Brachychampsa

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Brachychampsa
Temporal range: Late Cretaceous, 72.1–66  Ma [1]
Brachychampsa sp. - Natural History Museum of Utah - DSC07238.JPG
Brachychampsa sp.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauromorpha
Clade: Archosauriformes
Order: Crocodilia
Family: Alligatoridae
Subfamily: Caimaninae
Genus: Brachychampsa
Gilmore, 1911 [2]
Species
  • B. montanaGilmore, 1911 (type) [2]
  • B. perrugosus(Cope, 1875)
  • ?B. sealeyiWilliamson, 1996 [3]

Brachychampsa is an extinct genus of alligatoroid, possibly a basal caiman. [4] Specimens have been reported from New Mexico, Colorado, Wyoming, [5] Montana, [2] [6] [7] North and South Dakota, [8] New Jersey, and Saskatchewan, though only those from Montana, Utah, and New Mexico are based on material sufficient to justify the referral. One specimen has been reported from the Darbasa Formation of Kazakhstan, although the species status is indeterminate for the fossil. The genus first appeared during the late Campanian stage of the Late Cretaceous (Judithian North American stage) and became extinct during the late Maastrichtian stage of the Cretaceous (Lancian North American Land Mammal "Age"). Brachychampsa is distinguished by an enlarged fifth maxillary tooth in the upper jaw.

Contents

Life reconstruction of Brachychampsa montana Brachychampsa NT small.jpg
Life reconstruction of Brachychampsa montana
Skull Brachychampsa sp. - Natural History Museum of Utah - DSC07244.JPG
Skull

Species

The type species of Brachychampsa is B. montana, first discovered from the Hell Creek Formation of Montana and described by Charles W. Gilmore in a paper in 1911. In that same paper, Gilmore recombined Bottosaurus perrugosus as a new species of Brachychampsa, called B. perrugosus. [2] The holotype specimen of B. perrugosus went missing as the paper was being written, but it was later rediscovered and soon afterward designated as a nomen dubium due to a lack of diagnostic features that distinguish it from other alligatorids discovered since the paper was published. [9] Another species from the Allison Member of the Menefee Formation of the San Juan Basin, B. sealeyi, was discovered in 1996, [3] but was later argued to be synonymous with B. montana by interpreting it as an immature specimen of the latter species. [10] However, other studies have shown that some of the variation seen between the two species, such as the orientation of the maxillary tooth row, are not ontogenic, thus making B. sealeyi a valid taxon. [11]

Phylogeny

Brachychampsa's position within Alligatoroidea has undergone many revisions since it was first named. Originally it was placed within Alligatoridae, and was later refined to the Alligatorinae in 1964, [5] only to be placed outside both Alligatorinae and Alligatoridae (but still within Alligatoroidea) in 1994. [9] Accordingly, studies have shown Brachychampsa as a basal member of Alligatoroidea, within the clade Globidonta, [12] [13] as shown in the cladogram below. [13]

Alligatoroidea

Alternatively, other phylogenetic studies have recovered Brachychampsa as an alligatorid, specifically as a stem-caiman, [14] [15] as shown in the cladogram below. [14]

Alligatoroidea
(stem-based group)

Related Research Articles

<span class="mw-page-title-main">Alligatoridae</span> Family of crocodilians including alligators, caimans and kin

The family Alligatoridae of crocodylians includes alligators, caimans and their extinct relatives.

<i>Ceratosuchus</i> Extinct genus of reptiles

Ceratosuchus is an extinct genus of alligatorine crocodylian from latest Paleocene rocks of Colorado's Piceance Basin and earliest Eocene rocks of Wyoming's Bighorn Basin in North America, a slice of time known as the Clarkforkian North American Land Mammal Age. Like its modern relatives, Ceratosuchus was a swamp-dwelling predator. It is named for the pair of flattened, triangular bony plates that extend from the back of its head.

<i>Diplocynodon</i> Extinct genus of reptiles

Diplocynodon is an extinct genus of alligatoroid crocodilian that lived during the Paleocene to Middle Miocene in Europe. Some species may have reached lengths of 3 metres (9.8 ft), while others probably did not exceed 1 metre (3.3 ft). They are almost exclusively found in freshwater environments. The various species are thought to have been opportunistic aquatic predators.

Wannaganosuchus is an extinct genus of small alligatorid crocodilian. It was found in Late Paleocene-age rocks of Billings County, North Dakota, United States.

<i>Procaimanoidea</i> Extinct genus of reptiles

Procaimanoidea is an extinct genus of alligatorid from the Eocene of North America. It was named posthumously in 1946 by Charles W. Gilmore; the type species is P. utahensis, from the Uintan of Utah. It is based on USNM 15996, a nearly complete skull and partial left hind leg. A second species, P. kayi, was named in 1941 by C.C. Mook as a species of Hassiacosuchus, for remains from the Bridgerian of Wyoming. It was reassigned to Procaimanoidea in 1967 by Wassersug and Hecht.

<i>Leidyosuchus</i> Genus of reptiles

Leidyosuchus is an extinct genus of alligatoroid from the Late Cretaceous of Alberta. It was named in 1907 by Lawrence Lambe, and the type species is L. canadensis. It is known from a number of specimens from the middle Campanian age Dinosaur Park Formation. It was a medium-sized alligatorid, with a maximum skull length greater than 40 centimeters (16 in).

<span class="mw-page-title-main">Caiman</span> Subfamily of reptiles

A caiman is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico and Central and South America from marshes and swamps to mangrove rivers and lakes. They have scaly skin and live a fairly nocturnal existence. They are relatively small-sized crocodilians with an average maximum weight of 6 to 40 kg depending on species, with the exception of the black caiman, which can grow more than 4 m (13 ft) in length and weigh in excess of 1,000 kg. The black caiman is the largest caiman species in the world and is found in the slow-moving rivers and lakes that surround the Amazon basin. The smallest species is the Cuvier's dwarf caiman, which grows to 1.2 to 1.5 m long. There are six different species of caiman found throughout the watery jungle habitats of Central and Southern America. The average length for most of the other caiman species is about 2 to 2.5 m long.

<span class="mw-page-title-main">Alligatoroidea</span> Superfamily of reptiles

Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.

Eocaiman is an extinct genus of caiman containing species living from the Early Paleocene to Miocene in what is now Argentina, Itaboraí Formation of Brazil and Colombia. Eocaiman contains three described species: E. cavernensis, E. palaeocenicus, and E. itaboraiensis, and is typically recovered as one of the more basal members of Caimaninae. Notocaiman was synonymized with Eocaiman paleocenicus in 2022.

<i>Allognathosuchus</i> Extinct genus of reptiles

Allognathosuchus is an extinct genus of alligatorine crocodylian with a complicated taxonomic history. It was named in 1921.

<i>Navajosuchus</i> Extinct genus of reptiles

Navajosuchus is an extinct genus of alligatorine crocodylian. Its fossils have been found in the Paleocene-age Nacimiento Formation of the San Juan Basin, New Mexico. It was named in 1942 by Charles C. Mook, and the original type species was N. novomexicanus. N. novomexicanus was based on AMNH 5186, a partial skull collected in 1913. Later research showed that Navajosuchus novomexicanus was the same as the earlier-named Allognathosuchus mooki. However, A. mooki does not belong to the genus Allognathosuchus, and so the name of the crocodilian becomes Navajosuchus mooki. Under whichever name is used, this animal would have been a generalized predator of the Nacimiento floodplains. It was the most common Nacimiento Formation crocodilian, found in both the Puercan and Torrejonian faunal assemblages.

<i>Acynodon</i> Extinct genus of reptiles

Acynodon is an extinct genus of eusuchian crocodylomorph from the Late Cretaceous, with fossils found throughout Southern Europe.

Arambourgia is an extinct monotypic genus of alligatorine crocodylian from Europe. It was named in 1905 as Allognathosuchus gaudryi. It was made a separate genus Arambourgia in 1940. This was synonymized with Allognathosuchus haupti in 1990, but later reassigned as its own genus once again in 2004. Arambourgia was likely to have been part of an early dispersal event of alligatorines from North America to Europe during the Eocene epoch. Arambourgia had non-serrated teeth and a deep orienirostral snout, unlike the flatter snouts of most other alligatorids.

<span class="mw-page-title-main">Menefee Formation</span> Geologic formation in New Mexico and Colorado

The Menefee Formation is a lower Campanian geologic formation found in Colorado and New Mexico, United States.

<i>Planocrania</i> Extinct genus of reptiles

Planocrania is an extinct genus of eusuchian crocodyliforms from what is now China. Two species are currently known to belong to the genus.

<i>Stangerochampsa</i> Extinct genus of reptiles

Stangerochampsa is an extinct genus of globidontan alligatoroid, possibly an alligatorine or a stem-caiman, from the Late Cretaceous of Alberta. It is based on RTMP.86.61.1, a skull, partial lower jaws, and partial postcranial skeleton discovered in the late Campanian–early Maastrichtian-age Horseshoe Canyon Formation. Stangerochampsa was described in 1996 by Wu and colleagues. The type species is S. mccabei. The generic name honors the Stanger family, the owners of the ranch where the specimen was found, and the species name honors James Ross McCabe, who discovered, collected, and prepared it. Stangerochampsa is described as "small to medium–sized"; the type skull is 20.0 centimetres (7.9 in) long from the tip of the snout to the occipital condyle, and is 13.0 centimetres (5.1 in) wide at its greatest, while the thigh bone is 14.2 centimetres (5.6 in) long. It had heterodont dentition, with large crushing teeth at the rear of the jaws.

<span class="mw-page-title-main">Globidonta</span> Clade of reptiles

Globidonta is a clade of alligatoroids that includes alligators, caimans, and closely related extinct forms. It is defined as a stem-based clade including Alligator mississippiensis and all forms more closely related to it than to Diplocynodon. The group's fossil range extends back into the Late Cretaceous with early alligatoroids such as Albertochampsa and Brachychampsa. Extinct globidontans were particularly common in North America and Eurasia, and their modern range also includes South America.

Alligator mefferdi is an extinct species of alligator described by Charles Craig Mook. They lived in the Miocene period, and their range was principally in what is now Nebraska, United States. The type specimen was discovered in the Ash Hollow Formation at Ash Hollow State Historical Park.

<i>Menefeeceratops</i> Extinct genus of dinosaurs

Menefeeceratops is a genus of ceratopsid dinosaur from the Menefee Formation in New Mexico, United States. It is potentially the oldest known member of the ceratopsids, as well as the centrosaurine subfamily, related to animals including Yehuecauhceratops and Crittendenceratops. The type and only species is Menefeeceratops sealeyi, known from a partial, non-articulated skeleton.

Jiangxisuchus is an extinct genus of crocodylian that lived during the Late Cretaceous in what is now China. It was described in 2019, and was proposed to be a basal member of Crocodyloidea. However, another concurrent 2019 study recovered Jiangxisuchus instead as a basal member of Alligatoroidea, within the newly named clade Orientalosuchina.

References

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