Loricatosaurus Temporal range: Middle Jurassic, | |
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Fossils originally referred to Lexovisaurus, Muséum national d'histoire naturelle, Paris. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Thyreophora |
Clade: | † Stegosauria |
Family: | † Stegosauridae |
Genus: | † Loricatosaurus Maidment et al., 2008 |
Type species | |
†Loricatosaurus priscus Nopcsa, 1911 | |
Synonyms | |
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Loricatosaurus (meaning "armored lizard") is a Stegosaurid genus from Callovian-age (Middle Jurassic) rocks of England and France. [1]
Loricatosaurus is known from remains previously assigned to Lexovisaurus that were first discovered at the Fletton brick pit in the Oxford Clay Formation, Fletton, Peterborough, England, by Alfred Leeds during November 1901. [2] Leeds collected a semi articulated skeleton consisting of two cervical, six dorsal, and 16 caudal vertebrae, a partial right forelimb, partial left hindlimb, partial pelves, and some fragmentary dermal armor. One piece of armor was described as a parascapular spine, but it is actually a fragmentary tail spine.
In 1957, another specimen now referred to Loricatosaurus was briefly referred to Lexovisaurus until a more proper description was made in 1985 by Peter Galton. The specimen was a partial postcranial skeleton, including many vertebrae and a dorsal spine, from the Lower Callovian of Le Fresne d'Argences (Calvados), Normandy, France.
The material was later named Stegosaurus priscus (“ancient roofed lizard”) by Baron Franz Nopcsa in 1911 and Lexovisaurus by Hoffstetter, 1957 and Peter Galton, 1985. However, upon later review of the material in 2008, Susannah Maidment and colleagues determined that Lexovisaurus was based upon nondiagnostic remains, and coined a new genus for Lexovisaurus priscus and its referred specimen. The genus name Loricatosaurus derives from the Latin "loricatus" (meaning armoured) and the Greek "sauros" (meaning lizard).
It is known from only two partial skeletons. Contrary to previous reports (as Lexovisaurus), a shoulder spine is not present in the known material; the possible spine is more likely from the tail. [1]
The holotype of L. priscus was digitally scanned in January 2023 at the Natural History Museum in London.
The holotype (NHMUK R3167) is a partial skeleton, including two cervical, six dorsal, and 16 caudal vertebrae, a right humerus and right ulna, some carpus, two metacarpals (one incomplete), a partial ilia, partial right ischium, and pubis, the left femur, partial tibia and fibula, fused tarsals, and some armour plates. This material now resides in the British Museum of Natural History's palaeontology collection. Another referred specimen (MHBR 001), from an unnamed unit referred to as the Marnes a Belemnopsis latesulcatus Formation from the lower Callovian, Middle Jurassic of Le Fresne d’Argences, Calvados, Normandy, France, includes neck, back and tail vertebrae, a left upper arm, right thigh, shin and calf, and some armour plates. [1]
Multiple phylogenetic analyses carried out by S. Maidment and O. Mateus between 2008 and 2010 place Loricatosaurus as a basal genus within the family Stegosauridae. [1] [3] [4]
Stegosaurus is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been found in the western United States and in Portugal, where they are found in Kimmeridgian- to Tithonian-aged strata, dating to between 155 and 145 million years ago. Of the species that have been classified in the upper Morrison Formation of the western US, only three are universally recognized: S. stenops, S. ungulatus and S. sulcatus. The remains of over 80 individual animals of this genus have been found. Stegosaurus would have lived alongside dinosaurs such as Apatosaurus, Diplodocus, Camarasaurus and Allosaurus, the latter of which may have preyed on it.
Kentrosaurus is a genus of stegosaurid dinosaur from the Late Jurassic in Lindi Region of Tanzania. The type species is K. aethiopicus, named and described by German palaeontologist Edwin Hennig in 1915. Often thought to be a "primitive" member of the Stegosauria, several recent cladistic analyses find it as more derived than many other stegosaurs, and a close relative of Stegosaurus from the North American Morrison Formation within the Stegosauridae.
Hesperosaurus is a herbivorous stegosaurian dinosaur from the Kimmeridgian age of the Jurassic period, approximately 156 million years ago.
Dacentrurus, originally known as Omosaurus, is a genus of stegosaurian dinosaur from the Late Jurassic to Early Cretaceous of Europe. Its type species, Omosaurus armatus, was named in 1875, based on a skeleton found in a clay pit in the Kimmeridge Clay in Swindon, England. In 1902 the genus was renamed Dacentrurus because the name Omosaurus had already been used for a crocodylian. After 1875, half a dozen other species would be named but perhaps only Dacentrurus armatus is valid. Finds of this animal have been limited and much of its appearance is uncertain. It was a heavily built quadrupedal herbivore, adorned with plates and spikes, reaching 8–9 metres (26–30 ft) in length and 5 metric tons in body mass.
Chialingosaurus is a genus of herbivorous stegosaurian dinosaur similar to Kentrosaurus from the Upper Shaximiao Formation, Late Jurassic beds in Sichuan Province in China. Its age makes it one of the oldest species of stegosaurs, living about 160 million years ago. Since it was an herbivore, scientists think that Chialingosaurus probably ate ferns and cycads, which were plentiful during the period when Chialingosaurus was alive.
Wuerhosaurus is a genus of stegosaurid dinosaur from the Early Cretaceous Period of China and Mongolia. As such, it was one of the last genera of stegosaurians known to have existed.
Huayangosaurus is a genus of stegosaurian dinosaur from the Middle Jurassic of China. The name derives from "Huayang" (華陽), an alternate name for Sichuan, and "saurus", meaning "lizard". It lived during the Bathonian to Callovian stages, around 165 million years ago, some 20 million years before its famous relative, Stegosaurus appeared in North America. At only approximately 4 metres (13 ft) long, it was also much smaller than its famous cousin. Found in the Lower Shaximiao Formation, Huayangosaurus shared the local Middle Jurassic landscape with the sauropods Shunosaurus, Datousaurus, Omeisaurus and Protognathosaurus, the ornithopod Xiaosaurus and the carnivorous Gasosaurus.
Lexovisaurus is a genus of stegosaur from mid-to-Late Jurassic Europe, 165.7-164.7 mya. Fossils of limb bones and armor fragments have been found in middle to late Jurassic-aged strata of England and France.
Tuojiangosaurus is a genus of herbivorous stegosaurian dinosaur from the Late Jurassic Period, recovered from the Upper Shaximiao Formation of what is now Sichuan Province in China.
Paranthodon is a genus of stegosaurian dinosaur that lived in what is now South Africa during the Early Cretaceous, between 139 and 131 million years ago. Discovered in 1845, it was one of the first stegosaurians found. Its only remains, a partial skull, isolated teeth, and fragments of vertebrae, were found in the Kirkwood Formation. British paleontologist Richard Owen initially identified the fragments as those of the pareiasaur Anthodon. After remaining untouched for years in the British Museum of Natural History, the partial skull was identified by South African paleontologist Robert Broom as belonging to a different genus; he named the specimen Palaeoscincus africanus. Several years later, Hungarian paleontologist Franz Nopcsa, unaware of Broom's new name, similarly concluded that it represented a new taxon, and named it Paranthodon owenii. Since Nopcsa's species name was assigned after Broom's, and Broom did not assign a new genus, both names are now synonyms of the current binomial, Paranthodon africanus. The genus name combines the Ancient Greek para (near) with the genus name Anthodon, to represent the initial referral of the remains.
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, Africa, South America and Asia. Their geographical origins are unclear; the earliest unequivocal stegosaurian, Huayangosaurus taibaii, lived in China.
Stegosauridae is a family of thyreophoran dinosaurs within the suborder Stegosauria. The clade is defined as all species of dinosaurs more closely related to Stegosaurus than Huayangosaurus. The name ‘Stegosauridae’ is thus a stem-based name taken from the well-represented genus – Stegosaurus. Fossil evidence of stegosaurids, dating from the Middle Jurassic through the Early Cretaceous, have been recovered from North America, Eurasia and Africa.
Chungkingosaurus, meaning "Chongqing Lizard", is a genus of herbivorous dinosaur from the Late Jurassic Upper Shaximiao Formation in what is now China. It is a member of the Stegosauria.
Gigantspinosaurus is a genus of herbivorous ornithischian dinosaur from the Late Jurassic. It was a stegosaur found in China.
Yingshanosaurus is a genus of stegosaurian dinosaur from the Late Jurassic, around 155 million years ago. It was a herbivore that lived in what is now China. The type species is Yingshanosaurus jichuanensis.
Hypsirhophus is a genus of stegosaurian dinosaurs. It contains a single species, Hypsirhophus discurus, which is known only from a fragmentary specimen. The fossil consists of partial vertebrae from the back, three from the tail, and a piece of rib.
Miragaia is a long-necked stegosaurid dinosaur. Its fossils have been found in Upper Jurassic rocks in Portugal and possibly also Wyoming, United States. Miragaia has the longest neck known for any stegosaurian, which included at least seventeen vertebrae.
Alcovasaurus, alternatively known as Miragaia longispinus, is a genus of herbivorous stegosaurian dinosaur that lived in the Late Jurassic. It was found in the Morrison Formation of Natrona County, Wyoming, United States. The type species is Stegosaurus longispinus, later given the genus Alcovasaurus, and in 2019 recombined as Miragaia longispinus.
This timeline of stegosaur research is a chronological listing of events in the history of paleontology focused on the stegosaurs, the iconic plate-backed, spike-tailed herbivorous eurypod dinosaurs that predominated during the Jurassic period. The first scientifically documented stegosaur remains were recovered from Early Cretaceous strata in England during the mid-19th century. However, they would not be recognized as a distinct group of dinosaurs until Othniel Charles Marsh described the new genus and species Stegosaurus armatus in 1877, which he regarded as the founding member of the Stegosauria. This new taxon originally included all armored dinosaurs. It was not until 1927 that Alfred Sherwood Romer implemented the modern use of the name Stegosauria as specifically pertaining to the plate-backed and spike-tailed dinosaurs.
Isaberrysaura is a genus of stegosaurian ornithischian dinosaur from the Middle Jurassic Los Molles Formation of Patagonia, Argentina. The genus contains a single species, I. mollensis, described by Salgado et al. in 2017 from a single specimen. Although initially classified as a basal neornithischian, subsequent analysis has allied it with the Stegosauria; the morphology of its skull resembles those of other members of the group.