Nsungwe Formation

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Nsungwe Formation
Stratigraphic range: Oligocene
~25.2  Ma
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Type Geological formation
Unit ofRed Sandstone Group (Rukwa Rift Basin)
Sub-unitsUtengule Member, Songwe Member
UnderliesUnconformity with Lake Beds Sequence
OverliesUnconformity with Galula Formation
Thickness400 m (1,300 ft)
Lithology
Primary Sandstone
Other Conglomerate, mudstone, siltstone, tuff
Location
Location Rukwa, Mbeya Region
Region African Great Lakes
CountryFlag of Tanzania.svg  Tanzania
Type section
Named for Nsungwe River
Named byRoberts et al.
Year defined2010
Thickness at type section~400 m

The Nsungwe Formation is a formation in the Rukwa Rift Basin of the East African Rift System, it is Oligocene in age based on U-Pb dating of a tuff horizon within the formation. It is part of the Red Sandstone Group along with the uncomfortably underlying Mid-Cretaceous Galula Formation It is divided into two members, the lower Utengule Member, and the upper Songwe member. It is notable for being one of the most important Paleogene fossil deposits in Sub-Saharan Africa.

Contents

Geology

The lithology of the two members are quite different, representing different fluvial environments. The Utengule member is 85 m thick and predominantly consists of red-orange sandstones and matrix to clast supported conglomerates. The overlying Songwe Member is approximately twice as thick as the Utengule member, being 310–320 m thick in the type section. It is much finer grained, consisting of red-orange and grey green claystones, siltstones, mudstones, lenticular sandstones and tuffs. The sediments of the Songwe Member are noted for their fossil content. [1]

Fossil content

Important fossils have come out of the formation, including the rodent Kahawamys , [2] some of the oldest records of the frog genus Xenopus , [3] the hyaenodont Pakakali , [4] the elephant shrews Oligorhynchocyon and Rukwasengi , [5] and the primates Nsungwepithecus and Rukwapithecus , some of the earliest crown catarrhines. [6]

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<i>Galulatherium</i> Fossil taxon

Galulatherium is an extinct genus of possibly gondwanathere mammal, from the Late Cretaceous (Turonian-Campanian)-aged Galula Formation of Tanzania. It is known solely from the type specimen TNM 02067 a fragmentary fossil dentary. The short, deep bone is about 19.5 mm (0.77 in) long, but the back part is broken off. It contains a large, forward-inclined incisor with a root that extends deep into the jaw, separated by a diastema (gap) from five cheekteeth. Very little remains of the teeth, but enough to determine that they are hypsodont (high-crowned). The third cheektooth is the largest and the roots of the teeth are curved. First described in 2003, TNM 02067 has been tentatively identified as a sudamericid—an extinct family of high-crowned gondwanathere mammals otherwise known from South America, Madagascar, India, and Antarctica. If truly a gondwanathere, it would be the only African member of the group and may be the oldest. The describers could not exclude other possibilities, such as that the jaw represents some mammalian group known only from younger, Cenozoic times. In 2019 the fossil was CT scanned, which revealed additional details of the specimen.

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References

  1. Roberts, Eric M.; O’Connor, Patrick M.; Stevens, Nancy J.; Gottfried, Michael D.; Jinnah, Zubair A.; Ngasala, Sifael; Choh, Adeline M.; Armstrong, Richard A. (May 2010). "Sedimentology and depositional environments of the Red Sandstone Group, Rukwa Rift Basin, southwestern Tanzania: New insight into Cretaceous and Paleogene terrestrial ecosystems and tectonics in sub-equatorial Africa". Journal of African Earth Sciences. 57 (3): 179–212. Bibcode:2010JAfES..57..179R. doi:10.1016/j.jafrearsci.2009.09.002.
  2. Stevens, Nancy J.; Holroyd, Patricia A.; Roberts, Eric M.; O'connor, Patrick M.; Gottfried, Michael D. (2009-06-12). "Kahawamys mbeyaensis (n. gen., n. sp.) (Rodentia: Thryonomyoidea) from the late Oligocene Rukwa Rift Basin, Tanzania". Journal of Vertebrate Paleontology. 29 (2): 631–634. doi:10.1671/039.029.0219. ISSN   0272-4634. S2CID   140590080.
  3. Blackburn, David C.; Paluh, Daniel J.; Krone, Isaac; Roberts, Eric M.; Stanley, Edward L.; Stevens, Nancy J. (2019-05-15). "The Earliest Fossil of the African Clawed Frog (Genus Xenopus) from Sub-Saharan Africa". Journal of Herpetology. 53 (2): 125. doi: 10.1670/18-139 . ISSN   0022-1511.
  4. Borths, Matthew R.; Stevens, Nancy J. (2017-10-11). Smith, Thierry (ed.). "The first hyaenodont from the late Oligocene Nsungwe Formation of Tanzania: Paleoecological insights into the Paleogene-Neogene carnivore transition". PLOS ONE. 12 (10): e0185301. Bibcode:2017PLoSO..1285301B. doi: 10.1371/journal.pone.0185301 . ISSN   1932-6203. PMC   5636082 . PMID   29020030.
  5. Stevens, Nancy J.; O’Connor, Patrick M.; Mtelela, Cassy; Roberts, Eric M. (2021-07-23). "Macroscelideans (Myohyracinae and Rhynchocyoninae) from the late Oligocene Nsungwe formation of the Rukwa Rift Basin, southwestern Tanzania". Historical Biology: 1–7. doi: 10.1080/08912963.2021.1938565 . ISSN   0891-2963.
  6. Stevens, Nancy J.; Seiffert, Erik R.; O'Connor, Patrick M.; Roberts, Eric M.; Schmitz, Mark D.; Krause, Cornelia; Gorscak, Eric; Ngasala, Sifa; Hieronymus, Tobin L.; Temu, Joseph (2013). "Palaeontological evidence for an Oligocene divergence between Old World monkeys and apes". Nature. 497 (7451): 611–614. doi:10.1038/nature12161.