Flexibilia

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Flexibilia
Temporal range: Middle Ordovician–Permian
Onychocrinus ramulosus.JPG
Onychocrinus ramulosus
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Echinodermata
Class: Crinoidea
Parvclass: Cladida
Superorder: Flexibilia
Zittel, 1895
Orders

Flexibilia is a superorder of specialized Paleozoic (Middle Ordovician to Permian) crinoids. They exhibited a conserved body plan and consistent suite of characteristics throughout their long history. Previously considered a subclass with unclear affinities, later investigation determined that flexibles are well-nested within Cladida, a broad group ancestral to living articulate crinoids. [1] The Ordovician cladid Cupulocrinus acts as an intermediate form linking the generalized anatomy of other early cladids with the distinctive anatomy of flexibles, and several studies have considered it to be ancestral to the rest of the group. [2] [1] [3] Although flexibles never reached the same abundance or diversity as many other crinoid groups, they remained a reliable component of crinoid faunas, particularly from the Silurian onwards. Flexible fossils are very rare in the Ordovician (the most common taxa being Cupulocrinus and Protaxocrinus ), but the Late Ordovician appears to have been an interval of rapid diversification for the group. [3]

Major traits of Flexibilia include: [1] [4]

Flexibilia is strongly supported as a monophyletic clade. [1] [3] It is traditionally split into the orders Sagenocrinida and Taxocrinida, [4] though the monophyly of these orders has been called into question. The most recent discussion of Flexibilia taxonomy considers Taxocrinida to be paraphyletic and ancestral to Sagenocrinida. [3]

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<span class="mw-page-title-main">Cincta</span> Extinct class of marine invertebrates

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References

  1. 1 2 3 4 Wright, David F.; Ausich, William I.; Cole, Selina R.; Peter, Mark E.; Rhenberg, Elizabeth C. (2017). "Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata)". Journal of Paleontology. 91 (4): 829–846. doi: 10.1017/jpa.2016.142 . ISSN   0022-3360. S2CID   13806992.
  2. Peter, M. E. (2019-05-15). "Aberrations in the infrabasal circlet of the cladid crinoid genus Cupulocrinus (Echinodermata) and implications for the origin of flexible crinoids". Palaeogeography, Palaeoclimatology, Palaeoecology. 522: 52–61. doi: 10.1016/j.palaeo.2019.03.002 . ISSN   0031-0182. S2CID   134102417.
  3. 1 2 3 4 Wright, David F.; Toom, Ursula (2017). Sevastopulo, George (ed.). "New crinoids from the Baltic region (Estonia): fossil tip‐dating phylogenetics constrains the origin and Ordovician–Silurian diversification of the Flexibilia (Echinodermata)". Palaeontology. 60 (6): 893–910. doi: 10.1111/pala.12324 . ISSN   0031-0239. S2CID   134049045.
  4. 1 2 Moore, Raymond C. (1978). Moore, Raymond C.; Teichert, Curt (eds.). Part T, Echinodermata 2, Crinoidea, vol. 2. Treatise on invertebrate paleontology. By Georges Ubaghs, R. C. Moore, H. Wienberg Rasmussen, N. Gary Lane, Albert Breimer, H. L. Strimple, J. C. Brower, Russell M. Jeffords, James Sprinkle, R. E. Peck, D. B. Macurda, Jr., D. L. Meyer, Michel Roux, Hertha Sieverts-Doreck, R. O. Fay, and R. A. Robison. Boulder, CO and Lawrence, KS: University of Kansas and Geological Society of America. ISBN   978-0-8137-3021-9. OCLC   531991.