Brille

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Clouded brille of a colubrid nearing moulting AB124 Brille.JPG
Clouded brille of a colubrid nearing moulting
Clear brille of the same snake - 20 days earlier AB125 - Chequered Keelback head.JPG
Clear brille of the same snake - 20 days earlier
Exuvia of grass snake Natrix natrix, showing brille scales Ormskinn (Natrix natrix) skannat.jpg
Exuvia of grass snake Natrix natrix , showing brille scales

The brille (also called the ocular scale, eye cap or spectacle) is the layer of transparent, immovable disc-shaped skin or scale covering the eyes of some animals for protection, especially in animals without eyelids. In squamate reptiles both the nictitating membrane and the eyelids have been suggested as the evolutionary origin of the brille, but embryonic studies supports the fusion of the dorsal and ventral eyelids. [1] Brille means "spectacles" or "glasses" in German, Norwegian, and Danish.

In snakes, there are no eyelids and the brille is clear and cannot be distinguished, except when the animal is becoming ready for ecdysis. At that time, it becomes cloudy and is visible as a cover over the eye. When the snake moults, the brille is also shed, generally inside out, as part of its skin. The brilles protect their eyes from dust and dirt and give them a "glassy-eyed" blank appearance. [2]

Snakes, flap-footed lizards, night lizards, and some skinks have brilles. All geckos except those in the subfamily Eublepharinae (eyelid geckos) possess brilles.

Some groups of bony fish have a transparent eyelid known as the adipose eyelid. [3] Some reptiles, mammals and birds have a translucent third eyelid that moves horizontally across the eye called the nictitating membrane. [4]

See also

Related Research Articles

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<span class="mw-page-title-main">Lizard</span> Informal group of reptiles

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<span class="mw-page-title-main">Snake</span> Limbless, scaly, elongate reptile

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<span class="mw-page-title-main">Tears</span> Clear liquid secreted from glands in eyes of mammals

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<span class="mw-page-title-main">Gymnophthalmidae</span> Family of lizards

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<span class="mw-page-title-main">Eyelid</span> Thin fold of skin that covers and protects the eye

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<span class="mw-page-title-main">Plica semilunaris of conjunctiva</span> Small fold of mucous membrane in the eye.

The plica semilunaris is a small fold of bulbar conjunctiva on the medial canthus of the eye. It functions during movement of the eye, to help maintain tear drainage via the lacrimal lake, and to permit greater rotation of the globe, for without the plica the conjunctiva would attach directly to the eyeball, restricting movement. It is the vestigial remnant of the nictitating membrane which is drawn across the eye for protection, and is present in other animals such as birds, reptiles, and fish, but is rare in mammals, mainly found in monotremes and marsupials. Its associated muscles are also vestigial. It is loose, thus eye movements are not restricted by it. Only one species of primate, the Calabar angwantibo, is known to have a functioning nictitating membrane.

<span class="mw-page-title-main">Snake scale</span> Scales covering the skin of snakes

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In scaled reptiles, the ocular scales are those forming the margin of the eye. The name originates from the term oculus which is Latin for 'eye' and, in the broadest sense, refers to a scale associated with the eye. The numbers of these scales present, and sometimes the shapes and sizes, are some of many characteristics used to differentiate species from one another.

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<span class="mw-page-title-main">Eagle eye</span>

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<span class="mw-page-title-main">Albinism</span> Disorder causing lack of pigmentation

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References

  1. Through the Looking Glass: The Spectacle in Gymnophthalmid Lizards
  2. "Encyclopedia of Animals (Milk snake entry)", EBSCO Animals, EBSCO Publishing
  3. Froese, Rainer. "Adipose eyelid". FishBase Glossary. Retrieved 2011-12-11.
  4. Anon. "Nictiiating membrane". The Free Dictionary. Farlex Inc. Retrieved 3 November 2013.