Clelandina

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Clelandina
Temporal range: Late Permian 259–254  Ma
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Clelandina.png
Holotype skull of C. rubidgei
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Gorgonopsia
Family: Gorgonopsidae
Tribe: Rubidgeini
Genus: Clelandina
Broom, 1948
Type species
Clelandina rubidgei
Broom, 1948
Synonyms

Genus-level

  • Tigrisaurus
    Broom & George, 1950
  • Dragocephalus
    Brink & Kitching, 1953

Species-level

  • Clelandina majorBroom, 1940
  • Tigrisaurus priceiBroom & George, 1950
  • Dracocephalus scheepersiBrink & Kitching, 1953
  • Dinogorgon (Dracocephalus) scheepersiWatson & Romer, 1956
  • Clelandina scheepersiSigogneau, 1970
  • Dinogorgon priceiSigogneau, 1970
  • Rubidgea priceiGebauer, 2007
Restoration of C. rubidgei Clelandina1DB.jpg
Restoration of C. rubidgei
Referred skull, the holotype of Tigrisaurus pricei Tigrisaurus pricei.png
Referred skull, the holotype of Tigrisaurus pricei

Clelandina is an extinct genus of rubidgeine gorgonopsian from the Late Permian of Cistecephalus Assemblage Zone of South Africa. It was first named by Broom in 1948. The type and only species is C. rubidgei. It is relatively rare, with only four known specimens. [1] [2]

Contents

Description

Clelandina rubidgei has an extraordinarily small sclerotic ring relative to the size of its orbit, which implies that it was diurnal. It is the only rubidgeine with a preserved sclerotic ring, so it is unknown whether this trait was shared by other members of the subfamily. Like all rubidgeines, it was relatively large, with a skull up to 36 cm long. It had reduced dentition, with the teeth posterior to the canines being absent and replaced with a bony ridge. The skull has heavily pachyostosed, with massive rugose bosses. [1]

Classification

Clelandina shares many characteristics with the contemporary Rubidgea , and is currently recognized as the sister taxon of this genus. Together with Dinogorgon and Leontosaurus , these genera form Rubidgeini, a clade of large gorgonopsians with distinctively robust, broad, and pachyostosed skulls. [1]

Below is a cladogram recovered by Christian Kammerer in 2016. [1]

Rubidgeinae

Smilesaurus ferox

Aelurognathus tigriceps

Ruhuhucerberus haughtoni

Sycosaurus laticeps

Sycosaurus nowaki

Rubidgeini

Leontosaurus vanderhosti

Dinogorgon rubidgei

Rubidgea atrox

Clelandina rubidgei

See also

Related Research Articles

<span class="mw-page-title-main">Gorgonopsia</span> Extinct group of saber-toothed therapsids from the Permian

Gorgonopsia is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw.

<i>Cistecephalus</i> Extinct genus of dicynodonts

Cistecephalus is an extinct genus of dicynodont therapsid from the Late Permian of southern Africa. It was a small, specialised, burrowing dicynodont, possibly with habits similar to a modern mole. The head was flattened and wedge-shaped, the body long, and the forelimbs very strong, with similarities in structure to the forelimb of modern burrowing mammals.

<i>Inostrancevia</i> Extinct genus of therapsids

Inostrancevia is an extinct genus of large carnivorous therapsids which lived during the Late Permian in what is now European Russia and South Africa. The first known fossils of this gorgonopsian were discovered in the Northern Dvina, where two almost complete skeletons were exhumed. Subsequently, several other fossil materials were discovered in various oblasts, and these finds will lead to a confusion about the exact number of valid species in the country, before only three of them were officially recognized : I. alexandri, I. latifrons and I. uralensis. More recent research carried out in South Africa has discovered fairly well-preserved remains of the genus, being attributed to the species I. africana. The whole genus is named in honor of Alexander Inostrantsev, professor of Vladimir P. Amalitsky, the paleontologist who described the taxon.

<i>Dinogorgon</i> Extinct genus of therapsids

Dinogorgon is a genus of gorgonopsid from the Late Permian of South Africa and Tanzania. The generic name Dinogorgon is derived from Greek, meaning "terrible gorgon", while its species name rubidgei is taken from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. The type species of the genus is D. rubidgei.

Sclerotic rings are rings of bone found in the eyes of many animals in several groups of vertebrates, except for mammals, amphibians, and crocodilians. The ring is located within the soft tissue of the sclera, commonly known as the "white" of the eye.

<i>Sauroctonus</i> Extinct genus of therapsids

Sauroctonus is an extinct genus of gorgonopsian therapsids who lived during the end of the Middle Permian in what is now European Russia. The first fossils, discovered in Tatarstan, were first considered to belong to a new species of the South African genus Arctognathus, named A. progressus in 1938. The taxon will be designated as such until 1940, when it will be assigned to the genus Inostrancevia by Ivan Yefremov, before being definitively classified in a separate genus erected by Alexey Bystrow in 1955. The most complete known fossils of S. progressus include cranial and postcranial elements currently all recorded in Tatarstan. These elements show that the animal is a rather medium-sized gorgonopsian.

<i>Arctops</i> Extinct genus of therapsids

Arctops is an extinct genus of gorgonopsian therapsids known from the Late Permian of South Africa. It measured up to 2 metres in length and its skull was 30 centimetres (12 in) long. The type species is Arctops willistoni. A second species, A. watsoni, may be synonymous with A. willistoni. A. kitchingi may be a third species of Arctops, but it was only tentatively assigned to the genus when it was first named. Both were formally synonymized with A. willistoni by Christian Kammerer in 2017.

<i>Viatkogorgon</i> Extinct genus of therapsids

Viatkogorgon is a genus of gorgonopsian that lived during the Permian period in what is now Russia. The first fossil was found at the Kotelnich locality near the Vyatka River and was made the holotype of the new genus and species V. ivachnenkoi in 1999. The generic name refers to the river and the related genus Gorgonops—the gorgons of Greek mythology are often referenced in the names of the group. The specific name honors the paleontologist Mikhail F. Ivakhnenko. The holotype skeleton is one of the most complete gorgonopsian specimens known and includes rarely preserved elements such as gastralia and a sclerotic ring. A larger, but poorly preserved specimen has also been assigned to the species.

<i>Rubidgea</i> Extinct genus of therapsids

Rubidgea is a genus of gorgonopsid from the upper Permian of South Africa and Tanzania, containing the species Rubidgea atrox. The generic name Rubidgea is sometimes believed to be derived from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. However, this generic name was actually erected in honor of Rubidge's paternal grandfather, Sydney Rubidge, who was a renowned fossil hunter. Its species name atrox is derived from Latin, meaning “fierce, savage, terrible”. Rubidgea is part of the gorgonopsian subfamily Rubidgeinae, a derived group of large-bodied gorgonopsians restricted to the Late Permian (Lopingian). The subfamily Rubidgeinae first appeared in the Tropidostoma Assemblage Zone. They reached their highest diversity in the Cistecephalus and Daptocephalus assemblage zones of the Beaufort Group in South Africa.

<i>Aelurosaurus</i> Extinct genus of therapsids

Aelurosaurus is a small, carnivorous, extinct genus of gorgonopsian therapsids from the Late Permian of South Africa. It was discovered in the Karoo Basin of South Africa, and first named by Richard Owen in 1881. It was named so because it appeared to be an ancestor for cat-like marsupials, but not yet a mammal itself. It contains five species, A. felinus, A. whaitsi, A. polyodon, A. wilmanae, and A.? watermeyeri. A. felinus, the type species, is generally well described with established features, while the other four species are not due to their poorly preserved holotypes.

<i>Pravoslavlevia</i> Extinct genus of therapsids

Pravoslavlevia is an extinct genus of gorgonopsian therapsids that lived in the late Permian and is part of the Sokolki subcomplex of Russia. It had a skull 22 centimetres (8.7 in) long. The total length of the animal was about 1.4 metres. Only one species is known.

<i>Eriphostoma</i> Extinct genus of therapsids

Eriphostoma is an extinct genus of gorgonopsian therapsids known from the Middle Permian of Tapinocephalus Assemblage Zone, South Africa. It has one known species, Eriphostoma microdon, and was first named by Robert Broom in 1911. It is the oldest known gorgonopsian and among the smallest and most basal members of the clade.

<i>Pelanomodon</i> Extinct genus of dicynodonts

Pelanomodon is an extinct genus of dicynodont therapsids that lived in the Late Permian period. Fossil evidence of this genus is principally found in the Karoo Basin of South Africa, in the Dicynodon Assemblage Zone. Lack of fossil record after the Late Permian epoch suggests that Pelanomodon fell victim to the Permian-Triassic extinction event.

<i>Smilesaurus</i> Extinct genus of therapsids

Smilesaurus is an extinct genus of gorgonopsian known from Africa. It lived during the Late Permian. It contains the single species S. ferox.

"Dixeya" nasuta is a species of the predatory gorgonopsian therapsids from the Late Permian of East Africa, known from fossils found in what is now Tanzania. The species has a complicated taxonomic history, it was originally named as a second species of the genus Dixeya which is now considered a junior synonym of Aelurognathus. "D." nasuta itself, however, was not moved to Aelurognathus, and although it was instead tentatively referred to Arctognathus at first it has since been recognised to not belong to this genus either. This situation leaves "Dixeya" nasuta without a formal genus name. It was proposed to belong to a new distinct genus, named "Njalila", that was informally proposed for the species in a PhD thesis, but this name has not yet been formally published and is currently a nomen nudum. "D." nasuta has been characterised from other gorgonopsians by a combination of its straight snout profile, upturned and 'pinched' nose, and curved jaw margin.

<span class="mw-page-title-main">Rubidgeinae</span> Extinct subfamily of mammals

Rubidgeinae is an extinct subfamily of gorgonopsid therapsids known only from Africa. They were among the largest gorgonopsians, and their fossils are common in the Cistecephalus and Daptocephalus assemblage zones of the Karoo Basin. They lived during the Late Permian, and became extinct at the end of the Permian.

<i>Bulbasaurus</i> Extinct genus of dicynodonts

Bulbasaurus is an extinct genus of dicynodont that is known from the Lopingian epoch of the Late Permian period of what is now South Africa, containing the type and only species B. phylloxyron. It was formerly considered as belonging to Tropidostoma; however, due to numerous differences from Tropidostoma in terms of skull morphology and size, it has been reclassified the earliest known member of the family Geikiidae, and the only member of the group known from the Tropidostoma Assemblage Zone. Within the Geikiidae, it has been placed close to Aulacephalodon, although a more basal position is not implausible.

Leogorgon is an extinct genus of dubious therapsid from the Late Permian Sokolki Faunal Assemblage of Russia. It was originally classified as a rubidgeine gorgonopsian, and would have been the first member of that clade from outside of Africa if that identification had been valid. However, it may instead be a combination of the tooth of a gorgonopsian and the braincase of a dicynodont, and may be a wastebin taxon.

<i>Nochnitsa</i> Extinct genus of therapsids

Nochnitsa is an extinct genus of gorgonopsian therapsids who lived during an uncertain stage of the Permian in what is now European Russia. Only one species is known, N. geminidens, described in 2018 from a single specimen including a complete skull and some postcranial remains, discovered in the red beds of Kotelnich, Kirov Oblast. The genus is named in reference to Nocnitsa, a nocturnal creature from Slavic mythology. This name is intended as a parallel to the Gorgons, which are named after many genera among gorgonopsians, as well as for the nocturnal behavior inferred for the animal. The only known specimen of Nochnitsa is one of the smallest gorgonopsians identified to date, with a skull measuring close to 8 cm (3.1 in) in length. The rare postcranial elements indicate that the animal's skeleton should be particularly slender.

Phorcys is an extinct genus of gorgonopsian that lived during the Middle Permian period (Guadalupian) of what is now South Africa. It is known from two specimens, both portions from the back of the skull, that were described and named in 2022 as a new genus and species P. dubei by Christian Kammerer and Bruce Rubidge. The generic name is from Phorcys of Greek mythology, the father of the Gorgons from which the gorgonopsians are named after, and refers to its status as one of the oldest representatives of the group in the fossil record. Phorcys was recovered from the lowest strata of the Tapinocephalus Assemblage Zone (AZ) of the Beaufort Group, making it one of the oldest known gorgonopsians in the fossil record—second only to fragmentary remains of an indeterminate gorgonopsian from the older underlying Eodicynodon Assemblage Zone.

References

  1. 1 2 3 4 Kammerer, Christian F. (2016). "Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)". PeerJ. 4: e1608. doi: 10.7717/peerj.1608 . PMC   4730894 . PMID   26823998.
  2. "Fossilworks: Clelandina".

Further reading