Giant tapir

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Giant tapir
Temporal range: Middle to Late Pleistocene
MegatapirusskeletalAugustiosPaleo.png
Skeletal reconstruction of the holotype.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Tapiridae
Genus: Tapirus
Species:
T. augustus
Binomial name
Tapirus augustus
Matthew & Granger, 1923
Synonyms
  • Megatapirus augustus(Matthew & Granger, 1923)
A restoration of Tapirus augustus MegatapirusDB.jpg
A restoration of Tapirus augustus

The giant tapir (Tapirus augustus) [1] [2] [3] is an extinct species of tapir that lived in southern China, Vietnam and Laos, [4] with reports suggesting it also lived in Taiwan, [5] Java, and potentially Borneo. [6] The species has been recorded from Middle and Late Pleistocene. [7] There is only weak evidence for a Holocene survival. [8] Tapirus augustus was larger than any living tapir, [1] with an estimated weight of about 623 kilograms (1,373 lb). [9] The species was also placed in its own genus of Megatapirus, however, it is now conventionally placed within Tapirus .

Discovery and taxonomy

Despite not being named until 1923, the Palaeontological Museum, Munich Paleontologist Max Schlosser described several teeth purchased from Chinese drug stores in 1903 that he assigned to Tapirus sinensis. [10] Some of the teeth had been unearthed at the Chang I locality in Wanzhou, Eastern Sichuan, China that come from the Pleistocene strata of the area. [10] Tapirus augustus was first described in 1923 William Diller Matthew and Walter Granger based on fossils found by the American Museum of Natural History during the Central Asiatic Expeditions of 1920–1930. The fossils had been recovered just a few miles from the site where many of the teeth described by Schlosser had been found.

Related Research Articles

<span class="mw-page-title-main">Perissodactyla</span> Order of hoofed mammals

Perissodactyla is an order of ungulates. The order includes about 17 living species divided into three families: Equidae, Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines, rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do.

<span class="mw-page-title-main">Tapir</span> Herbivorous mammal native to South and Central America, and Southeast Asia

Tapirs are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America and Southeast Asia. They are one of three extant branches of Perissodactyla, alongside equines and rhinoceroses. Only a single genus, Tapirus, is currently extant. Tapirs migrated into South America during the Pleistocene epoch from North America after the formation of the Isthmus of Panama as part of the Great American Interchange. Tapirs were formerly present across North America, but became extinct in the region at the end of the Late Pleistocene, around 12,000 years ago.

<i>Megaloceros</i> Extinct genus of deer

Megaloceros is an extinct genus of deer whose members lived throughout Eurasia from the Pleistocene to the early Holocene. The type and only undisputed member of the genus, Megaloceros giganteus, vernacularly known as the "Irish elk" or "giant deer", is also the best known. Fallow deer are thought to be their closest living relatives. Megaloceros is thought to be closely related to the East Asian genus Sinomegaceros, as well as possibly other extinct genera of "giant deer".

<i>Palaeoloxodon</i> Genus of extinct elephants

Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.

<i>Tapirus</i> Genus of tapir

Tapirus is a genus of tapir which contains the living tapir species. The Malayan tapir is usually included in Tapirus as well, although some authorities have moved it into its own genus, Acrocodia.

<span class="mw-page-title-main">Litopterna</span> Extinct order of hoofed mammals

Litopterna is an extinct order of South American native ungulates that lived from the Paleocene to the end of the Pleistocene-early Holocene around 63 million-12,000 years ago, and were also present in Antarctica during the Eocene. They represent the second most diverse group of South American ungulates after Notoungulata. It is divided into nine families, with Proterotheriidae and Macraucheniidae being the most diverse and last surviving families.

<i>Stegodon</i> Genus of extinct proboscidean

Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon remained widespread in South, Southeast and East Asia until the end of the Pleistocene.

<i>Toxodon</i> Extinct genus of notoungulates

Toxodon is an extinct genus of large ungulate native to South America from the Late Miocene to early Holocene epochs. Toxodon is a member of Notoungulata, an order of extinct South American native ungulates distinct from living ungulate groups that had been indigenous to the continent for over 60 million years since the early Cenozoic, prior to the arrival of living ungulates into South America around 2.5 million years ago during the Great American Interchange. Toxodon is a member of the family Toxodontidae, which includes medium to large sized herbivores. Toxodon was one of the largest members of Toxodontidae and Notoungulata, with Toxodon platensis having an estimated body mass of 1,000–1,200 kilograms (2,200–2,600 lb).

<i>Macrauchenia</i> Extinct genus of litopterns

Macrauchenia is an extinct genus of large ungulate native to South America from the late Pliocene to the end of the Pleistocene. It is a member of the extinct order Litopterna, a group of South American native ungulates which had been present in South America since the early Cenozoic, over 60 million years ago, prior to the arrival of living ungulates in South America around 2.5 million years ago as part of the Great American Interchange. The bodyform of Macrauchenia has been described as similar to a camel, being one of the largest known litopterns, with an estimated body mass of around 1 tonne. The genus gives its name to its family, Macraucheniidae, which like Macrauchenia typically had long necks and three toed feet, as well a retracted nasal region, which in Macrauchenia manifests as the nasal opening being on the top of the skull behind the eye sockets. This has historically been argued to correspond to the presence of a tapir-like proboscis, but some recent authors suggest a moose-like prehensile lip is more likely.

<i>Doedicurus</i> An extinct genus of mammals belonging to the armadillo order, Cingulata

Doedicurus is an extinct genus of glyptodont from South America containing one species, D. clavicaudatus. Glyptodonts are a member of the family Chlamyphoridae, which also includes some modern armadillo species, and they are classified in the superorder Xenarthra alongside sloths and anteaters. Being a glyptodont, it was a rotund animal with heavy armor and a carapace. Averaging at an approximate 1,400 kg (3,100 lb), it was one of the largest glyptodonts to have ever lived. Though glyptodonts were quadrupeds, large ones like Doedicurus may have been able to stand on two legs like other xenarthrans. It notably sported a spiked tail club, which may have weighed 40 or 65 kg in life, and it may have swung this in defense against predators or in fights with other Doedicurus at speeds of perhaps 11 m/s.

<i>Xenorhinotherium</i> Extinct genus of litopterns

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<span class="mw-page-title-main">Late Pleistocene extinctions</span> Extinction event occurring during the late Quaternary period

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<span class="mw-page-title-main">Stegodontidae</span> Family of extinct elephant-like mammals

Stegodontidae is an extinct family of proboscideans from Africa and Asia from the Early Miocene to the Late Pleistocene. It contains two genera, the earlier Stegolophodon, known from the Miocene of Asia and the later Stegodon, from the Late Miocene to Late Pleistocene of Africa and Asia which is thought to have evolved from the former. The group is noted for their plate-like lophs on their teeth, which are similar to elephants and different from those of other extinct proboscideans like gomphotheres and mammutids, with both groups having a proal jaw movement utilizing forward strokes of the lower jaw. These similarities with modern elephants were probably convergently evolved. Like elephantids, stegodontids are thought to have evolved from gomphothere ancestors.

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<i>Tapirus californicus</i> Extinct species of tapir

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<i>Tapirus veroensis</i> Extinct species of mammal

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References

  1. 1 2 Matthew, William Diller; Granger, Walter; Andrews, Roy Chapman (1923). "New fossil mammals from the Pliocene of Sze-chuan, China". Bulletin of the AMNH. 48: 17. hdl:2246/1308.
  2. Janis, Christine M.; Scott, Kathleen M.; Jacobs, Louis L. (1998-05-28). Evolution of Tertiary Mammals of North America: Volume 1, Terrestrial Carnivores, Ungulates, and Ungulate Like Mammals. Cambridge University Press. ISBN   9780521355193.
  3. "Fossilworks: Tapirus (Megatapirus) augustus". fossilworks.org. Retrieved 17 December 2021.
  4. Louys, Julien; Curnoe, Darren; Tong, Haowen (January 2007). "Characteristics of Pleistocene megafauna extinctions in Southeast Asia" (PDF). Palaeogeography, Palaeoclimatology, Palaeoecology. 243 (1–2): 152–173. Bibcode:2007PPP...243..152L. doi:10.1016/j.palaeo.2006.07.011.
  5. http://www2.ihp.sinica.edu.tw/file/3111QQPWEMU.pdf Archived 2021-02-25 at the Wayback Machine [ bare URL PDF ]
  6. Chazine, J.-M. (2005). "Rock art, burials, and habitations: Caves in East Kalimantan" (PDF). Asian Perspectives. 44 (1): 219–230. doi:10.1353/asi.2005.0006. hdl: 10125/17232 . S2CID   53372873.
  7. Tong, Haowen (January 2005). "Dental characters of the Quaternary tapirs in China, their significance in classification and phylogenetic assessment". Geobios. 38 (1): 139–150. Bibcode:2005Geobi..38..139T. doi:10.1016/j.geobios.2003.07.006.
  8. Turvey, Samuel T.; Tong, Haowen; Stuart, Anthony J.; Lister, Adrian M. (September 2013). "Holocene survival of Late Pleistocene megafauna in China: a critical review of the evidence". Quaternary Science Reviews. 76: 156–166. Bibcode:2013QSRv...76..156T. doi:10.1016/j.quascirev.2013.06.030.
  9. Maclaren, Jamie A; Hulbert, Richard C; Wallace, Steven C; Nauwelaerts, Sandra (2018-10-05). "A morphometric analysis of the forelimb in the genus Tapirus (Perissodactyla: Tapiridae) reveals influences of habitat, phylogeny and size through time and across geographical space". Zoological Journal of the Linnean Society. 184 (2): 499–515. doi:10.1093/zoolinnean/zly019.
  10. 1 2 Hooijer, D. A. (1947). On fossil and prehistoric remains of Tapirus from Java, Sumatra and China. Zoologische Mededelingen, 27(3), 253–299.