Pakicetidae is an extinct family of Archaeoceti that lived during the Early Eocene in Pakistan.
The evolution of cetaceans is thought to have begun in the Indian subcontinent from even-toed ungulates (Artiodactyla) 50 million years ago (mya) and to have proceeded over a period of at least 15 million years. Cetaceans are fully aquatic marine mammals belonging to the order Artiodactyla and branched off from other artiodactyls around 50 mya. Cetaceans are thought to have evolved during the Eocene, the second epoch of the present-extending Cenozoic Era. Molecular and morphological analyses suggest Cetacea share a relatively recent closest common ancestor with hippopotami and that they are sister groups. Being mammals, they surface to breathe air; they have 5 finger bones (even-toed) in their fins; they nurse their young; and, despite their fully aquatic life style, they retain many skeletal features from their terrestrial ancestors. Research conducted in the late 1970s in Pakistan revealed several stages in the transition of cetaceans from land to sea.
Ambulocetus is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene (Lutetian). It contains one species, Ambulocetus natans, known solely from a near-complete skeleton. Ambulocetus is among the best-studied of Eocene cetaceans, and serves as an instrumental find in the study of cetacean evolution and their transition from land to sea, as it was the first cetacean discovered to preserve a suite of adaptations consistent with an amphibious lifestyle. Ambulocetus is classified in the group Archaeoceti—the ancient forerunners of modern cetaceans whose members span the transition from land to sea—and in the family Ambulocetidae, which includes Himalayacetus and Gandakasia.
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, which prompted attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Basilosauridae is a family of extinct cetaceans. They lived during the middle to the early late Eocene and are known from all continents, including Antarctica. They were probably the first fully aquatic cetaceans. The group is noted to be a paraphyletic assemblage of stem group whales from which the monophyletic Neoceti are derived.
Pakicetus is an extinct genus of amphibious cetacean of the family Pakicetidae, which was endemic to Pakistan during the Ypresian period, about 50 million years ago. It was a wolf-like animal, about 1 metre to 2 metres long, and lived in and around water where it ate fish and other small animals. The vast majority of paleontologists regard it as the most basal whale, representing a transitional stage between land mammals and whales. It belongs to the even-toed ungulates with the closest living non-cetacean relative being the hippopotamus.
Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.
Himalayacetus is an extinct genus of carnivorous aquatic mammal of the family Ambulocetidae. The holotype was found in Himachal Pradesh, India, in what was the remnants of the ancient Tethys Ocean during the Early Eocene. This makes Himalayacetus the oldest archaeocete known, extending the fossil record of whales some 3.5 million years.
Gandakasia is an extinct genus of ambulocetid from Pakistan, that lived in the Eocene epoch. It probably caught its prey near rivers or streams.
Ichthyolestes is an extinct genus of archaic cetacean that was endemic to Indo-Pakistan during the Lutetian stage. To date, this monotypic genus is only represented by Ichthyolestes pinfoldi.
Kutchicetus is an extinct genus of early whale of the family Remingtonocetidae that lived during Early-Middle Eocene in what is now the coastal border of Pakistan and India. It is closely related to Andrewsiphius with which it was synonymized by Gingerich et al. 2001. Thewissen & Bajpai 2009 proposed a new clade, Andrewsiphiinae, for the two species. Later authors, however, still accept both as separate genera.
Indohyus is an extinct genus of digitigrade even-toed ungulates known from Eocene fossils in Asia. This small chevrotain-like animal found in the Himalayas is one of the earliest known non-cetacean ancestors of whales.
Remingtonocetus is an extinct genus of early cetacean freshwater aquatic mammals of the family Remingtonocetidae endemic to the coastline of the ancient Tethys Ocean during the Eocene. It was named after naturalist Remington Kellogg.
Eotheroides is an extinct genus of Eocene sirenian. It is an early member of the family Dugongidae, which includes the extant dugong. Fossils have been found from Egypt, India, and Madagascar. Eotheroides was first described by Richard Owen in 1875 under the name Eotherium, which was replaced by the current name in 1899.
Chonecetus is an extinct genus of primitive baleen whale of the family Aetiocetidae that lived in the Oligocene period. Its fossils have been found in Canada, in the northeast Pacific. It was first named by L.S. Russell in 1968, and contains one species, C. sookensis.
Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg.
Dalanistes is an extinct genus of remingtonocetid early whale known from the late early Eocene of Kutch, India and Punjab and Balochistan, Pakistan. Dalanistes is closely related to Remingtonocetus, but also shares several features with Ambulocetus, and, with its combination of terrestrial and amphibious adaptations, Dalanistes apparently is an intermediate form between these two groups. Isotopic evidence suggest that Dalanistes had a marine diet.
Gaviacetus is an extinct archaeocete whale that lived approximately 45 million years ago. Gaviacetus was named for its characteristic narrow rostrum and the fast pursuit predation suggested by its unfused sacral vertebrae.
Saghacetus is an extinct genus of basilosaurid early whale, fossils of which have been found in the Upper Eocene Qasr el Sagha Formation, Egypt.
Johannes Gerardus Marie (Hans) Thewissen is a Dutch-American paleontologist known for his significant contributions to the field of whale evolution. Thewissen's fieldwork has led to the discovery of key fossils that have shed light on the transition of whales from land to water, including the discovery of Ambulocetus, Pakicetus, Indohyus, and Kutchicetus. In addition to his work on fossil discoveries, Thewissen also studies modern bowhead and beluga whales in Alaska, focusing on their biology and the implications of this knowledge for management and conservation efforts. His research has been instrumental in deepening our understanding of cetacean evolution and the adaptations that allowed these mammals to transition from terrestrial to fully aquatic lifestyles.
