The Pennsylvanian is, on the ICS geologic timescale, the younger of two subperiods of the Carboniferous Period. It lasted from roughly 323.2 million years ago to 298.9 million years ago. As with most other geochronologic units, the rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal-producing beds of this age are widespread.
Reptiliomorpha is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes Homo sapiens, but not Ascaphus truei. Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing Homo sapiens, but not Pipa pipa, Caecilia tentaculata, and Siren lacertina.
Anthracosauria is an order of extinct reptile-like amphibians that flourished during the Carboniferous and early Permian periods, although precisely which species are included depends on one's definition of the taxon. "Anthracosauria" is sometimes used to refer to all tetrapods more closely related to amniotes such as reptiles, mammals, and birds, than to lissamphibians such as frogs and salamanders. An equivalent term to this definition would be Reptiliomorpha. Anthracosauria has also been used to refer to a smaller group of large, crocodilian-like aquatic tetrapods also known as embolomeres.
Diadectomorpha is a clade of large tetrapods that lived in Euramerica during the Carboniferous and Early Permian periods and in Asia during Late Permian (Wuchiapingian), They have typically been classified as advanced reptiliomorphs positioned close to, but outside of the clade Amniota, though some recent research has recovered them as the sister group to the traditional Synapsida within Amniota, based on inner ear anatomy and cladistic analyses. They include both large carnivorous and even larger herbivorous forms, some semi-aquatic and others fully terrestrial. The diadectomorphs seem to have originated during late Mississippian times, although they only became common after the Carboniferous rainforest collapse and flourished during the Late Pennsylvanian and Early Permian periods.
Silvanerpeton is an extinct genus of early reptiliomorph found by Stan Wood in the East Kirkton Quarry of West Lothian, Scotland, in a sequence from the Brigantian substage of the Viséan. The find is important, as the quarry represents terrestrial deposits from Romer's gap, a period poor in fossils where the higher groups "labyrinthodonts" evolved.
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
Pholiderpeton is an extinct genus of embolomere amphibian which lived in the Late Carboniferous period (Bashkirian) of England. The genus was first named by Thomas Henry Huxley in 1869 to include the species P. scutigerum, based on the disarticulated front half of a skeleton discovered near Bradford, Yorkshire. Associated fossil wood suggests that this specimen died inside a Lepidodendron tree trunk.
Diplovertebron is an extinct genus of embolomere that lived in the Late Carboniferous period (Moscovian), about 310 million years ago. Diplovertebron was a medium-sized animal, around 50 cm in length. Members of the genus inhabited European Carboniferous swamps in what is now the Czech Republic. They were closely related to larger swamp-dwelling tetrapods like Proterogyrinus and Anthracosaurus. However, Diplovertebron were much smaller than these large, crocodile-like creatures. Known from a single species, Diplovertebron punctatum, this genus has had a complicated history closely tied to Gephyrostegus, another genus of small, reptile-like amphibians.
Anthracosaurus is an extinct genus of embolomere, a possible distant relative of reptiles that lived during the Late Carboniferous in what is now Scotland, England, and Ohio. It was a large, aquatic eel-like predator. It has a robust skull about 40 centimetres (1.3 ft) in length with large teeth in the jaws and on the roof of the mouth. Anthracosaurus probably inhabited swamps, rivers and lakes. Its name is Greek for "coal lizard".
Pteroplax is an extinct genus of embolomerous anthracosaur. Only one species has been described; the skull table noted is the lectotype of this species. Pteroplax dates from the late Carboniferous Period, about 315 million years ago. It is known with certainty only from Newsham in Blyth, Northumberland, England. At that site, it shared a coal-swamp lake habitat with the larger embolomere, "Eogyrinus". Pteroplax probably grew up to about 10 ft (3.0 m) in length and was largely aquatic, feeding upon fish and smaller tetrapods. It likely had a long, eel-like body, with short limbs and a long tail.
Megalocephalus is an extinct genus of baphetid amphibian from the late Carboniferous of the British Isles and the United States (Ohio). It contains two species, M. pachycephalus and M. lineolatus.
The Allegheny Group, often termed the Allegheny Formation, is a Pennsylvanian-age geological unit in the Appalachian Plateau. It is a major coal-bearing unit in the eastern United States, extending through western and central Pennsylvania, western Maryland and West Virginia, and southeastern Ohio. Fossils of fishes such as Bandringa are known from the Kittaning Formation, which is part of the Allegheny Group.
Neopteroplax is an extinct genus of eogyrinid embolomere closely related to European genera such as Eogyrinus and Pteroplax. Members of this genus were among the largest embolomeres in North America. Neopteroplax is primarily known from a large skull found in Ohio, although fragmentary embolomere fossils from Texas and New Mexico have also been tentatively referred to the genus. Despite its similarities to specific European embolomeres, it can be distinguished from them due to a small number of skull and jaw features, most notably a lower surangular at the upper rear portion of the lower jaw.
Acherontiscus is an extinct genus of stegocephalians that lived in the Early Carboniferous of Scotland. The type and only species is Acherontiscus caledoniae, named by paleontologist Robert Carroll in 1969. Members of this genus have an unusual combination of features which makes their placement within amphibian-grade tetrapods uncertain. They possess multi-bone vertebrae similar to those of embolomeres, but also a skull similar to lepospondyls. The only known specimen of Acherontiscus possessed an elongated body similar to that of a snake or eel. No limbs were preserved, and evidence for their presence in close relatives of Acherontiscus is dubious at best. Phylogenetic analyses created by Marcello Ruta and other paleontologists in the 2000s indicate that Acherontiscus is part of Adelospondyli, closely related to other snake-like animals such as Adelogyrinus and Dolichopareias. Adelospondyls are traditionally placed within the group Lepospondyli due to their fused vertebrae. Some analyses published since 2007 have argued that adelospondyls such as Acherontiscus may not actually be lepospondyls, instead being close relatives or members of the family Colosteidae. This would indicate that they evolved prior to the split between the tetrapod lineage that leads to reptiles (Reptiliomorpha) and the one that leads to modern amphibians (Batrachomorpha). Members of this genus were probably aquatic animals that were able to swim using snake-like movements.
Odonterpeton is an extinct genus of "microsaur" from the Late Carboniferous of Ohio, containing the lone species Odonterpeton triangulare. It is known from a single partial skeleton preserving the skull, forelimbs, and the front part of the torso. The specimen was found in the abandoned Diamond Coal Mine of Linton, Ohio, a fossiliferous coal deposit dated to the late Moscovian stage, about 310 million years ago.
Adamanterpeton is a genus of Edopoid Temnospondyl within the family Cochleosauridae. The type species A. ohioensis was named in 1998 and is currently the only known species within this genus. Adamanterpeton is rare in the Linton vertebrate assemblage, with other amphibians like Sauropleura, Ophiderpeton, and Colosteus being more common. Unlike other Linton vertebrates, Adamanterpeton may have been adapted to a terrestrial lifestyle.
Eldeceeon is an extinct genus of reptiliomorph from the Mississippian of Scotland. It is known from two fossil specimens found within the Viséan-age East Kirkton Quarry in West Lothian. The type and only species, E. rolfei, was named in 1994. Eldeceeon is thought to be closely related to embolomeres, but it has several distinguishing features including long limbs and a short trunk. Initially known from two crushed partial skeletons, additional specimens have been reported by Ruta & Clack (2006). Eldeceeon was redescribed by Ruta, Clack, & Smithson (2020). The redescription supported affinities with Silvanerpeton, reconstructed a skull with larger eyes and a shorter snout, and emphasized potential correlations for an enlarge puboischiofemoralis internus 2 muscle.
Spondylerpeton is an extinct genus of tetrapod closely related to "Cricotus" (Archeria) in the family Archeriidae. This genus is known from fragmentary remains, namely a short series of tail vertebrae preserved in an ironstone nodule. These remains were found in the Mazon Creek beds of Illinois, an area famed for its preservation of Carboniferous plants and animals. Spondylerpeton individuals were probably about three to four feet in length, by far the largest animals known to have inhabited the Mazon Creek area during this era.
Palaeoherpeton is an extinct genus of eogyrinid embolomere which lived in the Pennsylvanian of Scotland. It is primarily known from a series of relatively small but well-preserved skulls. Some of these have among the best braincase and middle ear material known in embolomeres. Originally given the species name Palaeogyrinus decorus, this was later corrected to Palaeoherpeton decorum when it was determined that Palaeogyrinus was a name preoccupied by a genus of beetles.
The Scottish Coal Measures Group is a lithostratigraphic unit referring to the coal-bearing succession of rock strata which occur in Scotland during the Westphalian regional stage of the Carboniferous Period. It is the Scottish portion of the informal Coal Measures Group, which also includes the South Wales and Pennine coal measures. The Scottish coal measures are well-exposed across the Midland Valley. The group is also found within small grabens in the vicinity of Sanquhar and Thornhill.
