Picea engelmannii

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Picea engelmannii
Engelmann spruce on the PCT, Washington.JPG
Mature tree with female cones
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Gymnospermae
Division: Pinophyta
Class: Pinopsida
Order: Pinales
Family: Pinaceae
Genus: Picea
Species:
P. engelmannii
Binomial name
Picea engelmannii
Picea engelmannii levila.png

Picea engelmannii, with the common names Engelmann spruce, [2] white spruce, [2] mountain spruce, [2] and silver spruce, [2] is a species of spruce native to western North America. It is mostly a high-elevation mountain tree but also appears in watered canyons. [3] [4]

Contents

Description

Picea engelmannii is a medium-sized to large evergreen tree growing to 25–40 metres (82–131 feet) tall, exceptionally to 65 m (213 ft) tall, and with a trunk diameter of up to 1.5 m (4 ft 11 in). The reddish bark is thin and scaly, [5] flaking off in small circular plates 5–10 centimetres (2–4 in) across. The crown is narrow conic in young trees, becoming cylindric in older trees. The shoots are buff-brown to orange-brown, usually densely pubescent, and with prominent pulvini. The leaves are needle-like, 15–30 millimetres (581+18 in) long, flexible, [5] rhombic in cross-section, glaucous blue-green above with several thin lines of stomata, and blue-white below with two broad bands of stomata. The needles have a pungent odour when crushed. [5]

Purple cones of about 1 cm appear in spring, releasing yellow pollen when windy. [5] The cones are pendulous, slender cylindrical, 2.5–8 cm long [5] and 1.5 cm broad when closed, opening to 3 cm broad. They have thin, flexible scales 15–20 mm long, with a wavy margin. They are reddish to dark purple, maturing to light brown [5] 4–7 months after pollination. The seeds are black, 2–3 mm long, with a slender, 5–8 mm long light brown wing.

The tree grows in a krummholz form along the fringe of alpine tundras. [5]

Distribution

Engelmann spruce is native to western North America, primarily in the Rocky Mountains and east slopes of the Cascade range from central British Columbia to Southern Oregon in the Cascades and commonly in Montana, Idaho, and Colorado, and more sparsely towards Arizona and New Mexico in the Sky islands; [5] there are also two isolated populations in Northern Mexico. It is mostly a high-elevation mountain tree, in many areas reaching the tree line, but at lower elevations occupies cool watered canyons. [5] It grows from 520–3,650 m (1,710–11,980 ft) above sea level, [5] rarely lower towards the northwest. It appears in the canyons of the Idaho Panhandle and more limitedly in the northeastern Olympic Mountains; the latter includes exceptionally large specimens, e.g. one 2.1 m (7 ft) thick and 55 m (179 ft) tall. [5] It can be found in the Cascade Range (mostly on the eastern slopes) from elevations of 900–1,800 m (3,000–5,900 ft) and liberally in the Rocky Mountains. [5] It can also be found in the Monashee and Selkirk Mountains, as well as the highlands surrounding the Interior Plateau.

Ecology

Both water uptake and water stored in roots appear to be critical for the survival of subalpine Engelmann spruce saplings that are exposed above the snowpack in later winter to early spring. [6] Transpiration is greatly reduced in small saplings while engulfed in snowpack. For exposed trees, the availability of soil water may be critical in late winter, when transpirational demands increase. Increased rates of transpiration in response to loss of snowpack, coupled with low sapwood water reserves and an extended period of soil frost in windswept areas, may prevent Engelmann spruce from regenerating in open areas both above and below the tree line. Cuticular damage by windblown ice is probably more important at the tree line, [7] [8] but damage caused by desiccation is likely to be more important at lower elevations. [6]

Despite wind damage, the species tends to grow taller than others at the tree line. [5] It is shade tolerant, but not so much as subalpine fir. Thus, it is somewhat dependent on fires to outgrow competitors, although its thin bark and shallow roots make it vulnerable to fire as well. [5] Spruce bark beetles attack the tree, being particularly deadly to groups which have stood for centuries. [5] It is also susceptible to avalanches. [5]

Although older spruce forests are not very useful to animals for forage, they are so in the aftermath of fires, which allow many other plants to rise. [5] Engelmann spruce-shaded streams are exploited by trout. [5] Additionally, aphids produce galls which hang from the tree and look similar to cones when they dry out. [5]

Subspecies and hybrids

Two geographical subspecies (treated as varieties by some authors, and as distinct species by others) occur:

The Engelmann spruce hybridises and intergrades extensively with the closely related white spruce ( Picea glauca ), [5] found further north and east in the Rockies, and to a lesser extent with the closely related Sitka spruce where they meet on the western fringes of the Cascades.

Uses

Native Americans made various medicines from the resin and foliage. [5]

Engelmann spruce is of economic importance for its wood, being light and fairly strong. [5] It harvested for paper-making and general construction. [5] Wood from slow-grown trees at high elevation has a specialized use in making musical instruments such as acoustic guitars, harps, violins, and pianos. [5] Because it is odourless and has little resin, it has been used for food containers such as barrels. [5] It is also used to a small extent as a Christmas tree.

Related Research Articles

<span class="mw-page-title-main">Spruce</span> Genus of evergreen, coniferous tree

A spruce is a tree of the genus Picea, a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. Picea is the sole genus in the subfamily Piceoideae. Spruces are large trees, from about 20 to 60 m tall when mature, and have whorled branches and conical form. They can be distinguished from other members of the pine family by their needles (leaves), which are four-sided and attached singly to small persistent peg-like structures on the branches, and by their cones, which hang downwards after they are pollinated. The needles are shed when 4–10 years old, leaving the branches rough with the retained pegs. In other similar genera, the branches are fairly smooth.

<i>Pinus albicaulis</i> Pine tree species found in North America

Pinus albicaulis, known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine areas of the Sierra Nevada, Cascade Range, Pacific Coast Ranges, and Rocky Mountains. It shares the common name "creeping pine" with several other plants.

<i>Picea mariana</i> North American species of spruce tree

Picea mariana, the black spruce, is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and all 3 territories. It is the official tree of the province of Newfoundland and Labrador and is that province's most numerous tree. The range of the black spruce extends into northern parts of the United States: in Alaska, the Great Lakes region, and the upper Northeast. It is a frequent part of the biome known as taiga or boreal forest.

<span class="mw-page-title-main">Tree line</span> Edge of the habitat at which trees are capable of growing

The tree line is the edge of a habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions. The tree line is sometimes distinguished from a lower timberline, which is the line below which trees form a forest with a closed canopy.

<i>Picea sitchensis</i> Species of large coniferous tree

Picea sitchensis, the Sitka spruce, is a large, coniferous, evergreen tree growing to almost 100 meters (330 ft) tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth-largest conifer in the world, and the third-tallest conifer species. The Sitka spruce is one of the few species documented to exceed 90 m (300 ft) in height. Its name is derived from the community of Sitka in southeast Alaska, where it is prevalent. Its range hugs the western coast of Canada and the US, continuing south into northernmost California.

<i>Abies grandis</i> Species of conifer tree

Abies grandis is a fir native to the Pacific Northwest and Northern California of North America, occurring at altitudes of sea level to 1,700 metres (5,600 ft). It is a major constituent of the Grand Fir/Douglas Fir Ecoregion of the Cascade Range.

<i>Abies concolor</i> Species of conifer tree

Abies concolor, the white fir, concolor fir, or Colorado fir, is a coniferous tree in the pine family Pinaceae. This tree is native to the mountains of western North America, including the Cascade Range and southern Rocky Mountains, and into the isolated mountain ranges of southern Arizona, New Mexico, and Northern Mexico. It naturally occurs at elevations between 900 and 3,400 metres.

<i>Pinus edulis</i> Species of plant

Pinus edulis, the Colorado pinyon, two-needle piñon, pinyon pine, or simply piñon, is a pine in the pinyon pine group whose ancestor was a member of the Madro-Tertiary Geoflora and is native to the United States.

<i>Tsuga heterophylla</i> Species of conifer

Tsuga heterophylla, the western hemlock or western hemlock-spruce, is a species of hemlock native to the west coast of North America, with its northwestern limit on the Kenai Peninsula, Alaska, and its southeastern limit in northern Sonoma County, California. The Latin species name means 'variable leaves'.

<i>Tsuga mertensiana</i> Species of tree found in western North America

Tsuga mertensiana, known as mountain hemlock, is a species of hemlock native to the west coast of North America, found between Southcentral Alaska and south-central California.

<i>Picea breweriana</i> Species of conifer

Picea breweriana, known as Brewer spruce, Brewer's weeping spruce, or weeping spruce, is a species of spruce native to western North America, where it is one of the rarest on the continent. The specific epithet breweriana is in honor of the American botanist William Henry Brewer.

<span class="mw-page-title-main">Blue spruce</span> Species of tree

The blue spruce, also commonly known as green spruce, Colorado spruce, or Colorado blue spruce, is a species of spruce tree. It is native to North America, and is found in USDA growing zones 1 through 7. It is found naturally in Arizona, Colorado, Idaho, New Mexico, Utah and Wyoming. It has been widely introduced elsewhere and is used as an ornamental tree in many places far beyond its native range. It does well in zones warmer than 7 where summer heat is moderate, as at San Francisco. The blue spruce has blue-green colored needles and is a coniferous tree.

<i>Picea jezoensis</i> Species of conifer

Picea jezoensis, the dark-bark spruce, Ezo spruce, Yezo spruce, or Jezo spruce, is a large evergreen tree growing to 30–50 m tall and with a trunk diameter of up to 2 m. It is native to northeast Asia, from the mountains of central Japan and the Changbai Mountains on the China-North Korea border, north to eastern Siberia, including the Sikhote-Alin, Kuril Islands, Sakhalin and Kamchatka. It is found in cold but humid temperate rain forests, and nowhere does its range extend more than 400 km from the Pacific Ocean. The specific epithet jezoensis derives from Ezo, an old name for Hokkaido and other islands north of the Japanese island of Honshu, where the species is found.

<i>Picea martinezii</i> Species of conifer

Picea martinezii, the Martinez spruce, is a medium-sized evergreen tree growing to 25–35 m tall, and with a trunk diameter of up to 1 m. It is native to northeast Mexico, where it occurs at six localities in the Sierra Madre Oriental mountains in Nuevo León. It grows at moderate altitudes from 2150–2600 m, growing along streamsides in mountain valleys, where moisture levels in the soil are greater than the otherwise low rainfall in the area would suggest.

<i>Picea chihuahuana</i> Species of conifer

Picea chihuahuana, the Chihuahua spruce, is a medium-sized evergreen tree growing to 25–35 m tall, and with a trunk diameter of up to 1 m. It is native to northwest Mexico, where it occurs in 25 small populations in the Sierra Madre Occidental mountains in Chihuahua and Durango. It grows at moderate altitudes from 2300–3200 m, growing along streamsides in mountain valleys, where moisture levels in the soil are greater than the otherwise low rainfall in the area would suggest.

<span class="mw-page-title-main">Fraser fir</span> Species of conifer

The Fraser fir, sometimes spelled" Frasier fir, is an endangered species of fir native to the Appalachian Mountains of the Southeastern United States. They are endemic to only seven montane regions in the Appalachian Mountains.

<i>Abies amabilis</i> Species of conifer

Abies amabilis, commonly known as the Pacific silver fir, is a fir native to the Pacific Northwest of North America, occurring in the Pacific Coast Ranges and the Cascade Range. It is also commonly referred to in English as the white fir, red fir, lovely fir, amabilis fir, Cascades fir, or silver fir. The species name is Latin for 'lovely'.

<i>Abies vejarii</i> Species of conifer

Abies vejarii is a species of fir native to northeastern Mexico, in the states of Coahuila, Nuevo León, and Tamaulipas, where it grows at high altitudes in the Sierra Madre Oriental.

<span class="mw-page-title-main">Snowbank fungus</span> Species of fungus

A snowbank fungus is any one of a number of diverse species of fungi that occur adjacent to or within melting snow. They are most commonly found in the mountains of western North America where a deep snowpack accumulates during the winter and slowly melts through the spring and summer, often shaded by coniferous forest. They may be saprotrophic, mycorrhizal, or in the case of Caloscypha fulgens, pathogenic.

<span class="mw-page-title-main">Fraser Experimental Forest</span>

The Fraser Experimental Forest is an outdoor research laboratory to study timber, water, wildlife management, and their integration in the high elevation subalpine coniferous forests. The experimental forest was established in 1937, and encompasses 9,328 hectares (36.02 sq mi). It is situated on the west side of the Continental Divide in north-central Colorado and includes the entire watershed of main Saint Louis Creek, a tributary of the Fraser River.

References

  1. Farjon, A. (2013). "Picea engelmannii". IUCN Red List of Threatened Species . 2013: e.T42322A2972365. doi: 10.2305/IUCN.UK.2013-1.RLTS.T42322A2972365.en . Retrieved 12 November 2021.
  2. 1 2 3 4 "Montana Plant Life". Archived from the original on 2016-06-26. Retrieved 2013-06-09.
  3. "Picea engelmannifiParry ex Engelm". www.srs.fs.usda.gov. Retrieved 2023-12-24.
  4. (PDF) https://plants.usda.gov/DocumentLibrary/plantguide/pdf/pg_pien.pdf.{{cite web}}: Missing or empty |title= (help)
  5. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Arno, Stephen F.; Hammerly, Ramona P. (2020) [1977]. Northwest Trees: Identifying & Understanding the Region's Native Trees (field guide ed.). Seattle: Mountaineers Books. pp. 91–96. ISBN   978-1-68051-329-5. OCLC   1141235469.
  6. 1 2 Boyce, R.L. and Lucero, S.A. 1999. Role of roots in winter water relations of Engelmann spruce saplings. Tree Physiol. 19:893–898.
  7. Hadley, J.L.; Smith, W.K. 1983. Influence of wind exposure on needle desiccation and mortality for timberline conifers in Wyoming, USA. Arctic Alpine Res. 15:127–135. (Cited in Coates et al. 1994).
  8. Hadley, J.L.; Smith, W.K. 1986. Wind effects on needles of timberline conifers seasonal influence on mortality. Ecology 67:12–19. Cited in Coates et al. (1994).
  9. "Picea engelmannii (Engelmann spruce) description".

Further reading