| Lycopodium clavatum | |
|---|---|
 | |
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Lycophytes |
| Class: | Lycopodiopsida |
| Order: | Lycopodiales |
| Family: | Lycopodiaceae |
| Genus: | Lycopodium |
| Species: | L. clavatum |
| Binomial name | |
| Lycopodium clavatum | |
| Synonyms [1] [2] | |
Synonymy
| |
Lycopodium clavatum (common club moss, [3] [4] stag's-horn clubmoss, [5] running clubmoss, [6] or ground pine [7] ) is the most widespread species in the genus Lycopodium in the clubmoss family.
Lycopodium clavatum is a spore-bearing vascular plant, growing mainly prostrate along the ground with stems up to 1 m (39 in) long; the stems are much branched, and densely clothed with small, spirally arranged microphyll leaves. The leaves are 3–5 mm long and 0.7–1 mm broad, tapered to a fine hair-like white point. The branches bearing strobili or spore cones turn erect, reaching 5–15 cm (2.0–5.9 in) above ground, and their leaves are modified as sporophylls that enclose the spore capsules or sporangia. The spore cones are yellow-green, 2–3 cm (0.79–1.18 in) long, and 5 mm (0.20 in) broad. The horizontal stems produce roots at frequent intervals along their length, allowing the stem to grow indefinitely along the ground. The stems superficially resemble small seedlings of coniferous trees, though it is not related to these.
Lycopodium clavatum has a widespread distribution across several continents. [8] [9] [10] [11] [12] [13] There are distinct subspecies and varieties in different parts of its range:
Although globally widespread, like many clubmosses, it is confined to undisturbed sites, disappearing from farmed areas and sites with regular burning. As a result, it is endangered in many areas. In the UK it is one of 101 species named as a high priority for conservation by the wild plant charity Plantlife.
Common names for this species include wolf's-foot clubmoss, common clubmoss, wolf-paw clubmoss, running ground-pine, running pine, [7] running moss, princess pine, and others.
The spores of this moss, "lycopodium powder", are explosive if present in high density air. They were used as flash powder in early photography and magic acts.
The plant has been used in folk science as a diuretic and, like Huperzia selago, as a remedy for rickets. [14]
Bioactive secondary metabolites in clubmosses include triterpenoids with acetylcholinesterase inhibitor activity isolated from this species. [15]
The Lycopodiaceae are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 16 accepted genera and about 400 known species. This family originated about 380 million years ago in the early Devonian, though the diversity within the family has been much more recent. "Wolf foot" is another common name for this family due to the resemblance of either the roots or branch tips to a wolf's paw.
Lycopodium is a genus of clubmosses, also known as ground pines or creeping cedars, in the family Lycopodiaceae. Two very different circumscriptions of the genus are in use. In the Pteridophyte Phylogeny Group classification of 2016, Lycopodium is one of nine genera in the subfamily Lycopodioideae, and has from nine to 15 species. In other classifications, the genus is equivalent to the whole of the subfamily, since it includes all of the other genera. More than 40 species are accepted.
Diphasiastrum is a genus of clubmosses in the plant family Lycopodiaceae. In the Pteridophyte Phylogeny Group classification of 2016, it is placed in the subfamily Lycopodioideae. It is closely related to the genus Lycopodium, and some botanists treat it within a broad view of that genus as a section, Lycopodium sect. Complanata. Some species superficially resemble diminutive gymnosperms and have been given common names such as ground-pine or ground-cedar.
Lycopodiella is a genus in the clubmoss family Lycopodiaceae. The genus members are commonly called bog clubmosses, describing their wetland habitat. The genus has a cosmopolitan distribution, with centers of diversity in the tropical New World and New Guinea. In the past, the genus was often incorporated within the related genus Lycopodium, but was segregated in 1964. In the Pteridophyte Phylogeny Group classification of 2016, Lycopodiella is placed in the subfamily Lycopodielloideae, along with three other genera. In this circumscription, the genus has about 15 species. Other sources use a wider circumscription, in which the genus is equivalent to the Lycopodielloideae of PPG I, in which case about 40 species and hybrids are accepted.
Huperzia selago, the northern firmoss or fir clubmoss, is a vascular plant in the family Lycopodiaceae. It is small-ish, sturdy, stiff and upright and densely scale-leaved. This plant is an evergreen, perennial pteridophyte. The spores are produced June to September. It has a circumpolar distribution.
Phlegmariurus phlegmaria, synonym Huperzia phlegmaria, commonly known as either coarse tassel fern or common tassel fern, is an epiphytic species native to rainforests in Madagascar, some islands in the Indian Ocean, Asia, Australasia and many Pacific Islands. Phlegmariurus phlegmaria is commonly found in moist forests and rainforests at high altitudes, in and amongst mosses and other epiphytes. Members of the order Lycopodiales are commonly referred to as clubmosses.
Diphasiastrum complanatum, common names groundcedar, creeping jenny, or northern running-pine, is a species of clubmoss native to dry coniferous forests in colder northerly parts of the world. Under the original name Lycopodium complanatum, this was an inclusive superspecies that included a number of other species now known to be biologically separate. This plant is an evergreen, perennial pteridophyte. The spores are produced June to September.
Spinulum annotinum, synonym Lycopodium annotinum, known as interrupted club-moss, or stiff clubmoss, is a species of clubmoss native to forests of the colder parts of North America, as well as Asia, and most of Europe. It is an evergreen perennial pteridophyte. The genus Spinulum is accepted in the Pteridophyte Phylogeny Group classification of 2016, but not in other classifications, which submerge the genus in Lycopodium.
Lycopodiella inundata is a species of club moss known by the common names inundated club moss, marsh clubmoss and northern bog club moss. It has a circumpolar and circumboreal distribution, occurring throughout the northern Northern Hemisphere from the Arctic to montane temperate regions in Eurasia and North America. It grows in wet habitat, such as bogs, ponds, moist spots on the tundra, and long-standing borrow pits.
Diphasiastrum digitatum is known as groundcedar, running cedar or crowsfoot, along with other members of its genus, but the common name fan clubmoss can be used to refer to it specifically. It is the most common species of Diphasiastrum in North America. It is a type of plant known as a clubmoss, which is within one of the three main divisions of living vascular plants. It was formerly included in the superspecies Diphasiastrum complanatum. For many years, this species was known as Lycopodium flabelliforme or Lycopodium digitatum.
Dendrolycopodium obscurum, synonym Lycopodium obscurum, commonly called rare clubmoss, ground pine, or princess pine, is a North American species of clubmoss in the family Lycopodiaceae. It is a close relative of other species such as D. dendroideum and D. hickeyi, also treelike. It is native to the eastern United States and southeastern Canada from Georgia to Minnesota to Nova Scotia. It grows in the understory of temperate coniferous and deciduous forests, where it is involved in seral secondary succession, growing in clonal colonies some years after disturbance has occurred. It has also been found in Japan, Taiwan, Korea, Russian Far East, and northeastern China.
Diphasiastrum alpinum, the alpine clubmoss, is a species of clubmoss. This plant is a glaucous scale-leaved perennial pteridophyte. The spores are produced June to September. It was first described by Carl Linnaeus in his Flora Lapponica, 1737, from specimens obtained in Finland.
Diphasiastrum sitchense, the Sitka clubmoss, is a pteridophyte species native to northern North America and northeastern Asia. It is a terrestrial herb spreading by stolons running on the surface or the ground or just slightly below the surface. Leaves are appressed, broadly lanceolate, up to 3.2 mm long. Strobili are solitary on the ends of shoots. It is known from every province in Canada, plus the US States of Alaska, Oregon, Washington, Idaho, Montana, Maine, New Hampshire, Vermont, and New York. It is also found in Greenland, St. Pierre and Miquelon, Yukon, Japan, and the Kamchatka Peninsula of Asiatic Russia. It can be found in alpine meadows, open rocky barrens, and coniferous woodlands.
Diphasiastrum tristachyum, commonly known as blue clubmoss, blue ground-cedar, ground pine, deep-rooted running-pine or ground cedar, is a North American and Eurasian species of clubmoss. In North America, it has been found from Newfoundland west to Manitoba, and south as far as Georgia and Alabama. In Eurasia, it ranges from southern Norway and Sweden south to France and Italy and it also occurs in the Caucasus.
Lycopodium lagopus, commonly known as one-cone club-moss, is an arctic and subarctic species of plants in the genus Lycopodium in the clubmoss family. It is widespread in cold, northerly regions: Canada, Greenland, Russia, Scandinavia, and the northern United States including Alaska.
Dendrolycopodium hickeyi known as Hickey's tree club-moss or Pennsylvania clubmoss, is a North American species of clubmoss in the family Lycopodiaceae. It is native to eastern and Central Canada and the eastern and north-central United States. The genus Dendrolycopodium is accepted in the Pteridophyte Phylogeny Group classification of 2016, but not in other classifications, which submerge the genus in a larger Lycopodium.
Lycopodiella alopecuroides, the foxtail clubmoss, is a species of perennial vascular plant in the club-moss family, Lycopodiaceae. It is commonly found along the Atlantic seaboard and has been recently been discovered in the state of Maine. The family, Lycopodiaceae contains nearly 15 genera and about 375 species
Lycopodioideae is a subfamily in the family Lycopodiaceae in the Pteridophyte Phylogeny Group classification of 2016. It is equivalent to a broad circumscription of the genus Lycopodium in other classifications. Like all lycophytes, members of the Lycopodioideae reproduce by spores. The oldest fossils of modern members of the subfamily date to the Early Cretaceous.
