Haringtonhippus

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Haringtonhippus
Temporal range: ~3–0.011  Ma
Haringtonhippus skulls.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Genus: Haringtonhippus
Heintzman et al., 2017
Species:
H. francisci
Binomial name
Haringtonhippus francisci
Hay, 1915 [1]
Synonyms
  • Equus francisciHay, 1915
  • Equus achates
  • Equus quinni
  • Equus cedralensis

Haringtonhippus is an extinct genus of equine from the Pleistocene of North America [2] The genus is monospecific, consisting of the species H. francisci, initially described in 1915 by Oliver Perry Hay as Equus francisci. Members of the genus are often referred to as stilt-legged horses, in reference to their slender distal limb bones, in contrast with those of contemporary "stout legged" caballine true horses. [3]

Contents

Haringtonhippus fossils have only been discovered in North America. [3] Specimens have been found from southern Mexico [4] to southern South Dakota and in Alberta, Canada, [2] at sites such as Gypsum Cave and Natural Trap Cave, as well as eastern Beringia in Yukon [3] A later study found that Equus cedralensis from the Late Pleistocene of Mexico also belonged to this species. [5] The earliest species of the lineage appeared in North America during the Late Pliocene to Early Pleistocene, around 2 to 3 Ma. [3] [6] It became extinct at the end of the Late Pleistocene, around 12,000 years ago as part of the end-Pleistocene extinctions, along with most other large mammals in the Americas. [3]

Taxonomy

Haringtonhippus is named after Charles Richard Harington. [3] It was originally described as a new Equus species, E. francisci, in 1915. [1] Dalquest (1979) considered Equus tauOwen, 1869, described from teeth in Mexico, a senior synonym of E. francisci, [7] while Equus quinni and E. arrelanoi were synonymized with E. francisci by Winans (1989). [8] The species Equus achates Hay and Cook, 1930 (synonymized with E. tau by Dalquest 1979) was synonymized with E. francisci by Hulbert (1995), who also declared E. tau and E. littoralis nomina dubia. [2] [9]

A 2017 paper placed Equus francisci outside of the crown group containing all living member of the genus Equus based on a phylogenetic analysis of DNA sequences, leading to erection of the new genus Haringtonhippus. The genus is phylogenetically closer to Equus than to Hippidion . It is estimated to have diverged from modern Equus around 4.1–5.7 million years ago, during the late Hemphillian or early Blancan. [3]

Equini

Hippidion

Haringtonhippus

Genus  Equus

Equus ferus (wild horse)

Equus ovodovi

Subgenus Asinus (asses)

Subgenus Hippotigris (zebras)

Some other authors have argued that the species should be retained in the genus Equus, due to its morphological similarity to members of that genus. [10]

Description

H. francisci was relatively small-sized, with two studies giving estimated body mass ranges of 167 to 251 kilograms (368 to 553 lb) [4] and 175.5–278.4 kilograms (387–614 lb). [11] The third metatarsal (metapodial) bones are slender [11] and similar to those of asses. [3]

Ecology

H. francisci is thought to have had a predominantly grazing based diet, similar to living equines. [4] Dental mesowear analysis of specimens from Natural Trap Cave in Wyoming suggest that the species regularly consumed abrasive vegetation. [12]

Related Research Articles

<span class="mw-page-title-main">Equidae</span> Family of hoofed mammals

Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families.

<i>Equus</i> (genus) Genus of mammals which includes horses, donkeys, and zebras

Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within the Equidae, Equus is the only recognized extant genus, comprising seven living species. Like Equidae more broadly, Equus has numerous extinct species known only from fossils. The genus most likely originated in North America and spread quickly to the Old World. Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes, and long tails. All species are herbivorous, and mostly grazers, with simpler digestive systems than ruminants but able to subsist on lower-quality vegetation.

<i>Equus scotti</i> Extinct species of mammal

Equus scotti is an extinct species of horse native to Pleistocene North America.

<span class="mw-page-title-main">Wild horse</span> Undomesticated four-footed mammal from the equine family

The wild horse is a species of the genus Equus, which includes as subspecies the modern domesticated horse as well as the endangered Przewalski's horse. The European wild horse, also known as the tarpan, that went extinct in the late 19th or early 20th century has previously been treated as the nominate subspecies of wild horse, Equus ferus ferus, but more recent studies have cast doubt on whether tarpans were truly wild or if they actually were feral horses or hybrids.

<i>Equus lambei</i> Extinct species of horse

Equus lambei, commonly known as the Yukon horse or Yukon wild horse, is an extinct species of the genus Equus. Equus lambei ranged across North America until approximately 10,000 years ago. Based on recent examinations of the mtDNA of Equus lambei remains, scientists have concluded that E. lambei was probably much like the extinct tarpan, also known as the Eurasian wild horse, and the living Przewalski's horse. A partial carcass of Equus lambei is on display at the Yukon Beringia Interpretive Centre in Whitehorse, Yukon.

<i>Equus sivalensis</i> Extinct species of mammal

Equus sivalensis is an extinct species of large equid native to the northern Indian subcontinent. Remains date from the beginning of the Pleistocene, c. 2.58 million years ago until around 600,000 years ago, during the Middle Pleistocene. It is considered a "stenonine horse", meaning that it is more closely related to zebras and asses than true horses. Based on isotopes and teeth morphology, it is thought to have been a grazer. The later species Equus namadicus from the same region has sometimes been suggested to be a synonym due to their similar teeth morphology.

<i>Equus namadicus</i> Extinct species of mammal

Equus namadicus is a prehistoric equid, known from remains dating to the Middle and Late Pleistocene from across the Indian subcontinent, with its last dated records being approximately 29-14,000 years ago. It is considered a "stenonine horse", meaning that it is probably more closely related to zebras and asses than true horses. It is relatively large in size. It is very similar to the earlier Equus sivalensis, also from the Indian subcontinent, from which it only differs in size and in subtle aspects of dental anatomy, and it has sometimes been suggested to be a synonym of it.

<i>Hippidion</i> Extinct genus of mammals

Hippidion is an extinct genus of equine that lived in South America from the Late Pliocene to the end of the Late Pleistocene (Lujanian), between 2.5 million and 11,000 years ago. They were one of two lineages of equines native to South America during the Pleistocene epoch, alongside Equus (Amerhippus) neogeus.

<span class="mw-page-title-main">Evolution of the horse</span>

The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the modern horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.

Plesippus is a genus of extinct horse from the Pleistocene of North America. Although commonly seen as a subgenus of Equus recent cladistic analysis considers it a distinct genus.

The Yunnan horse is an extinct species of equine that was present in East Asia during the Pleistocene very likely as a grazer on open tracts of grassland. It was a small equine comparable in size to the modern Przewalski's horse.

<span class="mw-page-title-main">Equinae</span> Subfamily of mammals

Equinae is a subfamily of the family Equidae, known from the Hemingfordian stage of the Early Miocene onwards. They originated in North America, before dispersing to every continent except Australia and Antarctica. They are thought to be a monophyletic grouping. Members of the subfamily are referred to as equines; the only extant equines are the horses, asses, and zebras of the genus Equus, with two other genera Haringtonhippus and Hippidion becoming extinct at the beginning of the Holocene, around 11-12,000 years ago.

<i>Equus conversidens</i> Extinct species of mammal

Equus conversidens, or the Mexican horse, was a Pleistocene species of horse, now extinct, that inhabited North America.

Equus alaskae was a Pleistocene species of horse, now extinct, that inhabited North America.

Equus semplicatus was a Pleistocene species of New World stilt-legged horse, and considered the type species for the stilt legged horses, one of three lineages of equids within the Americas, the other two being hippidionid and caballine horses. Now extinct, Equus semiplicatus once inhabited North America.

<i>Equus neogeus</i> Extinct species of equine native to South America

Equus neogeus is an extinct species of equine native to South America during the Pleistocene. It was formerly thought to be several distinct species within the subgenus Amerhippus, but was later shown to be a single morphologically variable species. It is thought to be closely related to true horses.

<i>Equus occidentalis</i> Extinct species of horse

Equus occidentalis is an extinct species of wild horse that once inhabited North America, specifically the Southwestern United States, during the Pleistocene epoch. It was first described from three teeth with insufficient diagnostic characters, one even being suggested to be a separate taxon related to the American Zebra, leading some researchers to consider it a nomen dubium, though this taxonomic debate is yet to be fully resolved.

<span class="mw-page-title-main">Horses in the United States</span>

Horses have been an important component of American life and culture since before the founding of the nation. In 2008, there were an estimated 9.2 million horses in the United States, with 4.6 million citizens involved in businesses related to horses. There are an estimated 82,000 feral horses that roam freely in the wild in certain parts of the country, mostly in the Western United States.

<i>Equus stenonis</i> Extinct species of mammal

Equus stenonis is an extinct species of equine that lived in Western Eurasia during the Early Pleistocene epoch.

Equus niobrarensis is an extinct species of Equus, the genus that includes the horse. E. niobrarensis may be synonymous with Equus scotti. It was "stout-legged" and belonged to the "big horses" category as defined by M. C. Winans. The skull of the horse was noted as being broader than Equus caballus.

References

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  2. 1 2 3 "†Equus francisci Hay 1915 (horse)". Fossilworks . Gateway to the Paleobiology Database . Retrieved 29 November 2017.
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  5. Jiménez-Hidalgo, E.; Díaz-Sibaja, R. (2020). "Was Equus cedralensis a Non-Stilt Legged Horse? Taxonomical Implications For the Mexican Pleistocene Horses". Ameghiniana. 57 (3): 284. doi:10.5710/amgh.06.01.2020.3262. ISSN   0002-7014. S2CID   213232576.
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  7. Dalquest, Walter W. (1979). "The Little Horses (Genus Equus) of the Pleistocene of North America". The American Midland Naturalist. 101 (1): 241–244. doi:10.2307/2424920. JSTOR   2424920.
  8. Winans, M. C. 1989. A quantitative study of the North American fossil species of the genus Equus. Pp. 262-297, in The evolution of perissodactyls (D. R. Prothero & R. M. Schoch, eds.), Oxford Monographs Geol. Geophysics, no. 15, 537 pp.
  9. Hulbert, R. C. 1995. Equus from Leisey Shell Pit 1A and other Irvingtonian localities from Florida. Bulletin of the Florida Museum of Natural History 37:553—602.
  10. Barrón-Ortiz, Christina I.; Avilla2, Leonardo S.; Jass, Christopher N.; Bravo-Cuevas, Víctor M.; Machado, Helena; Mothé, Dimila (12 September 2019). "What Is Equus? Reconciling Taxonomy and Phylogenetic Analyses". Frontiers in Ecology and Evolution. 7. doi: 10.3389/fevo.2019.00343 .{{cite journal}}: CS1 maint: numeric names: authors list (link)
  11. 1 2 Landry, Zoe; Roloson, Mathew J.; Fraser, Danielle (December 2022). "Investigating the reliability of metapodials as taxonomic Indicators for Beringian horses". Journal of Mammalian Evolution. 29 (4): 863–875. doi:10.1007/s10914-022-09626-4. ISSN   1064-7554. PMC   9684255 . PMID   36438779.
  12. Spencer, Lillian M.; Scott, Eric (February 2023). "Resource partitioning among late Pleistocene herbivores of Natural Trap Cave, Wyoming". Quaternary International. 647–648: 88–93. Bibcode:2023QuInt.647...88S. doi: 10.1016/j.quaint.2022.05.019 .