also known from the Callovian and Bajocian of France (S.keileni), and the Tithonian of India (S.indicus). The referral of these two species to Simolestes is dubious, however.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Megacephalosaurus is an extinct genus of short-necked pliosaur that inhabited the Western Interior Seaway of North America about 94 to 93 million years ago during the Turonian stage of the Late Cretaceous, containing the single species M. eulerti. It is named after its large head, which is the largest of any plesiosaur in the continent and measures up to 1.75 meters (5.7 ft) in length. Megacephalosaurus was one of the largest marine reptiles of its time with an estimated length of 6–9 meters (20–30 ft). Its long snout and consistently sized teeth suggest that it preferred a diet on smaller-sized prey.
Stenorhynchosaurus is an extinct genus of pliosaurid plesiosaurs which lived in South America during the Early Cretaceous. The type species and only known is Stenorhynchosaurus munozi. It was a medium-sized pliosaur, reaching an adult body length of 7 m (23 ft).
Luskhan is an extinct genus of brachauchenine pliosaur from the Cretaceous of Russia. The type and only species is Luskhan itilensis, named by Valentin Fischer and colleagues in 2017 from a well-preserved and nearly complete skeleton. As an early-diverging brachauchenine, Luskhan consequently exhibits an intermediate combination of traits seen in more basal and more derived pliosaurs. However, Luskhan departs significantly from other pliosaurs in that it exhibits a lack of adaptations in its skull to feeding on large prey; its slender snout, small teeth, and short tooth rows instead indicate a skull adapted for feeding on small, soft prey. In these features, it is the pliosaur that approaches closest to the distantly-related piscivorous polycotylids, having convergently evolved these traits more than 10 million years apart.
María Euridice Páramo Fonseca is a Colombian paleontologist and geologist. She has contributed on the paleontology in Colombia in the fields of describing various Cretaceous reptiles, most notably the mosasaurs Eonatator and Yaguarasaurus, the ichthyosaurs Kyhytysuka and Stenorhynchosaurus and the plesiosaur Leivanectes.
Sachicasaurus is an extinct genus of brachauchenine pliosaurid known from the Barremian of the Paja Formation, Altiplano Cundiboyacense in the Colombian Eastern Ranges of the Andes. The type species is S. vitae.