Kannemeyeria

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Kannemeyeria
Temporal range: Anisian
~247–242  Ma
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Kannemeyeria erithrea.JPG
Skull of Kannemeyeria simocephalus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Suborder: Anomodontia
Clade: Dicynodontia
Family: Kannemeyeriidae
Genus: Kannemeyeria
Seeley, 1908
Type species
Kannemeyeria proboscoides
Seeley, 1908
[ Junior synonym of K. simocephalus]
Species
  • K. aganosteusKammerer & Ordoñez, 2021
  • K. lophorhinusRenaut et al., 2003
  • K. simocephalus (Weithofer, 1888)
Synonyms
Genus synonymy
Species synonymy
  • Synonyms of K. simocephalus:
    • Dicynodon latifronsBroom, 1889
    • Kannemeyeria erithreaHaughton, 1915
    • K. proboscoidesSeeley, 1909
    • K. wilsoniCruickshank, 1970
    • Proplacerias vanhoepeni(Camp, 1956)
    • ?Ptychocynodon pentangulatusSeeley, 1904 [ nomen nudum?]
    • Sagecephalus pachyrhynchusJaekel, 1911

Kannemeyeria is a genus of dicynodont that lived during the Anisian age of Middle Triassic period in what is now Africa and South America. [1] The generic name is given in honor of Daniel Rossouw Kannemeyer, the South African fossil collector who discovered the original specimen. [2] It is one of the first representatives of the family, and hence one of the first large herbivores of the Triassic.

Contents

Description

Restoration of Kannemeyeria Kannemeyeria DB.jpg
Restoration of Kannemeyeria

Kannemeyeria was about 3 metres (9.8 ft) in length, about the size of an ox. Although it had a large head, it was lightweight due to the size of the eye sockets and nasal cavity. It also had limb girdles which formed massive plates of bone that helped support its heavily built body. [3]

Kannemeyeria was well-adapted to living as a herbivore; it had a powerful beak and strong jaw muscles built for shearing plant material. [3] Kannemeyeria had a massive head with unusually large openings for the eyes, nostrils and jaw muscles. It evidently tore up roots, stripped leaves from the vegetation with its horny beak and ground them up with its toothless jaws. [4]

In a zone of Karoo Supergroup, Kannemeyeria is found alongside large carnivorous archosaur Erythrosuchus . [5]

Distribution

Kannemeyeria skull Kannemeyeria skull.JPG
Kannemeyeria skull

Kannemeyeria is known from the Subzone B of Burgersdorp Formation of South Africa, the Ntawere Formation of Zambia, the Omingonde Formation of Namibia, the Lifua member of Manda Formation of Tanzania, and the Quebrada de los Fósiles Formation of Argentina. [6] [7]

Classification

Life reconstruction of K. simocephalus Kannemeyeria NT small.jpg
Life reconstruction of K. simocephalus

Below is a cladogram from Kammerer et al. (2013): [8]

Kannemeyeriiformes

See also

Related Research Articles

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Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers.

<i>Robertia</i> Extinct genus of dicynodonts

Robertia is an extinct genus of small herbivorous dicynodonts from the Middle to Late Permian of South Africa, between 260 and 265 million years ago. It is a monospecific genus, consisting of the type-species R. broomiana, which was classified by Lieuwe Dirk Boonstra in 1948 and named in honor of Robert Broom for his study of South African mammal-like reptiles.

<i>Diictodon</i> Extinct genus of dicynodonts

Diictodon is an extinct genus of pylaecephalid dicynodont that lived during the Late Permian period, approximately 255 million years ago. Fossils have been found in the Cistecephalus Assemblage Zone of the Madumabisa Mudstone of the Luangwa Basin in Zambia and the Tropidostoma Assemblage Zone of the Teekloof Formation, Tapinocephalus Assemblage Zone of the Abrahamskraal Formation, Dicynodon Assemblage Zone of the Balfour Formation, Cistecephalus Assemblage Zone of the Middleton or Balfour Formation of South Africa and the Guodikeng Formation of China. Roughly half of all Permian vertebrate specimens found in South Africa are those of Diictodon. This small herbivorous animal was one of the most successful synapsids in the Permian period.

<i>Sinokannemeyeria</i> Extinct genus of dicynodonts

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<i>Emydops</i> Extinct genus of dicynodonts

Emydops is an extinct genus of dicynodont therapsids from the Middle Permian to Late Permian of what is now South Africa. The genus is generally small and herbivorous, sharing the dicynodont synapomorphy of bearing two tusks. In the following years, the genus grew to include fourteen species. Many of these species were erected on the basis of differences in the teeth and the positioning of the frontal and parietal bones. A 2008 study narrowed Emydops down to two species, E. arctatus and the newly described E. oweni.

<i>Cynognathus</i> Assemblage Zone Biozone which correlates to the Burgersdorp Formation of the Beaufort Group

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<i>Stahleckeria</i> Extinct genus of dicynodonts

Stahleckeria is an extinct genus of Middle Triassic (Ladinian) dicynodonts. It lived about 240 million years ago in what is now Brazil and Namibia. As a member of the group Kannemeyeriiformes, it was similar to the genus Kannemeyeria. The genus is known from the type species Stahleckeria potens, which was first collected from the Ladinian-age Santa Maria Formation in the Paleorrota fossil site of Brazil. Stahleckeria was named in honor of Rudolf Stahlecker, who discovered the first specimens during a 1935 expedition led by paleontologist Friedrich von Huene to the Chiniquá fossil site.

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<i>Myosaurus</i> Extinct genus of dicynodont from the lower Triassic

Myosaurus is a genus of Anomodontia in the order Therapsida. They are also classified as Dicynodontia, which is a subclade of Anomodontia. The Mysosaurus was a small, herbivorous synapsid that existed around the early Triassic period. All of the fossils found of this species were found in Antarctica and South Africa. Compared to other fossils found from species that existed during this time, the Myosaurus is not common in the fossil record. This is due to a shortage of discovered fossils that possess characteristics unique to the Myosaurus. Notably, under 130 fossil fragments have been found that have been classified as Myosauridae, and almost all have been skulls. These skulls can be classified as Myosaurus because this species, unlike other dicynodonts, do not possess tusks or postfrontal teeth. The only species identified in the family Myosauridae is the Myosaurus gracilis, or M. gracilis. It should be recognized that the Myosaurus is almost always referred to as the M. gracilis in scientific research.

Angonisaurus is an extinct genus of kannemeyeriiform dicynodont from the Middle Triassic of Africa between 247 and 242 million years ago. Only one species, Angonisaurus cruickshanki has been assigned to this genus. This genus is thought to have been widely spread but rare in southern Gondwana. Though few in number, the fossil record of Angonisaurus cruickshanki contains multiple specimens giving it a measurable stratigraphic range. Sexually dimorphic features are found in Angonisaurus which include presence or absence of tusks and difference is size and robustness of the temporal arch and the rostral.

<i>Dicynodontoides</i> Extinct genus of dicynodonts

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<i>Eosimops</i> Extinct genus of dicynodonts

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<i>Rastodon</i> Extinct genus of dicynodonts

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<i>Pentasaurus</i> Genus of dicynodont therapsid from the late Triassic of South Africa

Pentasaurus is an extinct genus of dicynodont of the family Stahleckeriidae, closely related to the well known Placerias. It was found in the Lower Elliot Formation of South Africa, dated to the Norian of the Late Triassic period. The genus contains the type and only species, Pentasaurus goggai. Pentasaurus is named after the ichnogenus Pentasauropus, fossil footprints that were originally described from the lower Elliot Formation in 1970 decades before the body fossils of Pentasaurus itself were recognised. Pentasauropus footprints were likely made by dicynodonts, and in South Africa Pentasaurus itself was the likely trackmaker. The name reflects the fact that a large dicynodont was predicted to have existed in the lower Elliot Formation before any body fossils were recognised, and so Pentasaurus was named after its probable footprints. This is a reversal of the more typical occurrence where fossil footprints are named after their presumed trackmakers. The name of the species honours its collector Alfred Brown, nicknamed "Gogga", which means "bug" in Afrikaans.

<i>Thliptosaurus</i> Extinct genus of dicynodonts

Thliptosaurus is an extinct genus of small kingoriid dicynodont from the latest Permian period of the Karoo Basin in KwaZulu-Natal, South Africa. It contains the type and only known species T. imperforatus. Thliptosaurus is from the upper Daptocephalus Assemblage Zone, making it one of the youngest Permian dicynodonts known, living just prior to the Permian mass extinction. It also represents one of the few small bodied dicynodonts to exist at this time, when most other dicynodonts had large body sizes and many small dicynodonts had gone extinct. The unexpected discovery of Thliptosaurus in a region of the Karoo outside of the historically sampled localities suggests that it may have been part of an endemic local fauna not found in these historic sites. Such under-sampled localities may contain 'hidden diversities' of Permian faunas that are unknown from traditional samples. Thliptosaurus is also unusual for dicynodonts as it lacks a pineal foramen, suggesting that it played a much less important role in thermoregulation than it did for other dicynodonts.

<i>Ufudocyclops</i> Extinct genus of dicynodonts

Ufudocyclops is an extinct genus of stahleckeriid dicynodont from the Middle Triassic of South Africa. It was found in the Burgersdorp Formation, part of the uppermost Cynognathus Assemblage Zone of the Beaufort Group in the Karoo Basin. The type and only known species is U. mukanelai. It was a large, beaked herbivore like other Triassic dicynodonts, lacking tusks, and is mostly characterised by unique features of the skull. It is known from three specimens, two of which were previously referred to the Tanzanian dicynodont Angonisaurus. The separation of Ufudocyclops from Angonisaurus indicates that the Middle Triassic fauna of the Beaufort Group in South Africa was not part of a larger shared fauna with those of the Manda Beds in Tanzania, as was previously supposed, and suggests that they were separated as more localised faunas, possibly by geographic barriers or in time. Ufudocyclops then would have been a unique part of the uppermost Cynognathus Assemblage Zone in South Africa. It is also the oldest known member of the family Stahleckeriidae, and implies that the family was already diversifying in the Middle Triassic alongside other kannemeyeriiforms, not just in the Late Triassic after other families died out.

<i>Acratophorus argentinensis</i> Extinct species of dicynodont

Acratophorus is an extinct genus of dicynodont that lived during the Anisian age of the Middle Triassic-aged Río Seco de la Quebrada Formation in what is now Argentina. The type species, A. argentinensis, was originally placed in the genus Kannemeyeria by Jose Bonaparte in 1966, and later sometimes referred to Vinceria, before being transferred to a new, distinct genus, Acratophorus, in 2021 by Christian Kammerer and Angi Ordoñez. The species Vinceria vieja was also made a synonym of A. argentinensis in 2021. The holotype is PVL 3645, a partial skeleton discovered near a farm house in Puesto Viejo.

References

  1. "Kannemeyeria". Fossilworks. Retrieved 17 December 2021.
  2. "Kannemeyeria". Paleofile. Retrieved 12 March 2020.
  3. 1 2 Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 191. ISBN   1-84028-152-9.
  4. Cloudsley-Thompson, John L. (2005). Ecology and Behaviour of Mesozoic Reptiles (2005 ed.). Springer. pp. 113–114. ISBN   978-3540224211 . Retrieved 12 March 2020.
  5. Hancox, P.J.; Neveling, J.; Rubidge, B.S. (2020-06-01). "Biostratigraphy of the Cynognathus Assemblage Zone (Beaufort Group, Karoo Supergroup), South Africa". South African Journal of Geology. 123 (2): 217–238. Bibcode:2020SAJG..123..217H. doi:10.25131/sajg.123.0016. ISSN   1012-0750. S2CID   225828531.
  6. Chinsamy-Turan, Anusuya (2011). Forerunners of Mammals: Radiation • Histology • Biology. Indiana University Press. p. 53. ISBN   978-0-253-00533-5 . Retrieved 12 March 2020.
  7. Kammerer, C. F.; Ordoñez, M. D. (2021). "Dicynodonts (Therapsida: Anomodontia) of South America". Journal of South American Earth Sciences . 108: 103171. Bibcode:2021JSAES.10803171K. doi:10.1016/j.jsames.2021.103171. S2CID   233565963.
  8. Kammerer, C. F.; Fröbisch, J. R.; Angielczyk, K. D. (2013). Farke, Andrew A (ed.). "On the Validity and Phylogenetic Position of Eubrachiosaurus browni, a Kannemeyeriiform Dicynodont (Anomodontia) from Triassic North America". PLOS ONE. 8 (5): e64203. Bibcode:2013PLoSO...864203K. doi: 10.1371/journal.pone.0064203 . PMC   3669350 . PMID   23741307.