Macrobaenidae

Last updated

Macrobaenidae
Temporal range: Early Cretaceous–Paleocene
The American journal of science (1904) (18152226245).jpg
Osteopygis emarginatus carapace
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pantestudines
Clade: Testudinata
Clade: Perichelydia
Family: Macrobaenidae
Sukhanov, 1964
Genera

See text

Macrobaenidae is an extinct family of turtles, known from the Early Cretaceous to Paleogene of Laurasia. Their relationships to other turtles and whether they form a monophlyletic group are controversial. They are typically interpreted as stem or crown group cryptodires, but some more recent analyses have found them to lie outside crown group Testudines. Macrobaenids can be distinguished from other testudinatans by the presence of a carotid fenestra, cruciform plastron with strap-like epiplastra, and a lack of extragulars. [1]

Genera

Hongkongochelys from the Middle-Late Jurassic of China has sometimes been attributed to the family, [4] but other times has been attriuted to Sinemydidae, a group which has an unresolved relationship with Macrobaenidae.

Related Research Articles

<span class="mw-page-title-main">Carettochelyidae</span> Family of turtles

Carettochelyidae is a family of cryptodiran turtles belonging to the Trionychia. It contains only a single living species, the pig-nosed turtle native to New Guinea and Northern Australia. Stem-group carettochelyids are known from the Cretaceous of Asia, with the family being widely distributed across North America, Europe and Asia during the Paleogene.

<span class="mw-page-title-main">Chelidae</span> Family of turtles

Chelidae is one of three living families of the turtle suborder Pleurodira, and are commonly called Austro-South American side-neck turtles. The family is distributed in Australia, New Guinea, parts of Indonesia, and throughout most of South America. It is a large family of turtles with a significant fossil history dating back to the Cretaceous. The family is entirely Gondwanan in origin, with no members found outside Gondwana, either in the present day or as a fossil.

<span class="mw-page-title-main">Podocnemididae</span> Family of turtles

Podocnemididae is a family of pleurodire (side-necked) turtles, once widely distributed. Most of its 20 genera and 30 species are now extinct. Seven of its eight surviving species are native to South America: the genus Peltocephalus, with only one species ; and the genus Podocnemis, with six living species of South American side-necked river turtles. There is also one genus native to Madagascar: Erymnochelys, the Madagascan big-headed turtle, whose single species E. madagascariensis.

<span class="mw-page-title-main">Neuquén Group</span> Group of geologic formations in Argentina

The Neuquén Group is a group of geologic formations found in Argentina. Rocks in the Neuquén Group fall within the Cenomanian to early Campanian stages of the Late Cretaceous Period. It overlies the older Lohan Cura Formation and is itself overlain by the younger Allen Formation of the Malargüe Group, separated from both by unconformities, dated to 98 and 79 Ma respectively.

<span class="mw-page-title-main">Bothremydidae</span> Extinct family of turtles

Bothremydidae is an extinct family of side-necked turtles (Pleurodira) known from the Cretaceous and Cenozoic. They are closely related to Podocnemididae, and are amongst the most widely distributed pleurodire groups, with their fossils having been found in Africa, India, the Middle East, Europe, North America and South America. Bothremydids were aquatic turtles with a high morphological diversity, indicative of generalist, molluscivorous, piscivorous and possibly herbivorous grazing diets, with some probably capable of suction feeding. Unlike modern pleurodires, which are exclusively freshwater, bothremydids inhabited freshwater, marine and coastal environments. Their marine habits allowed bothremydids to disperse across oceanic barriers into Europe and North America during the early Late Cretaceous (Cenomanian). The youngest records of the group are indeterminate remains from Saudi Arabia and Oman, dating to the Miocene.

<span class="mw-page-title-main">Paracryptodira</span> Extinct clade of turtles

Paracryptodira is an extinct group of reptiles in the clade Testudinata, known from the Jurassic to Paleogene of North America and Europe. Initially treated as a suborder sister to Cryptodira, they were then thought to be a very primitive lineage inside the Cryptodira according to the most common use of the latter taxon. They are now often regarded as late-diverging stem-turtles, lying outside the clade formed by Cryptodira and Pleurodira. The paracryptodires are said to have phylogenic relationships, noted as primary subclades, within the Baenidae and Pleurosternidae. Within each subclade, lies many biodiverse turtles that are continuously being investigated and added to the fossil record. Paracryptodires are divided into three main groups, Compsemydidae, known from the Late Jurassic to Paleocene of North America and Europe, Pleurosternidae, known from the Late Jurassic to Early Cretaceous of North America and Europe, and Baenidae, known from the Early Cretaceous to Eocene of North America. The latter two groups are more closely related to each other than to Compsemys, forming the clade Baenoidea.

<span class="mw-page-title-main">Berothidae</span> Family of insects

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.

Eoscapherpeton is an extinct genus of giant salamander, known from the Late Cretaceous of Central Asia. Fossils have been found in the Cenomanian aged Khodzhakul Formation and Dzharakuduk Formation, Turonian aged Bissekty Formation and the Coniacian-Santonian aged Aitym Formation of Uzbekistan, the Santonian aged Yalovach Formation of Tajikistan, and the Santonian-lower Campanian aged Bostobe Formation and Campanian aged Darbasa Formation of Kazakhstan.

<span class="mw-page-title-main">Hemiphlebiidae</span> Family of damselflies

Hemiphlebiidae is a family of damselflies. It contains only one extant species, the ancient greenling, native to Southern Australia and Tasmania. The fossil record of the group extends back to the Late Jurassic, making them the oldest known crown group damselflies.

<span class="mw-page-title-main">Mesoraphidiidae</span> Extinct family of insects

Mesoraphidiidae is an extinct family of snakeflies in the suborder Raphidiomorpha. The family lived from the Late Jurassic through the Late Cretaceous and is known from twenty-five genera. Mesoraphidiids have been found as both compression fossils and as inclusions in amber. The family was first proposed in 1925 by the Russian paleoentomologist Andrey Vasilyevich Martynov based on Upper Jurassic fossils recovered in Kazakhstan. The family was expanded in 2002 by the synonymizing of several other proposed snakefly families. The family was divided into three subfamilies and one tribe in a 2011 paper, further clarifying the relationships of the included genera.

<span class="mw-page-title-main">Maimetshidae</span> Extinct family of wasps

Maimetshidae is an extinct family of wasps, known from the Cretaceous period. While originally considered relatives of Megalyridae, they are now considered to probably be close relatives of Trigonalidae.

<span class="mw-page-title-main">Sinemydidae</span> Extinct family of turtles

Sinemydidae is an extinct family of turtles from Cretaceous to Paleocene deposits in Asia and North America. Their exact position is engimatic, they have alternatively been considered stem-group cryptodires, but also "crownward stem-turtles" alongside Macrobaenidae, Paracryptodira, Xinjiangchelyidae, Thalassochelydia and Sandownidae outside of crown Testudines.

<span class="mw-page-title-main">Helochelydridae</span> Extinct family of reptiles

The Helochelydridae are an extinct family of stem-turtles known from fossils found in North America and Europe spanning the Early to Late Cretaceous.

<i>Naomichelys</i> Extinct genus of turtles

Naomichelys is an extinct genus of helochelydrid stem turtle known from the Cretaceous (Aptian-Campanian) of North America. It is the only member of the family known to be native to North America.

Anatolemys is an extinct turtle genus in the family Macrobaenidae. Two species are known, both of which lived in the Late Cretaceous. Fossils were discovered in the Yalovach Formation of Tajikistan, the Kulbikin Member and Khodzhakul and Bissekty Formations of Uzbekistan and the Bostobe Formation of Kazakhstan. With 70 cm (2.3 ft) in carapace length, Anatolemys maximus was one of the three largest macrobaenids along with Early Cretaceous Yakemys multiporcata and Paleocene Judithemys backmani.

<span class="mw-page-title-main">Perichelydia</span> Clade of reptiles

Perichelydia is a clade within Pantestudines known from the Middle Jurassic to Holocene. Alongside crown group Testudines, it also contains Helochelydridae, which is known from the Cretaceous of Europe and North America, Sichuanchelyidae from the Middle Jurassic to Paleocene of Asia and Europe, Meiolaniformes, which is known from the Cretaceous to Holocene of South America, Australia and Oceania, and Spoochelys, known from the Mid-Cretaceous Griman Creek Formation of Australia. Kallokibotion from the Late Cretaceous of Europe is also considered part of this group. Several other groups, including the proposed clade Angolachelonia, Paracryptodira, Macrobaenidae, Sinemydidae and Xinjiangchelyidae, which are sometimes considered members of Cryptodira, have also been found outside crown Testudines in several analyses. These groups are usually considered to be closer to the crown group than the other members of Perichelydia.

The Ulaanoosh Formation, formerly Baruunbayan Formation, is a geologic formation in the Ömnögovi Province of southern Mongolia. The formation dates to the Albian to Cenomanian stages of the Cretaceous. The Ulaanoosh Formation has provided fossils of dinosaurs, turtles and dinosaur eggs assigned to Parafaveoloolithus sp.. In 2020, the neoceratopsian Beg tse was described from the alluvial sandstones, mudstones and conglomerates of the formation.

Mesoblattinidae is an extinct family of cockroaches known from the Mesozoic. It was formerly considered a wastebasket taxon for Mesozoic cockroaches, but the family has subsequently been better defined, with many taxa transferred to Caloblattinidae. It is considered to have close affinities with Blattidae and Ectobiidae, as well as possibly Blaberidae. The family first appeared by the Early Jurassic. They are considered to represent amongst the oldest groups of modern cockroaches, and like them are thought to have laid oothecae.

Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae. They were originally considered to be beetles due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces.

<i>Yakemys</i> Extinct genus of turtles

Yakemys is an extinct genus of macrobaenid turtles from the Late Jurassic-Early Cretaceous (Tithonian-Valanginian?) lower Phu Kradung Formation of Thailand. It is monotypic, containing a single species, Yakemys multiporcata.

References

  1. Joyce, Walter G.; Anquetin, Jérémy; Cadena, Edwin-Alberto; Claude, Julien; Danilov, Igor G.; Evers, Serjoscha W.; Ferreira, Gabriel S.; Gentry, Andrew D.; Georgalis, Georgios L.; Lyson, Tyler R.; Pérez-García, Adán (December 2021). "A nomenclature for fossil and living turtles using phylogenetically defined clade names". Swiss Journal of Palaeontology. 140 (1): 5. doi: 10.1186/s13358-020-00211-x . hdl: 11336/155192 . ISSN   1664-2376.
  2. 1 2 3 4 5 6 7 "Macrobaenidae". FossilWorks. Retrieved 17 December 2021.
  3. Gentry, A. D.; Kiernan, C. R.; Parham, J. F. (2022). "A large non-marine turtle from the Upper Cretaceous of Alabama and a review of North American "Macrobaenids"". The Anatomical Record. doi:10.1002/ar.25054.
  4. 1 2 Adán Pérez-García (2020). "A European Cenozoic 'Macrobaenid:' New Data about the Paleocene Arrival of Several Turtle Lineages to Europe". Journal of Vertebrate Paleontology. 40 (4): e1795874. doi:10.1080/02724634.2020.1795874. S2CID   225151817.